Ctenognathus intermedius ( Broun, 1908 )

Larochelle & Larivière, Marie-Claude, 2021, Synopsis of the tribe Platynini in New Zealand (Coleoptera: Carabidae), Insecta Mundi 2021 (864), pp. 1-96 : 52-53

Ctenognathus intermedius ( Broun, 1908)

Fig. 73 , 116 , 135

Anchomenus intermedius Broun, 1908: 347 . Lectotype (here designated): male (BMNH) labeled “Type (circular red-bordered label; typed) / 2666. [male symbol] (hand-written) / [WI/WN] Manawatu Flats (hand-written) / Anchomenus intermedius . (hand-written) / New Zealand. Broun. Coll. Brit. Mus. 1922–482. (white label with red horizontal line; typed) / LECTOTYPE [male symbol] Anchomenus intermedius Broun, 1908 designated by Larochelle & Larivière, 2021 (red label; typed).”

Agonum (Anchomenus) intermedium: Csiki 1931: 865 .

Anchomenus intermedius: Hudson 1934: 174 .

Ctenognathus intermedius: Larochelle and Larivière 2007: 81 , 111.

Description. Body length 10.5–12.0 mm. Head, pronotum, and elytra mostly piceous black; lateral margins of pronotum and elytra reddish; abdomen piceous black, yellowish lateroapically; antennae and palpi rufotestaceous; femora and tibiae pale yellow; tarsi reddish. Microsculpture isodiametric on head, very transverse (with microlines) on pronotum, and isodiametric on elytra. Iridescence present on pronotum. Very shiny. Forebody narrow in comparison to elytra. Head. Moderately wide. Mandibles moderately long and curved anteriorly. Labrum strongly transverse, slightly emarginate anteriorly. Eyes strongly convex; two setiferous punctures on inner side of each eye. Tempora not inflated. Neck constriction shallow dorsolaterally. Mentum: medial tooth subtruncate apically; two setae. Submentum with six setae. Palpi with terminal segment truncate apically. Thorax. Pronotum moderately convex, impunctate, obsoletely wrinkled on disc and across base, narrow, moderately cordate, widest before middle; apex slightly emarginate; anterolateral angles poorly developed, obtuse; anterior bead complete; sides moderately rounded, moderately sinuate posteriorly; lateral beads absent; lateral depressions wide; two setiferous punctures (anterior and posterior) on each side; posterolateral angles rectangular; laterobasal foveae very deep, moderately wide, prolonged forward; posterior bead complete; base truncate. Legs. Very long. Metafemora with two posteroventral setae. Meso- and metatarsomeres 1–4 well developed, tricarinate dorsally, deeply bisulcate laterally. Metatarsomeres 4 moderately bilobed, symmetrical apically. Metatarsomeres 5 glabrous ventrally. Elytra. Slightly convex, not sloping down toward apex, subovate, widest about middle. Shoulders strongly rounded. Sides strongly rounded. Scutellar setiferous pore visible. Scutellar striole moderately long, impunctate. Striae deep, finely punctate. Intervals subdepressed; interval 3 with three moderately deep, small setiferous punctures. Umbilicate series with 17–19 setiferous punctures. Subapical sinuations feeble. Apices obtuse. Abdomen. Sterna IV–VI: both sexes with two long apical ambulatory setae. Sternum VII (last visible sternum): male with two long apical ambulatory setae; female with four long apical ambulatory setae. Aedeagus. Lateral view ( Fig. 116 ): moderately arcuate, slender; base moderately convex dorsally, with basal lobe narrow; middle subparallel, moderately convex dorsally, moderately concave ventrally, with dorsal membranous area moderately wide and moderately long; apex moderately convex dorsally, slightly concave ventrally, with extreme tip narrow and moderately long. Dorsal view: moderately wide, asymmetrical (ostium of membranous area deflected to the left); basal bulb moderately distant from membranous area.

Material examined. 54 specimens ( BMNH, JNNZ, MONZ, NZAC).

Geographic distribution ( Fig. 135 ). North Island: RI, TO, WI, WN.

Ecology. Lowland, montane. Epigean. Wet forests (beech, broadleaf). Associated with streams. Shaded ground; wet soil. Nocturnal; hides during the day under logs, fallen branches, and stones. Gregarious.

Biology. Seasonality: September–October, December–February. Mating observed in January (WI). Tenerals: February–April. Predacious (based on mouthpart morphology).

Dispersal power. Subapterous (incapable of flight). Swift runner.

Collecting techniques. Turning logs, fallen branches, and stones.

References. Larochelle and Larivière 2001: 130 (as “ Anchomenus ” intermedius ; catalogue; biology, dispersal power, ecology, geographic distribution, references), 2007: 81, 111 (as Ctenognathus intermedius ), 2016: 37 (list).

Remarks. Broun described Anchomenus intermedius from two specimens, one of which (a male) could be located (BMNH) and is here designated as lectotype. This type designation is made to preserve stability of nomenclature in the future.

Figures

Figures 73–76. Dorsal habitus. 73) Ctenognathus intermedius. 74) C. otagoensis. 75) C. colensonis. 76) C. earlyi new species. Scale line = 1 mm.

Figures 115–122. Aedeagus, lateral. 115) Ctenognathus xanthomelus. 116) C. intermedius. 117) C. colensonis. 118) C. otagoensis. 119) C. earlyi new species.120) C. davidsoni new species.121) C. oreobius. 122) C. hoarei new species.

Figures 130–135. Species distribution maps. 130) Ctenognathus edwardsii. 131) C. elevatus. 132) C. garnerae new species. 133) C. helmsi. 134) C. hoarei new species. 135) C. intermedius.