Willema Grishin, 2019

Cong, Qian, Zhang, Jing, Shen, Jinhui & Grishin, Nick V., 2019, Fifty new genera of Hesperiidae (Lepidoptera), Insecta Mundi 731 (731), pp. 1-56 : 20

Willema Grishin , new genus

http://zoobank.org/ D92BF2E1-F842-44C6-A924-023F495F9589

Type species. Heteropterus willemi Wallengren, 1857.

Definition. A sister genus to Hovala Evans, 1937 (type species Cyclopides pardalina Butler, 1879 ), more distant from Metisella Hemming, 1934 (type species Papilio metis Linnaeus, 1764), where it was placed previously ( Fig. 8 ). Keys to IV.20.B in Evans (1937). Shares many features with Metisella , such as hindwing discal cell longer than half of the wing, forewing veins R 1 and R 2 very near each other or touching (not separated). Distinguished from its relatives by the pattern of hindwing underside, which is with dark web-like pattern along veins with some streaks across and orange, ochreous, yellow or white spots. Hindwing below pale-yellow with dark web-like pattern along veins with some streaks across, but without darker markings. Genitalia with narrow and long penis, uncus with tegumen bulbous in dorsal view, uncus narrows to a point (except in W. willemi where distal end is mushroom-shaped and base of uncus with side processes), gnathos better developed than in Metisella , ampulla separated from harpe, protruding beyond its caudal end (except in W. willemi where harpe upturned embracing ampulla). In DNA, a combination of the following base pairs is diagnostic: aly363.37.1:A929C, aly 1450.10.1:A576G, aly283.4.1:T1129C, aly3555.2.2:T116C, aly451.23.2:C1177A.

Etymology. The name is a feminine noun in the nominative singular, derived from the type species name.

Species included. The type species, Cyclopides formosus var. tsadicus Aurivillius, 1905 , Cyclopides angolana Karsch, 1896 , Metisella kumbona Evans, 1937, Cyclopides carsoni Butler, 1898 , Heteropterus formosus Butler, 1894, and Cyclopides perexcellens Butler, 1896 .

Parent taxon. Tribe Heteropterini Aurivillius, 1925 .

Figures

Figure 8. Genomic trees of Heteropterinae and Trapezitinae. See Fig. 2 legend for notations.