Anthonomus (Anthomorphus) cruraluma, Poinar, G & Legalov, AA, 2015

Poinar, G & Legalov, AA, 2015, New Curculioninae (Coleoptera: Curculionidae) in Dominican amber, Palaeontologia Electronica 677 (1), pp. 1-15 : 3-7

Anthonomus (Anthomorphus) cruraluma sp. nov.

Figure 3 , Figure 4 , Figure 5

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Holotype. Female; deposited in the PACO (accession # C-125).

Paratype. Male; deposited in the AMNH (accession # AMNH DR-8-414 ) .

Since many diagnostic features are obscured in the male specimen, we have designated the female as holotype.

Description. Length body, 2.0- 2.2 mm; length rostrum, 0.8-0.9 mm. Body reddish brown, with dense covering of light, recumbent, setiform scales.

Rostrum elongate, 1.3 (male) - 1.8 (female) times as long as pronotum, 7.5 (male) - 9.6 (female) times longer than wide at apex, 6.4 (male) - 8.4 (female) times longer than wide in middle, 6.0 (male) - 7.4 (female) times longer than wide at base, distinctly curved, without carinae, finely and slightly punctate; antennal scrobes lateral and directed toward middle of eye; forehead narrow, flattened, punctate; eyes large, rounded, distinctly convex, diameter much greater than rostrum at base; vertex weakly flattened, punctate; temples

POINAR & LEGALOV: NEW CURCULIONINAE quite short, 0.4 times as long as eye, punctate; antennae inserted in apical third of rostrum, elongate, almost reaching base of pronotum; scape elongate, 15.2 times longer than wide, 1.2 times as long as funicle length; funicle with first to seventh antennomeres conical-elongate; first antennomere 2.7 times longer than wide, 0.2 times as long as and 1.2 times as wide as scape; second to sixth antennomeres almost of equal width; second antennomere 3.0 times longer than wide, 0.8 times as long as first antennomere; third antennomere 2.0 times longer than wide, 0.7 times as long as

PALAEO- ELECTRONICA.ORG second antennomere; fourth antennomere equal to third antennomere; fifth antennomere 1.5 times longer than wide, 0.8 times as long as fourth antennomere; sixth antennomere equal to fifth antennomere; seventh antennomere 1.6 times longer than wide, 1.3 times as long as and 1.3 times as wide as sixth antennomere; club compact, 2.4 times longer than wide, 0.4 times as long as funicle; first club almost of equal length and width, equal to length and 1.6 times as wide as seventh antennomere; second club equal to first club article; third club article distinctly acuminate at apex.

Pronotum bell shaped; 1.8 times longer than wide at apex, 0.94 times longer than wide in middle, 0.91 times longer than wide at base; disk weakly convex, weakly narrowed at apex and base, densely punctate; scutellum trapezoidal, convex.

Elytra elongate and distinctly convex, 1.6 times longer than wide at base, 1.5 times longer than wide in middle, 2.2 times longer than wide at apical fourth, 2.1-2.4 times as long as pronotum, without transverse basal patch of scales; greatest width behind the middle, humeri distinctly convex; striae regularly and distinctly punctate; strial punctures oval, quite dense; elytral intervals convex, 1.2-2.0 times as long as striae; apex of elytra separately acuminate.

Thorax punctate; procoxal cavities more or less equidistant from anterior and posterior margin of prosternum; precoxal portion 0.4 times as long as procoxal cavity length; procoxal cavities contiguous; mesocoxal cavities widely separated, metepisternum narrow, 5.0 times longer than wide in middle.

Abdomen convex ventrally; ventrites almost homogeneous; first and second ventrites equal in length; third and fourth ventrites slightly shorter than previous; fifth ventrite elongate, 1.4 times as long as

POINAR & LEGALOV: NEW CURCULIONINAE fourth ventrite; fifth ventrite subquadrately emarginate of male; pygidium sulcate; pygidial channel broad and deep.

Legs long; pro- and mesocoxae conical; metacoxae transverse; femora weakly clavate; trochanters obconical; profemora bidentate, first tooth large and second tooth small; meso- and metafemora toothed; profemora 3.9-5.4 times longer than wide; mesofemora 6.1 times longer than wide; metafemora 4.6-4.8 times longer than wide; tibiae with weak prominence on ventral margin; protibia almost straight, with large mucro; meso- and metatibiae slightly curved, with mucro; protibiae 7.0-7.8 times longer than wide in middle; metatibiae 5.5 times longer than wide in middle, without ventral fringe of setiform scales; tarsi long; first to third tarsomeres conical; fifth elongate; tarsomeres with pulvilli on underside; tarsal claws free, large, with basal teeth.

Type Locality. Amber mine in the Cordillera Septentrional of the northern portion of the Dominican Republic.

Etymology. The specific epithet is from the Latin “luma” = thorne and the Latin “cruris” = leg in regards to the large mucro on the fore tibia.

Comparison. The new species is similar to Anthonomus filicornis Hustache, 1929 from Guadeloupe and South America but differs by the tibiae with mucro, body elongate, rostrum shorter, and the smaller body size.

Remarks. Placement of Anthonomus cruraluma sp. nov. (AMNH DR-8-414 and C-125) in the tribe Anthonomini was based on the antennal scrobes directed toward the eyes, the short prosternum,

PALAEO- ELECTRONICA.ORG convex eyes, and the nature of the scutellum. Placement of this species in the genus Anthonomus is based on the tarsal claws with basal teeth, contiguous procoxae, widely separated mesocoxae, slender rostrum, longer than pronotum along dorsal midline, 7-articled antennal funicle, bidentate femora lateral antennal scrobes directed toward the middle of the eye, large eyes with the diameter much greater than the rostrum at base, elytra without a transverse basal patch of scales and straight protibia. The new species belongs to the subgenus Anthomorphus ( Clark and Burke, 1987) based on the fifth ventrite subquadrately emarginate and pygidium sulcate.

Figures

FIGURE 3. Lateral view of Holotype Anthonomus cruraluma sp. nov. in Dominican amber. Bar equals 0.730 mm.

FIGURE 4. Dorsal view of Holotype Anthonomus cruraluma sp. nov. in Dominican amber. Bar equals 0.780 mm.

FIGURE 5. Lateral view of Paratype Anthonomus cruraluma sp. nov. in Dominican amber. Bar equals 0.500 mm.