Pardosa fomichevi, Kronestedt & Marusik & Omelko, 2014

Kronestedt, Torbjörn, Marusik, Yuri M. & Omelko, Mikhail M., 2014, Studies on species of Holarctic Pardosa groups (Araneae, Lycosidae). VIII. The Palearctic species of the Pardosa nigra group, Zootaxa 3894 (1), pp. 33-60 : 38-42

Pardosa fomichevi sp. n.

Figs 2 , 10 , 17–18 , 21 , 29 , 37 , 46–47 , 60, 68, 77–78, 91, 100–101 , 129–130

Type material. Holotype ♂ and allotype ♀ from Russia: Primorsky Kray, Krasnoarmeisk District, slope of Ozernoe Plateau , glades near road, 1359 m (45º50’46”N 136º37’31”E) (M.M. Omelko), in ZMMU .— Paratypes. RUSSIA. Primorsky Kray: same data as holotype ( ISEA, NHRS), 2♂ .

Etymology. The specific epithet is a patronym for our colleague Alexander Fomichev from Novosibirsk.

Diagnosis. Males can be distinguished from all other members of this group by the curvature of the embolus ( Figs 46–47 ); females by the lateral elevations in the posterior part of the epigyne being at some distance from the septum and their margins diverging at the rear ends ( Fig. 18 ).

Description. Male (holotype): Total length 6.2. Carapace 3.10 long, 2.35 wide.

Prosoma. Carapace brownish. Median band yellowish, narrowed at cephalic-thoracic junction, wide behind PLEs. Lateral bands indistinct, yellowish brown, narrow, broken. Carapace with dark and adpressed light hairs on thoracic sides, median band with adpressed white hairs. Clypeus light brownish. Chelicerae light brownish with longitudinal darker streaks and yellow inside. Sternum dark greyish brown with narrow light stripe in front, furnished with light hairs.

Eyes. Width of row I (slightly procurved) 48, row II 69, row III 94, row II-III 67. Diameter of AME 10, ALE 9, PME 25, PLE 21. Distance between AME 5, between AME and ALE 3.

Opisthosoma. Dorsum brownish, covered with adpressed light hairs and more erect black hairs. Anterior part and lanceolate stripe with dense light pubescence. Dorsum patterned with black, notably laterally, forming two pairs of black blotches. Venter brown with adpressed light pubescence and numerous scattered short erect hairs. Spinnerets partly sooty with short black hairs.

Legs ( Table 1 ). Yellowish. Femora more or less pseudoannulated (=blotches and incomplete transverse markings) in greyish, Fe I and II with proximal half dark greyish. Ti III and IV with very weak darker annulation. Ti I with 3 pairs of ventral spines and two retrolateral spines.

Palp ( Figs 2 , 10 , 17 , 21 , 29 ). Pt 0.65, Ti 0.60, Cy 1.35. Brownish with darker markings. Longitudinal darker markings on Ti with with numerous strong black hairs. Cy blackish, distally light brown to yellowish. Tegular apophysis with ¾ of prolaterad rim of anteriorly directed branch very slightly concave ( Figs 17 , 22 , 37 ). Conductor as in Fig 78 , terminal apophysis as in Figs 60 , 68 & 77–78 . Embolus ( Figs 46–47 , 60, 68) wide, screw-shaped, apically curved inwards, narrowed, and tapering to pointed tip.

Female (allotype). Total length 6.8. Carapace 3.40 long, 2.60 wide.

Prosoma and opisthosoma. Carapace similar to that in the male, but with lateral bands more distinct. Opisthosoma patterned as in the male. Venter of opisthosoma brown with adpressed white pubescence.

Eyes. Width of row I (slightly procurved) 54, row II 76, row III 104, row II-III 74. Diameter of AME 11, ALE 10, PME 26, PLE 22. Distance between AME 7, between AME and ALE 4.

Legs ( Table 1 ). Yellowish, clearly annulated in grey (femora) and light brown (patellae-metatarsi). Ti I with two retrolateral spines.

Epigyne ( Figs 18 , 91 ). Lateral elevations diverging backwards (45º) from about half the length of the epigyne. Septum only partly filling the cavities in the posterior half. Spermathecae ovoid ( Figs 100–101 ).

Size variation. Carapace length: males 2.95–3.10 (n=3).

Distribution ( Figs 129–130 ). Russia: Primorsky Kray (only known from the type locality).

Figures

FIGURES 1–8. Left male palp, retrolateral. 1, Pardosa eiseni. 2, P. fomichevi sp. n. 3, P. giebeli. 4, P. lasciva. 5, P. lyrata. 6, P. nigra. 7, P. paramushirensis. 8, P. trailli. Scale lines 0.2 mm.

FIGURES 9–16. Left male palp, prolateral. 9, Pardosa eiseni. 10, P. fomichevi sp. n. 11, P. giebeli. 12, P. lasciva. 13, P. lyrata. 14, P. nigra. 15, P. paramushirensis. 16, P. trailli. Scale lines 0.2 mm.

FIGURES 17–19. 17-18, Pardosa fomichevi sp. n.: left male palp (17), epigyne (18). 19, P. trailli: left male palp. Scale lines 0.2 mm.

FIGURES 20–27. Bulbus of left male palp in ventral view. 20, Pardosa eiseni. 21, P. fomichevi sp. n. 22, P. giebeli. 23, P. lasciva. 24, P. lyrata. 25, P. nigra. 26, P. paramushirensis. 27, P. trailli. Scale lines 0.2 mm.

FIGURES 28–35. Bulbus of left male palp in oblique ventral-frontal view. 28, Pardosa eiseni. 29, P. fomichevi sp. n. 30, P. giebeli. 31, P. lasciva. 32, P. lyrata. 33, P. nigra. 34, P. paramushirensis. 35, P. trailli. Scale lines 0.2 mm.

FIGURES 36–43. Left male palp: tegulum with tegular apophysis. 36, Pardosa eiseni. 37, P. fomichevi sp. n. 38, P.giebeli. 39, P. lasciva. 40, P. lyrata. 41, P. nigra. 42, P. paramushirensis. 43, P. trailli. tg, tegulum; tg.ap, tegular apophysis. Scale line (applies to all) 300 µm.

FIGURES 44–58. Left male palp: embolus in frontal (44, 46, 48, 49, 51, 53, 55, 57) and ventral (45, 47, 50, 52, 54, 56, 58) view. 44–45, Pardosa eiseni. 46–47, P. fomichevi sp. n. 48, P. giebeli. 49–50, P. lasciva. 51–52, P. lyrata. 53–54, P. nigra. 55–56, P. paramushirensis. 57–58, P. trailli. Scale lines 0.2 mm.

FIGURES 98–113. Epigynes, cleared, in ventral (98, 100, 102, 104, 106, 108, 110, 112) and dorsal (99, 101, 103, 105, 107, 109, 111, 113) view. 98-99, Pardosa eiseni. 100–101, P. fomichevi sp. n. 102–103, P. giebeli. 104–105, P. lasciva. 106–107, P. lyrata. 108–109, P. nigra. 110–111, P. paramushirensis. 112–113, P. trailli. Scale lines 0.2 mm.

FIGURES 75–80. Left male palp: terminal part ventral view. 75, Pardosa eiseni. 76, P. giebeli. 77–78, P. fomichevi sp. n. from two oblique angles. 79–80, P. lasciva from two oblique angles. cond, conductor; emb, embolus; pal, palea; tl.ap, terminal apophysis. Scale line (applies to all) 300 µm

FIGURES 59–66. Left male palp: terminal part oblique ventral-frontal view. 59, Pardosa eiseni. 60, P. fomichevi sp. n. 61, P. giebeli. 62, P. lasciva. 63, P. lyrata. 64, P. nigra. 65, P. paramushirensis. 66, P. trailli. Scale lines 0.2 mm.

FIGURES 67–74. Left male palp: terminal part ventral view. 67, Pardosa eiseni. 68, P. fomichevi sp. n. 69, P. giebeli. 70, P. lasciva. 71, P. lyrata. 72, P. nigra. 73, P. paramushirensis. 74, P. trailli. Scale lines 0.2 mm.

FIGURES 90–97. Epigynes in ventral view. 90, Pardosa eiseni. 91, P. fomichevi sp. n. 92, P. giebeli. 93, P. lasciva. 94, P. lyrata. 95, P. nigra. 96, P. paramushirensis. 97, P. trailli. Arrows: see text. Scale lines 0.2 mm.

FIGURES 129–130. Distribution of Pardosa eiseni (black circles), P. fomichevi sp. n. (black square), P. lasciva (blue crosses), P. lyrata (green diamonds), P. paramushirensis (red stars) and P. trailli (red triangles) in Eurasia (129) and in Khabarovsk and Primorsky Krays (130). “?” in symbol indicates doubtful record.

Tables

TABLE 1. Leg I–IV measurements (mm) of Pardosa fomichevi sp. n., P. lyrata and P. paramushirensis.

Pardosa fomichevi Male
	Fe	Pt	Ti	Mt	Ta	Total
I	2.55	1.10	2.15	2.40	1.55	9.75
II	2.45	1.05	1.95	2.35	1.40	9.20
III	2.30	1.00	1.80	2.55	1.30	8.95
IV	3.00	1.10	2.40	3.90	1.75	12.15
Female
	Fe	Pt	Ti	Mt	Ta	Total
I	2.80	1.25	2.30	2.30	1.40	10.05
II	2.70	1.20	2.10	2.30	1.35	9.65
III	2.60	1.10	1.95	2.65	1.30	9.60
IV	3.50	1.30	2.85	4.45	1.80	13.90
Pardosa lyrata Male
	Fe	Pt	Ti	Mt	Ta	Total
I	2.50	1.10	2.40	2.65	1.60	10.25
II	2.40	1.05	2.05	2.50	1.40	9.40
III	2.30	1.00	1.80	2.60	1.30	9.00
IV	3.00	1.10	2.50	4.10	1.75	12.45
Female
	Fe	Pt	Ti	Mt	Ta	Total
I	2.70	1.20	2.30	2.30	1.35	9.85
II	2.60	1.15	2.05	2.25	1.30	9.35
III	2.45	1.05	1.90	2.60	1.25	9.25
IV	3.25	1.20	2.75	4.20	1.70	13.10
Pardosa paramushirensis Male
	Fe	Pt	Ti	Mt	Ta	Total
I	2.50	1.10	2.05	2.30	1.45	9.40
II	2.35	1.05	1.85	2.20	1.35	8.80
III	2.30	1.00	1.70	2.45	1.30	8.75
IV	3.00	1.10	2.35	3.80	1.75	12.00
Female
	Fe	Pt	Ti	Mt	Ta	Total
I	2.50	1.25	2.10	2.10	1.35	9.30
II	2.50	1.20	1.85	2.10	1.30	8.95
III	2.40	1.10	1.85	2.40	1.25	9.00
IV	3.15	1.25	2.70	3.95	1.70	12.75