Bonjeania jefferiesi, Winterton, Shaun L., 2007

Winterton, Shaun L., 2007, New species of Acraspisoides Hill & Winterton and Bonjeania Irwin & Lyneborg (Diptera: Therevidae: Agapophytinae), with the description of a new genus, Zootaxa 1438, pp. 1-25 : 14-15

Bonjeania jefferiesi sp. nov.

( Figure 5 )

Holotype male, AUSTRALIA: AUSTRALIAN CAPITAL TERRITORY: Mt. Ginini, 1700m, 4.i.1982, M.G. Jefferies (MEI 109594) ( ANIC).

Paratypes, AUSTRALIA: AUSTRALIAN CAPITAL TERRITORY: 2 males, same data as holotype (MEI 109595, 109598) ( ANIC); male, Picadilly Circus, 35°21’S, 148°47’E, 6.xii. 1994, M.G. Jefferies (MEI 109603) ( QDPI).

Diagnosis. Frons only slightly protruding, male frons slightly wider than ocellar tubercle at narrowest point; male abdomen without silver velutum; male genitalia with epandrium short, wider than long; aedeagus and gonocoxal apodemes not greatly elongate, gonocoxal apodemes about equal length to gonocoxites; distiphallus short, slightly curved ventrally; medial atrium relatively small.

Description. Male. Body length: 7.0 mm.

Head. Frons width slightly wider than ocellar tubercle at narrowest point; antennal base positioned low on frons; frons flat, rugose; frons and face only slightly protruding anteriorly, glossy black with elongate setae sparsely distributed; black setae on upper frons, pale setae laterally on lower frons and face; sparse silver pubescence on frons, denser laterally along margin of eye and on face beside and below antennal base; ocellar tubercle slightly raised, black with elongate, dark setae; occiput flat to slightly convex, black, overlain with grey pubescence; single row of relatively long, black, postocular setae dorsally; gena black, overlain with dense, silver-white pubescence admixed with long, white setae; palp and labellum brown with scattered, pale setae; antenna slightly shorter than head, brown, overlain with brown-grey pubescence; flagellum tinged with orange-brown pubescence; numerous elongate setae on scape, pedicel, and base of flagellum; setae pale ventrally, darker setae dorsally on scape and pedicel.

Thorax. Glossy black; scutum overlain with brown pubescence, pale grey to white pubescent laterally and posteriorly, pale, dorsocentral stripe; uniformly admixed with short, dark setae; scutal macrosetae black; scutellum overlain with olivaceous pubescence, pale laterally, admixed with dark, elongate setae; pleuron and coxae covered with dense, silver pubescence except on medial region of anepisternum, anepimeron and katatergite; elongate pale setae on anepisternum (few), katepisternum, katatergite and coxae; legs dark brown, tibia, basitarsi and hind femora paler basally; dark, slightly elongate setae ventrally on femora; wing hyaline with translucent brown infuscation distally in cells bm, d and basally in cell r5; venation brown; haltere brown; scutal chaetotaxy (pairs): np, 3–5; sa, 1; pa, 1; dc, 3–6; sc, 1 (+1–2 smaller macrosetae).

Abdomen. Uniform matte dark brown; dorsomedially overlain with olivaceous green to brown pubescence admixed with short, dark setae; tergites 1–4 with silver-white pubescence admixed with elongate, pale setae laterodorsally; intersegmental membrane of segments 2–4 white, brown medially; terminalia dark with pale setae.

Male genitalia. Genitalia similar shape to B. segnis and B. argentea sp. nov. Epandrium short, medially emarginate ( Fig. 5 A); cercus slightly elongate; tergite 8 medially emarginate ( Fig. 5 B); gonocoxal process shorter than gonostylus ( Figs 5 C–D); inner gonocoxal process equal length to gonostylus; gonocoxal apodeme shorter than gonocoxites; medial atrium very small; hypandrium triangular, fused to gonocoxite laterally; ventral lobe small, angular; sternite 8 broadly ovate to quadrangular ( Fig. 5 E); aedeagus relatively short, distiphallus only slightly curved dorsally along length ( Figs 5 F–G); dorsal apodeme of parameral sheath absent; lateral ejaculatory apodeme and ejaculatory apodeme enlarged; arms of ventral apodeme of aedeagal sheath relatively short, narrow.

Female. Similar in size and colouration to male.

Etymology. The specific epithet is a patronym in honour of Michael G. Jefferies, the collector of the specimens.

Comments. Bonjeania jefferiesi sp. nov. is closely related to B. argentea sp. nov. and B. segnis and is very similar in appearance to B. segnis . Bonjeania species typically have wing cell m3 closed, yet the type specimen of B. jefferiesi sp. nov. has wing cell m3 barely open in the left wing and closed in the right. Such variation is rare but does occur in other therevids. This species is known from the Australian Capital Territory.

Figures

FIGURE 5. Bonjeania jefferiesi sp. nov., male genitalia: A, epandrium, dorsal view; B, tergite 8, dorsal view; C, gonocoxite, lateral view; D, gonocoxites, ventral view; E, sternite 8, ventral view; F, aedeagus, lateral view; G, same, dorsal view. Scale line = 0.5 mm.