Chlamydatus becki Knight, 1968: 28

SCHUH, RANDALL T. & SCHWARTZ, MICHAEL D., 2005, Review of North American Chlamydatus Curtis Species, with New Synonymy and the Description of Two New Species (Heteroptera: Miridae: Phylinae), American Museum Novitates 3471 (1), pp. 1-56 : 26-37

Chlamydatus becki Knight, 1968: 28 (n.sp., host).

DIAGNOSIS: Along with uniformis , the only North American species with completely dark legs, including the tibiae. Easily distinguished from C. uniformis by its smaller size, total length 2.23 mm or less, whereas that species always 2.65 mm or more. Coloration in C. becki brownish black, whereas coloration in C. uniformis jet black, shining. Dorsal vestiture in C. becki shining, distinctly visible in dorsal perspective, whereas vestiture in C. uniformis black, not contrasting with coloration of dorsum.

HOST: Artemisia sp. (Asteraceae) .

DISTRIBUTION: Great Basin, including western Utah, Nevada , and eastern Mojave Desert .

SPECIMENS EXAMINED: Holotype: male: ‘‘ American Fork, Utah, June 28, 1965, Harry H. Knight’ ’; deposited in the National Museum of National History , Washington, D.C.

Additional specimens: USA: California: San Bernardino Co.: New York Mts., Live Oak Canyon, 5500 ft, May 25, 1977, J. D. Pinto, 23 (UCR). San Diego Co.: No specific locality, May 28, 1923, W. Benedict, 1♀ (KU). Nevada: Carson City Co. : Carson City, 6000 ft, July 11, 1965, H. H. Knight, 13 (USNM). Elko Co.: 30 mi W of Elko, June 18, 1982, T. J. Henry, Artemisia sp. (Asteraceae) , 33, 14♀ (USNM). Nye Co.: Atomic Test Site, 2 mi W of Tippapah Hwy on Mine Mt. Road, 4400 ft, June 7, 1983, Schuh, Schwartz, Stonedahl, 13, 1♀ (AMNH). Mercury, 6 M, June 17, 1965, H. H. Knight and J. Merino, paratype, 13 (USNM). Utah: Millard Co.: Kanosh, July 7, 1965, G. F. Knowlton, 1♀ (USU). Scipio, June 29, 1965, H. H. Knight, 1♀ (USNM). Tooele Co.: Benmore, environs, June 19, 1954, G. L. Nielson, 13 (USU). Utah Co.: American Fork, June 28, 1965, H. H. Knight, paratypes, 23, 1♀ (CNC). Wasatch Co.: Heber, July 5, 1922, E. P. Van Duzee, 3♀ (CAS).

Chlamydatus brevicornis Knight

Figures 1 , 3 , 4

Chlamydatus brevicornis Knight, 1964: 141 (n.sp.).

DIAGNOSIS: Recognized by the unicolorous dark coloration, the somewhat woolly, shining pubescence on the dorsum, and the dark femora and pale tibiae. Distinguished from C. keltoni , C. monilipes , and C. suavis , the three species with similar coloration, by the shining, somewhat woolly pubescence on the dorsum, whereas in those species the dorsal vestiture always dark.

HOST: Artemisia tridentata (Asteraceae) .

DISTRIBUTION: Western North America. Restricted to higher latitudes and altitudes.

SPECIMENS EXAMINED: Holotype: male: ‘‘ Shoshone N. For., Wyo., Aug. 14, 1927, H. H. Knight’ ’; deposited in the National Museum of Natural History , Washington, D.C.

Additional specimens: CANADA: British Columbia : Richter Pass, Osoyoos, June 25, 1959, L. A. Kelton, 1♀ (CNC). USA: California: Alpine Co. : E of Pacific Grade Summit on Rt 4, Sandy Meadow Trail head, near Toiyabe National Forest, 8000 ft, July 5, 1994, M. D. Schwartz, Artemisia sp. (Asteraceae) , 13, 1♀ (AMNH). Modoc Co.: 24.7 mi NW of Canby, 1375 m, July 1, 1979, R. T. Schuh and B. M. Massie, 7♀ (AMNH). Mono Co.: Tioga Pass, July 31, 1940, D. E. Hardy, 13 (KU). Nevada Co. : Sagehen Creek, June 27, 1972, M. L. Kok, 1♀ (UCD). Siskiyou Co.: Bray, June 20, 1935, R. H. Beamer, 1♀ (KU). Dorris, July 1, 1935, R. H. Beamer, 13, 1♀ (KU). Colorado: Jackson Co.: Gould, 9200 ft, August 12, 13, 1968, L. A. Kelton, 13, 3♀ (CNC). Routt Co.: Steamboat Springs, 6700–6900 ft, July 15, 1964, H. H. Knight, 53, 4♀ (CNC). Steamboat Springs, June 30, 1931, R. H. Beamer, 1♀ (KU). Saquache Co.: 23 mi E of Gunnison on Rt 50, 8100 ft, July 17, 1986, R. T. Schuh, Artemisia tridentata (Asteraceae) , 23, 1♀ (AMNH). Idaho: Gooding Co.: Wendell, September 6, 1933, 13, 1♀ (CNC). Oneida Co. : 6 mi NW of Holbrook, June 9, 1969, G. F. Knowlton, Artemisia tridentata (Asteraceae) , 23 (USU). Nevada: Elko Co. : 30 mi W of Elko on Rt 80, June 18, 1982, T. J. Henry, Artemisia sp. (Asteraceae) , 53, 3♀ (USNM). Oregon: Bak­ er Co.: 4 mi E of Unity, June 8, 1970, P. Oman, 13 (OSU). Crook Co.: Redmond, June 1, 1938, Schuh and Gray, 13 (AMNH). Klamath Co.: Chiloquin, 10 mi SE of, on Interstate 15, July 4, 1982, G. M. Stonedahl and T. J. Henry, Artemisia tridentata (Asteraceae) , 63, 8♀ (AMNH, USNM). Junction of Rts 97 and 31, 28 mi SE of, 4910 ft, July 25, 1979, M. D. Schwartz, 3♀ (AMNH). Keno, 6 mi SE of, June 27, 1979, M. D. Schwartz, Artemisia tridentata (Asteraceae) , 1♀ (AMNH). Klamath Falls, July 5, 1953, J. Schuh, 13, 1♀ (CNC). Lower Klamath Lake, Hill Area, July 10, 1953, J. Schuh, 33 (CNC). Swan Lake, July 12, 1953, J. Schuh, 13 (CNC). Lake Co.: Abert Lake, July 2, 1935, Joe Schuh, 13 (AMNH). Fort Rock, June 15, 1938, K. Gray and J. Schuh, Artemisia sp. (Asteraceae) , 13 (AMNH). Silver Lake, June 15, 1938, Schuh and Gray, 1♀ (AMNH). Silver Lake, June 16, 1938, J. Schuh, Gray, 33, 1♀ (CNC). Silver Lake, 11 mi NW of, on Hwy 31, 4500 ft, June 25, 1979, M. D. Schwartz and G. M. Stonedahl, Artemisia tridentata (Asteraceae) , 23, 2♀ (AMNH). Utah: Cache Co.: Franklin Basin Road, 0.4 mi N of Rt 89, 6575 ft, July 18, 2001, T. J. Henry and A. G. Wheeler, Jr., Artemisia tridentata (Asteraceae) , 33, 30♀ (USNM). Logan Canyon, July 24, 1948, G. G. Fleener, Artemisia sp. (Asteraceae) , 1♀ (USU). Rich Co.: Garden City, June 20, 1969, G. F. Knowlton, 1♀ (USU). Sevier Co.: Fish Lake National Forest, R2E T25S, sec. 24, 25, July 16, 1980, G. M. Stonedahl, Artemisia tridentata (Asteraceae) , 13 (AMNH). Wasatch Co.: 30 mi SE of Kamas on Rt 34, Uintah National Forest, Wolf Creek Campground, T4S, R10W, sec. 7, 9000 ft, August 15, 1986, Schwartz and Stonedahl, Artemisia tridentata (Asteraceae) , 83, 36♀ (AMNH). Wyoming: Lincoln Co.: 15 mi S of Afton on Rt 89, Salt River Pass, 7630 ft, July 21, 1981, M. D. Schwartz, Artemisia tridentata (Asteraceae) , 1♀ (AMNH). Sublette Co.: 25 mi SE of Jackson, Hobart River, June 27, 1966, W. Gagne and J. Haddock, 13 (UCB). Sweetwater Co.: 10 mi E of Rock Springs, July 4, 1968, H. H. Knight, 1♀ (USNM).

Chlamydatus californicus , new species

Figures 1 , 3D , 4

DIAGNOSIS: Recognized by the abbreviated hemelytra (staphylinoid) with rounded apices, the highly polished and shining frons, and the generally brown coloration. Most similar in structure of the wings to C. opacus , C. ruficornis (fig. 1), and C. wilkinsoni (fig. 2); separated from C. opacus by the dull vertex and face with only the clypeus and maxillary and mandibular plates shining in that species, whereas entire vertex and face shining in C. californicus ; C. opacus furthermore known only from high latitudes in Canada and C. californicus from the southern half of California; male genitalia in C. californicus with a much longer apical attenuation of the vesica than that in opacus ; separated from C. ruficornis by the red, weakly fusiform second antennal segment, uniform slate gray coloration, and entirely dull vertex and face in that species; separated from C. wilkinsoni by the smaller size (1.28–1.89 mm) and uniform, nearly black coloration in that species; C. wilkinsoni similar to C. californicus in possession of highly polished vertex and face. Females of C. californicus (fig. 1) much larger than males and largely pale, in contrast to mostly brown males.

DESCRIPTION: Male: Moderately small, all known specimens brachypterous, hemelytra cover about half of abdomen; total length 1.60–1.94, length apex clypeus–cuneal fracture 1.60, width across pronotum 0.72. COL­ ORATION (fig. 1): Body and antennae mostly deep brown to castaneous, legs somewhat paler, reddish. SURFACE AND VESTITU­ RE (figs. 1, 7C): Pronotum, scutellum, hemelytra, abdominal dorsum, thoracic pleuron, and abdominal venter granular, dull; vertex and face, including clypeus and mandibular and maxillary plates highly polished and shining; legs polished, but not so strongly as face; dorsum and abdominal venter covered with recumbent, shining, weakly woolly setae (fig. 1). STRUCTURE: Vertex and eyes more or less conforming to anterior margin of pronotum, eyes extending posteriorly along anterolateral angles of pronotum (fig. 1); posterior margin of vertex and eyes smoothly and rather broadly rounded (fig. 1); frons projecting well beyond anterior margin of eyes, broadly transversely rounded in dorsal view; clypeus not visible from above; antennae inserted just below ventral margin of eye; eyes occupying about half of height of head in lateral view; labium very long, reaching to about anterior margin of genital capsule; hemelytra truncate, posterior margin broadly rounded, devoid of membrane (fig. 1); claws of moderate length, bent at about 20° angle at apex of pulvilli, pulvilli moderately large and covering approximately half of the length of claw, parempodia setiform and attenuated; abdomen broad, genital capsule occupying about half the length of abdomen. MALE GENITALIA (fig. 4): Vesica composed of a single strap forming a single coil with a weakly curving apical attenuation; secondary gonopore well sclerotized, distant from apex of vesica, gonopore sclerite lacking.

Female: Distinctly larger than male; total length 2.57–2.77, length apex clypeus–cuneal fracture 1.76–1.87, width pronotum 0.73–0.85. COLORATION (fig. 1): Overall coloration much paler than in male, mostly pale with some darker markings. SURFACE AND VESTITURE (fig. 1): As in male. STRUCTURE (fig. 1): Similar to male.

ETYMOLOGY: Named for its apparently exclusive occurrence in California.

HOST: Hemizonia pungens (Asteraceae) .

DISTRIBUTION: California: San Joaquin Valley .

SPECIMENS EXAMINED: Holotype: male: ‘‘ California, San Benito, Big Panoche Creek , April 21, 1967, J. Powell and P. Opler’ ’; deposited in the California Academy of Sciences , San Francisco.

Paratypes: USA: California: Fresno Co. : Mendota, April 25, 1932, E. P. Van Duzee, Centro­ madia (= Hemizonia ) pungens (Asteraceae) , 18♀ (CAS, AMNH). San Benito, Big Panoche Creek, April 21, 1967, J. Powell and P. Opler, 23, 25♀ (UCB, AMNH).

Chlamydatus keltoni , new species

Figures 1 , 3H , 4 , 8

Chlamydatus pullus: Kelton, 1965: 1134 (misidentification).

DIAGNOSIS: Recognized by the uniformly dark coloration of the body, the dark setae on the dorsum, the dark femora with a pale band distally, the pale tibiae with dark spots at the base of the tibial spines, and the convoluted form of the vesica in the male. Similar to C. pulicarius and C. pullus in body form and coloration, but usually smaller than C. pulicarius , ranging in length from 2.29 to 2.61 whereas total length in C. pulicarius always 2.51 or more; males usually macropterous, an uncommon condition in C. pulicarius . Readily distinguished from C. pulicarius by the form of the male genitalia, the apical process of the vesica longer and the vesica more highly convoluted in C. pulicarius (fig. 5) than in C. keltoni (fig. 4); differences in vesical structure not so striking between C. keltoni and C. pullus , but C. pullus (fig. 6) with less highly convoluted vesica and apex somewhat shorter than in C. keltoni (fig. 4). Similar in size and coloration to C. brevicornis , C. monilipes , and C. suavis , relatively small species with all femora dark and the background coloration of the tibiae pale. Distinguished from C. suavis by generally brown coloration and faint brown bases of the tibial spines in that species (fig. 3G), from C. brevicornis by the shining, weakly woolly setae in that species (fig. 1) as opposed to the inconspicuous dorsal vestiture of dark common setae in C. keltoni (fig. 1), and from C. monilipes by the narrower pale area at the tibial articulation of the femora and white tibiae with contrasting heavy black bases of the tibial spines in that species (fig. 3). Male genitalia unlike those of C. brevicornis , C. monilipes , and C. suavis with their relatively short apical projection of the vesica (figs. 4–6).

DESCRIPTION: Male: Moderately small, broad bodied, somewhat flattened, total length 2.29–2.61, length apex clypeus–cuneal fracture 1.63–1.79, width across pronotum 0.82–0.88. COLORATION (figs. 1, 3H): Body and appendages castaneous to nearly black; femora with a moderately broad, pale, distal band; tibiae and tarsal segments 1 and 2 pale; tibial spines with dark bases, these being most distinct on hind tibiae; membrane of wing fumose, veins darker than surrounding areas. SURFACE AND VESTITURE (figs. 1, 7C): Dorsum granular, with weakly rugose appearance, weakly shining, covered with recumbent, dark, simple setae (fig. 1). STRUCTURE: Curvature of posterior margin of vertex and eyes more or less conforming to anterior margin of pronotum (fig. 1); posterior margin of vertex and eyes smoothly and rather broadly rounded (fig. 1); vertex and frons moderately rounded; clypeus at most barely visible from above; dorsal margin of antennal fossa at level of ventral margin of eye; eyes occupying about 60% of height of head in lateral view (fig. 8A); labium reaching to about posterior margin of hind trochanters; costal margin of hemelytra distinctly rounded, convex, cuneal fracture weakly incised, hemelytra weakly deflexed at fracture (fig. 1); claws of moderate length, bent at about 20° angle at apex of pulvilli, pulvilli moderately large and covering approximately three­fourths of the length of claw (fig. 8C,D), parempodia setiform and attenuated (fig. 8D); abdomen somewhat flattened, relatively broad, widest at midpoint, tapering toward relatively small genital capsule, the latter occupying than one­half length of abdomen. MALE GENITALIA (fig. 4): Vesica formed of a single strap curving in a convoluted manner and with a long, decurved, apical attenuation; secondary gonopore well sclerotized, distant from apex of vesica, gonopore sclerite lacking; phallotheca with elongate, more or less parallel­sided apical portion; left paramere boat shaped; right paramere lanceolate.

Female: Small; total length 2.04–2.45, length apex clypeus–cuneal fracture 1.58– 1.85, width pronotum 0.73–0.85. COLOR­ ATION (fig. 1): As in male, except distal half of antennal segment 2 pale. SURFACE AND VESTITURE (figs. 1, 8B): As in male. STRUCTURE (fig. 1): Usually submacropterous, membrane of wing shorter than in male, body broader, with lateral margins of hemelytra more strongly convexly rounded; eyes smaller than in male in dorsal view, vertex broader relative to width of head, but eyes of similar height to male in lateral view; head more strongly projecting anteriorly than in male, clypeus readily visible from above.

ETYMOLOGY: Named after Leonard A. Kelton, in recognition of his pioneering work on New World Chlamydatus , and North American Miridae in general.

HOSTS: Recorded from a variety of plant families, including—among others—the Fabaceae ( Astragalus , Lupinus , and Medicago spp. ), Ranunculaceae ( Ranunculus sp. ), and Rosaceae ( Potentilla spp. )

DISTRIBUTION: Distribution from Alaska to Greenland at higher latitudes; at lower latitudes restricted to the western United States, the most southern occurrence being in Colorado.

DISCUSSION: Kelton (1965) recorded this species from North America under the name Chlamydatus pullus . Comparison of male genitalia indicates that his concept for the species was based on misidentified specimens, as confirmed by I. M. Kerzhner (in litt.).

SPECIMENS EXAMINED: Holotype: male: ‘‘[CAN­ ADA:] Miller Lake, ONT.[,]12.VI.62[,] Kelton & Thorpe’’; deposited in the Canadian National Collection of Insects, Ottawa.

Paratypes: CANADA: Alberta: Banff, 5000– 6000 ft, July 4, 1925, O. Bryant, 13 (USNM). Banff, July 5, 1924, E. Hearle, 3♀ (CNC). Calgary, June 18, 1963, Carr, 1♀ (CNC). Calgary, 32 mi W of, July 18, 1970, P. Oman, 23, 4♀ (OSU). Canmore, August 27, 1952, L. A. Konotopetz, 13 (CNC). Coal Valley, August 31, 1970, L. A. Kelton, 23, 3♀ (CNC). Coults, June 15, 1952, L. A. Konotopetz, Lupinus argenteus (Fabaceae) , 143, 4♀ (CNC). Cypress Hills Provincial Park, Horseshoe Canyon Overlook, 4700 ft, July 18, 1990, M. D. Schwartz, 13 (CNC). Cypress Hills Provincial Park, Top Road, 1.2 km E of Spruce Coulee, 4700 ft, July 15, 1990, M. D. Schwartz, Potentilla fruticosa (Rosaceae) , 23 (CNC). Cypress Hills Provincial Park, Top Road, 9.6 km E of Rt 41 jct, 4700 ft, July 15, 1990, M. D. Schwartz, 33, 2♀ (CNC). Del Bonita, June 16, 1952, L. A. Konotopetz, 33 (CNC). Drumheller, June 18, 1957, Brooks, MacNay, 23, 1♀ (CNC). Eisenhower Junction, August 5, 1970, L. A. Kelton, 193, 38♀ (CNC). Eisenhower Junction, Banff National Park, 4700 ft, June 23, 1955, J. Coyles, 13 (CNC). Elkwater Lake, July 19, 1956, O. Peck, 2♀ (CNC). Elkwater Park, around marina, 3900 ft, July 16, 1990, M. D. Schwartz, Potentilla fruticosa (Rosaceae) , 33, 2♀ (CNC). Elkwater Park, July 7, 1952 and July 26, 1952, L. A. Konotopetz, 63, 6♀ (CNC). Fisher Creek A. F. S., July 26, 1973, L. A. Kelton, 83, 2♀ (CNC). Frank, June 18, 1952, A. R. Brooks, 13 (CNC). Geikie Road, Jasper National Park, August 29, 1932, R. H. Handford, 2♀ (CNC). Grande Prairie, July 4, 1940, C. L. Nelson, 2♀ (CNC). Grande Prairie, June 9, 1961 and July 26, 1961, A. R. Brooks, 33, 2♀ (CNC). High Prairie, June 22, 1961 and September 6, 1961, A. R. Brooks, 33, 2♀ (CNC). Jasper National Park, August 6, 1952, L. A. Konotopetz, 13 (CNC). Jumping Pound Creek, June 21, 1963, Carr, 1♀ (CNC). Kananaskis Highway, July 25, 1973, L. A. Kelton, 153, 14♀ (CNC). Kananaskis Road, July 20, 1974 and July 25, 1973, L. A. Kelton, 353, 32♀ (CNC). Kananaskis Valley, Pocaterra Creek, 5350 ft, July 15, 1964, H. B. Leach, 73, 1♀ (CAS). Lethbridge, June 20, 1952, L. A. Konotopetz, Trifolium repens (Fabaceae) , 23, 1♀ (CNC). Lethbridge, June 21, 1956, O. Peck, Medicago sativa (Fabaceae) , 13, 1♀ (CNC). Lethbridge, July 2, 1938, R. W. Salt, Medicago sativa (Fabaceae) , 33, 1♀ (CNC). Manyberries, June 21, 1952, A. R. Brooks, 1♀ (CNC). McMurray, August 7, 1953, G. E. Ball, 23 (CNC). Medicine Hat, July 28, 1930, F. S. Carr, 1♀ (USNM). Mile 14 Banff­ Jasper Highway, Banff, August 2, 1955, J. Coyles, 13 (CNC). Nordegg, August 31, 1970, L. A. Kelton, 13 (CNC). Nordegg, July 31, 1921, J. McDunnough, 13, 1♀ (USNM). Nordegg, July 31, 1921, J. McDunnough, 4♀ (CNC). Onefour, June 14, 1952, L. A. Konotopetz, 13 (CNC). Pincher, June 29, 1923, W. Carter, Medicago sativa (Fabaceae) , 23 (CNC). Rycroft, June 21, 1961, A. R. Brooks, 23 (CNC). Valleyview, June 6, 1961, A. R. Brooks, 23, 2♀ (CNC). Waterton Lakes National Park, July 4, 1972, L. A. Kelton, 203, 20♀ (CNC). British Columbia : Aspen Grove, July 10, 1975, L. A. Kelton, 13 (CNC). Atlin, July 4, 6, 7, 1955, B. A. Gibbard, 43, 1♀ (CNC). Buckinghorse River, Alaska Highway, June 24, 1985, S. G. Cannings, 23 (UBC). Cache Creek, 450–500 ft, June 1, 1978, N. L. H. Krauss, 13 (AMNH). Cathedral Provincial Park, Diamond Trail, E slope Scout Mountain, July 26, 1988, S. G. Cannings, 33 (UBC). Field, August 10, 1952, L. A. Konotopetz, 1♀ (CNC). Kaleden, July 2, 1975, L. A. Kelton, 13 (CNC). Kamloops, Paul Lake, June 29, 1950, B. P. Beirne, 33, 2♀ (CNC). Leanchoil, August 5, 1970, L. A. Kelton, 1♀ (CNC). Lumby, August 13, 1975, L. A. Kelton, 1♀ (CNC). Manning Provincial Park, Allison Pass, August 12, 1970, L. A. Kelton, 63, 5♀ (CNC). Manning Provincial Park, Blackwall, July 23, 1970, L. A. Kelton, 13, 3♀ (CNC). Manning Provincial Park, Gibson Pass, August 11, 1970, L. A. Kelton, 13, 1♀ (CNC). McLease Lake, June 28, 1950, B. P. Beirne, 1♀ (CNC). Merritt, 10 mi S of, July 19, 1959, L. A. Kelton, 13, 3♀ (CNC). Mi 392 Alaska Highway, Summit Lake, 4500 ft, June 26, 1959, E. E. MacDougall, 13 (CNC). Mi 496 Alaska Highway, Liard Hot Springs, 1500 ft, July 9, 1959, R. E. Leech, 193, 6♀ (CNC). Monaskie Pass, July 16, 1975, L. A. Kelton, 43 (CNC). Oliver, 7 mi E of, Indian Reservation, July 8, 1959, L. A. Kelton ( Asteraceae ), 23, 2♀ (CNC). Oliver, Vaseaux Lake, May 29, 1959, L. A. Kelton, 13, 1♀ (CNC). Osoyoos, Anarchist Mountain, July 13, 1970, L. A. Kelton, 33, 1♀ (CNC). Osoyoos, Anarchist Mountain, July 20, 1959, L. A. Kelton, 13, 2♀ (CNC). Osoyoos, Anarchist Mountain, July 6, 1959, R. Madge, 53, 10♀ (CNC). Osoyoos, Mount Kobau, July 18, 1996, J. Jarrett, 33, 1♀ (UBC). Osoyoos, Mount Kobau Rd., VK, Msxh pitfall trap K4 B­1, 5, July 20–August 18, 1996, J. Jarrett, 153, 3♀ (UBC). Osoyoos, Mount Kobau, VK, Msxh pitfall trap K4 A­2, 3, August 18–September 28, 1996, J. Jarrett, 33, 1♀ (UBC). Osoyoos, Mount Kobau, July 28– August, 18, 1997, J. Jarrett, 13 (UBC). Osoyoos, Richter Pass, June 28, 1959, L. A. Kelton, 13 (CNC). Osoyoos, Richter Pass, May 24, 1959, R. E. Leech, 13 (CNC). Parson, August 6, 1970, L. A. Kelton, 13, 1♀ (CNC). Pink Mountain, August 17, 1982, L. A. Kelton, 73, 4♀ (CNC). Prince George, July 10, 1947, G. J. Spencer ( Solanaceae ), 23 (CNC). Prophet River Provincial Park, June 24, 1985, E. Bijdemast, S. G. Cannings, 33, 1♀ (UBC). Quesnel: June 3, 1949, G. J. Spencer, alfalfa ( Fabaceae ), 13 (CNC); June 14, 1949, G. J. Spencer, 43, 3♀ (CNC); June 15, 1947, G. J. Spencer, 13 (CNC); June 15, 1949, G. J. Spencer, alfalfa ( Fabaceae ), 13, 1♀ (CNC); June 23, 1945, G. J. Spencer, 13, 1♀ (CNC); June 29, 1948, G. J. Spencer ( Solanaceae ), 13 (CNC); July 7, 1949, R. Stace­Smith, 1♀ (CNC); July 21, 1947, G. J. Spencer, 5♀ (CNC); July 31, 1947, G. J. Spencer, 4♀ (CNC). Soda Creek, June 13, 1950, G. J. Spencer, 33, 1♀ (CNC). Soda Creek, June 24, 1950, G. J. Spencer ( Solanaceae ), 1♀ (CNC). Soda Creek, July 12, 1950, G. J. Spencer, A. morloni 13 (CNC). Stanley, July 26, 1947, G. J. Spencer, 13 (CNC). Tete Jaune Cache, June 8, 1997, G. G. E. Scudder, 1♀ (UBC). Tunkwa Lake, August 15, 1951, D. G. Finlayson, 13 (CNC). Valemont, June 8, 1997, G. G. E. Scudder, 23 (UBC). Yolo National Park, July 16, 1970, L. A. Kelton, 33, 5♀ (CNC). Labrador: Hebron, July 10, 1954 and August 3, 1954, J. F. McAlpine, Ranunculus sp. (Ranunculaceae) , 153, 6♀ (CNC). Tessiujak Bay, Nagvak Fjord, August 15, 1954, J. F. McAlpine, 13 (CNC). Manitoba: Churchill, July 1, 1939, W. J. Brown (CNC). Churchill, July 12, 1952 and August 2, 1952, J. G. Chillcott, 133, 31♀ (CNC). Churchill, July 23, 1936 and August 9, 1937, H. E. McClure, D. G. Denning, 113, 6♀ (USNM). Churchill, July 27, 1936, C. T. Parsons, 1♀ (AMNH). Churchill, August 2, 1937, D. G. Dennig, 13, 1♀ (USNM). Fort Churchill, July 13, 1952, J. G. Chillcott, 53, 5♀ (CNC). Gillam, July 19, 1950, J. F. McAlpine, 13, 3♀ (CNC). Hudson Bay, August 3, 1952, J. G. Chillcott, 43 (CNC). Keld, August 8, 1937, R. H. Beamer, 1♀ (KU). Roblin, 30 mi N of, July 14, 1954, A. R. Brooks, Wallis, 53, 2♀ (CNC). Russell, July 21, 1954, A. R. Brooks, Wallis, 13 (CNC). Warkworth Creek, nr Churchill, July 7, 1952, J. G. Chillcott, 43 (CNC). Northwest Territories: Aklavik, June 18, 1931, Bryant, 1♀ (CNC). Aklavik, July 16, 1931, Bryant, 13 (CNC). Baffin Island, Lake Harbour, August 13, 1935, W. J. Brown, 1♀ (USNM). Geillini Lake, August 19, 1952, J. G. Chillcott, 83, 3♀ (CNC). Inuvik, July 25, 1983, L. A. Kelton, 23, 3♀ (CNC). Nicholson Lake, N end, July 27, 1966, J. G. Chillcott, 1♀ (CNC). Reindeer Depot, Mackenzie Delta, August 11, 1948, J. R. Vockeroth, 53, 4♀ (CNC). Reindeer Depot, Mackenzie Delta, July 7, 1948, W. J. Brown, 13 (CNC). Tuktoyaktuk, 20 mi E of, July 6, 1971, W. R. M. Mason, 113, 2♀ (CNC). Whatever Lake, July 1, 1989, J. Pilny, 13 (CNC). Nunavut: Lake Harbour, Baffin Island, August 3, 1935, L. A. Kelton, 49♀ (CNC). Ontario: Almonte, June 28, 1974, D. J. E. Brown, 23, 3♀ (CNC). Bell’s Corners, June 22, 1952, R. S. Bigelow, 13 (CNC). Calabogie, June 21, 1979, D. J. E. Brown, 63, 9♀ (CNC). Carp, June 22, 1981, D. J. E. Brown, 53, 1♀ (CNC). Flower Station, June 28, 1987, D. J. E. Brown, 1♀ (CNC). Gore Bay, July 10, 1961, G. Brumpton, 23 (CNC). Huntley, July 14, 1960, D. J. E. Brown, 1♀ (CNC). Mer Blue, August 9, 1932, G. S. Walley, 1♀ (CNC). Midland, June 14, 1962, Kelton, Thorpe, 53, 5♀ (CNC). Miller Lake, June 12, 1962, Kelton, Thorpe, 263, 14♀ (CNC). North Bay, June 19, 1963, L. A. Kelton, 123, 14♀ (CNC, USNM). Norway Point, Lake of Bays, July 10, 1922, J. McDunnough, 3♀ (CNC). Pautois Creek on Highway 17, 3 km E of Rt 630, July 1, 1990, M. D. Schwartz, Potentilla argentea (Rosaceae) , 63, 3♀ (CNC). South Gloucester, Blossom Park, Greenbelt, June 9, 1991, M. D. Schwartz, Potentilla argentea (Rosaceae) , 133, 10♀ (CNC, AMNH). Stittsville, June 20, 1962 and July 12, 1961, D. J. E. Brown, 33, 1♀ (CNC). Thessalon, June 15, 1965, W. Gagne, 83, 4♀ (UCB). Tobermory, June 12, 1962, Kelton, Thorpe, 1♀ (CNC). Wawa, July 7, 1961, G. Brumpton, 33, 1♀ (CNC). White Lake bog near White Lake , June 6, 1991, M. D. Schwartz, 23 (CNC). Quebec: Bradore Bay, July 29, 1930, W. J. Brown, 33, 4♀ (CNC). Bradore Bay, August 12, 1930, W. J. Brown, 13 (USNM). Fabre, June 13, 1963, L. A. Kelton, 463, 20♀ (CNC, USNM). Fort Chimo, July 2, 1954, W. R. Richards, 1♀ (CNC). Gatineau National Park, June 11, 1981, D. J. E. Brown, 133, 3♀ (CNC). Great Whale River, July 12, 1949 and August 18, 1949, J. R. Vockeroth, 43, 8♀ (CNC). Indian House Lake, July 6, 1954 and August 4, 1954, W. R. Richard, R. Coyles, 173, 2♀ (CNC). Laniel, June 17, 1963 and July 3, 1963, L. A. Kelton, 253, 10♀ (CNC). Laniel, June 26, 1963, L. A. Kelton, 13, 1♀ (CAS). Near Luskville, June 11, 1991, M. D. Schwartz, Potentilla argentea (Rosaceae) , 13, 3♀ (CNC). Petit­Gaspé beach, Cap­des­Rosiers, Forillon National Park, 5 m, July 20, 1995, M. D. Schwartz, Epilobium angustifolium (Onagraceae) , 13 (CNC). Shawville, July 22, 1958, L. A. Kelton, 43, 8♀ (CNC). Tasiujaq, Karivik, August 20, 1996, B. J. Sinclair, 2♀ (CNC). Yarms, July 23, 1958, L. A. Kelton, 1♀ (CNC). Saskatchewan: Big River, June 5, 1959 and July 3, 1959, A. R. and J. E. Brooks, 53, 2♀ (CNC). Christopher Lake, July 11, 1959, A. R. and J. E. Brooks, 63, 3♀ (CNC). Christopher Lake, July 13, 1959, A. and J. Brooks, 13 (USNM). Fish Creek, July 18, 1925, K. M. King, 23, 1♀ (CNC). Garrick, June 29, 1950, L. A. Konotopetz, Medicago sativa (Fabaceae) , 243, 5♀ (CNC). Good Spirit Lake, July 10, 1954, A. R. Brooks, Wallis, 13, 1♀ (CNC). Love, July 5, 1950, L. A. Konotopetz, Medicago sativa (Fabaceae) , 1♀ (CNC). Melfort, July 16, 1925, K. M. King, 13 (CNC). Preeceville, July 9, 1954, A. R. Brooks, Wallis, 83, 8♀ (CNC). Prince Albert, July 3, 1954, A. R. Brooks, Wallis, 73, 3♀ (CNC). Saskatoon, June 21, 1951, L. A. Konotopetz, 23, 2♀ (CNC). Saskatoon, June 28, 1950, A. R. Brooks, 103, 2♀ (CNC). Saskatoon, near Research Farm and Experimental Plots, July 11, 1990, M. D. Schwartz, 13 (CNC). Stoughton, 35.4 km N of, on Rt 7, July 9, 1990, M. D. Schwartz and R. G. Foottit, Brassica juncea (Brassicaceae) , 1♀ (CNC). Torch River, July 20, 1950, L. A. Konotopetz, 13 (CNC). Torch River Campground, 10.7 mi S of, on Rt 106, July 12, 1990, M. D. Schwartz, Achillea sp. (Asteraceae) , 13, 1♀ (CNC). White Fox , June 20, 1950 and August 7, 1950, L. A. Konotopetz, 143, 7♀ (CNC). Yukon Territory: Carcross, August 1, 1982, L. A. Kelton, Lupinus sp. (Fabaceae) , 43, 3♀ (CNC). Carcross, August 9, 1983, L. A. Kelton, Artemisia frigida (Asteraceae) , 2♀ (CNC). Carmacks, July 17, 1983, L. A. Kelton, Artemisia sp. (Asteraceae) , 4♀ (CNC). Dawson, July 21, 1982, L. A. Kelton, 53, 15♀ (CNC). Dawson, July 27, 1949, W. W. Judd, 23 (CNC). Destruction Bay, July 26, 1982, L. A. Kelton, Hedysarum mackenzii (Fabaceae) , 43, 2♀ (CNC). Destruction Bay, July 26, 1982, L. A. Kelton, Lathyrus sp. (Fabaceae) , 263, 23♀ (CNC). Dislocation Bay, July 25, 1982, L. A. Kelton, 33, 6♀ (CNC). Gravel Lake, 58 mi E of Dawson, 2050 ft, August 11, 1962, P. J. Skisko, Achillea sp. (Asteraceae) , 1♀ (CNC). Haines Junction, July 28, 1982, L. A. Kelton, 123, 20♀ (CNC). Jct Dempster and Klondike Hwy, July 20, 1982, L. A. Kelton, 13 (CNC). Koidern, July 24, 1982, L. A. Kelton (CNC). Mi 465 on Dempster Highway, 800 m, July 5, 1980, D. M. Wood, D. Lafontaine, 53, 4♀ (CNC). Moose Creek, July 18, 1982, L. A. Kelton, 43, 5♀ (CNC). Rampart House, July 19, 1951, J. E. H. Martin, 1♀ (CNC). Rancheria, August 11, 1982, L. A. Kelton, Lathyrus sp. (Fabaceae) , 13 (CNC). Silver City, July 27, 1982, L. A. Kelton, 83, 19♀ (CNC). Tagish, July 17, 1983, August 3, 1982, L. A. Kelton, Astragalus abniginum (Fabaceae) , 313, 26♀ (CNC). Takhini Hot Springs, 2400 ft, August 19, 1962, R. E. Leech, 23, 3♀ (CNC). Tatchun, August 18, 1982, L. A. Kelton, Lathyrus sp. (Fabaceae) , 13, 9♀ (CNC). Von Wilczek Lake, July 18, 1982, L. A. Kelton, 1♀ (CNC). Whitehorse, July 16, 1982, L. A. Kelton, Astragalus sp. (Fabaceae) , 323, 40♀ (CNC). Whitehorse, July 30, 1982, L. A. Kelton, Oxytropis campestris (Fabaceae) , 43, 14♀ (CNC). Whitehorse, August 6, 1974, N. L. H. Krauss, 1♀ (AMNH). DENMARK: Greenland: Sondrestrom Air Base, July 4, 1952, W. J. Brown, 1♀ (USNM). Sondrestrom Air Base, July 4, 1952 and July 30, 1952, W. J. Brown, 57♀ (CNC). USA: Alaska: Bristol Bay Co.: King Salmon, Naknek River, July 13, 1952, W. R. Mason, 13 (CNC). Denali Co.: Mount McKinley National Park, 1730 ft, June 19, 1957, G. W. Byers and party, 143, 16♀ (KU). Kenai Peninsula Co.: Hope, July 15, 1951, W. J. Brown, 13 (CNC). Nome Co.: Unalakleet, July 5, 1961, R. Madge, B. S. Heming, 313, 19♀ (CNC, USNM). South Fairbanks Co.: Big Delta, July 13, 1951, W. R. M. Mason, J. R. McGillis, 33, 1♀ (CNC). Big Delta, July 16, 1948, R. I. Sailer, 133, 2♀ (CNC, USNM). Chicken, July 22, 1982, L. A. Kelton, Aster sp. (Asteraceae) , 33, 9♀ (CNC). Shaw Creek, mi 289 Richardson Highway, July 11, 1951, Mason, McGillis, 1♀ (CNC). Colorado: Archuleta Co.: Pagosa Springs, July 22, 1968, L. A. Kelton, 1♀ (CNC). Eagle Co.: 9.2 mi S of Minturn, White River National Forest, 9080 ft, July 19, 1990, Zolnerowich, 13, 3♀ (TAMU). Jackson Co.: 4 mi N of Gould, August 5, 1975, J. C. Schaffner, 93, 10♀ (TAMU). Larimer Co.: Bennett Creek Picnic Grounds, Pingree Park Road, 46 mi W of Fort Collins, 7400 ft, July 14, 1986, R. T. Schuh and J. T. Polhemus, 23 (AMNH). Fish Creek Picnic Grounds, Pingree Park Road, 46 mi W of Fort Collins, 7700 ft, July 14, 1986, R. T. Schuh and J. T. Polhemus, 13, 2♀ (AMNH). Pingree Park, August 23, 1924, C. J. Drake, 1♀ (USNM). Pingree Park, August 25, 1935, Beamer, Lawson, 33 (CNC). Mesa Co. : Grand Mesa, Carson Lake , August 6, 1993, J. T. Polhemus, 1♀ (JTP). Routt Co.: Meadows, Routt National Forest, August 20, 1968, L. A. Kelton, Ranunculus sp. (Ranunculaceae) , 203, 15♀ (CNC). Steamboat Springs, August 22, 1968, L. A. Kelton, 1♀ (CNC). Idaho: Blaine Co.: Ketchum, July 30, 1955, B. P. Bliven, 1♀ (CAS). Montana: Beaverhead Co.: 5 mi E of Wise River, July 21, 1983, J. D. Pinto, 13 (UCR). Glacier Co.: 6 mi N of East Glacier Park on Rt 49, Blackfeet Indian Reservation, 5400 ft, August 3, 1994, M. D. Schwartz, 33 (CNC). Park Co.: 2 mi E of Cooke City on Rt 212, Soda Butte Campground, Gallatin National Forest, 7700 ft, August 11, 1986, Schuh, Schwartz, Stonedahl, 13 (AMNH). Utah: Uintah Co.: Oaks Park, August 2, 1965, G. F. Knowlton, 23, 4♀ (USU). Wyoming: Big Horn Co.: 24 mi W of Burgess Jct on road to radar station, 9000–9200 ft, August 12, 1986, Schuh, Schwartz, Stonedahl, Lupinus argenteus (Fabaceae) , 803, 75♀ (AMNH, USNM). Park Co.: E of Beartooth Pass on Hwy 212, 10,800 ft, August 1, 1994, M. D. Schwartz, Achillea millefolium (Asteraceae) , 123, 8♀ (CNC, AMNH). Sheridan Co.: Prune Creek, 42 mi W of Sheridan, at Prune Creek Campground, 7400 ft, July 25, 1978, N. L. Herman, 13 (AMNH).

Chlamydatus monilipes Van Duzee

Figures 1 , 3I , 5

Chlamydatus monilipes Van Duzee, 1921: 132 (n.sp.).

DIAGNOSIS: Recognized by the small size, black coloration of body and femora, and the nearly white background coloration of tibiae with strongly contrasting black spots at the bases of the tibial spines. Most similar in overall coloration to C. brevicornis , C. keltoni , and C. suavis ; distinguished from C. brevicornis by the conspicuously shining vestiture of the dorsum (fig. 1) and the bases of the tibial spines being pale in that species (fig. 3G); distinguished from C. keltoni by the brown bases of the tibial spines (fig. 3H) and convoluted vesica with a rather long apical spine (fig. 4) in that species as opposed to the heavy black tibial spines on a stark white background (fig. 3I) and the shorter vesica with a relatively short apical spine in C. monilipes (fig. 5); distinguished from C. suavis by generally brown coloration and faint brown bases of the tibial spines in that species (fig. 3G) as opposed to the nearly black coloration and heavy black bases of the tibial spines in C. monilipes (fig. 3I); coloration similar to C. pulicarius with femora almost totally dark, but that species always much larger, total length always more than 2.51 and with a long convoluted vesica with a very long apical process (fig. 5), whereas total length in C. monilipes never more than 2.60 and vesica relatively short with a short apical process (fig. 5).

HOSTS: Most frequently collected on Chrysothamnus spp. and Haplopappus spp. (Asteraceae) .

DISTRIBUTION: Restricted to western North America , ranging from Montana in the north to northern Baja California in the south .

DISCUSSION: The presumed holotype of C. monilipes is deposited in the California Academy of Sciences, San Francisco as type No. 3574. We have not examined it as the identity of this species seems not to be in question based on our examination of specimens identified by Van Duzee .

SPECIMENS EXAMINED: MEXICO: Baja California Norte: 8 mi E of Tecate, Highway 2, May 13, 1980, Brown and Faulkner, 13 (SDNH). junction Highway 3, Ojos Negros, April 4, 1981, Brown and Faulkner, 23 (SDNH). Baja California Sur : 3 mi N of San Jose Viejo, July 16, 1971, H. G. Real and R. E. Main, 13 (CAS). USA: Arizona: Gila Co. : 2 mi W of Miami, 3800 ft, May 4, 1979, E. M. Fisher, J. D. Pinto, 13 (UCR). Graham Co.: San Carlos Indian Reservation, April 16, 1969, D. and J. Schuh, 53 (AMNH). San Carlos Reservoir, S side of, April 17, 1969, D. and J. Schuh, 73 (AMNH). Maricopa Co. : Fort McDowell, March 15, 1982, J. T. Polhemus, 13 (JTP). Wickenburg, 4 mi SW of, on Vulture Mine Road, 700 m, April 1, 1981, R. T. Schuh and M. D. Schwartz, Haplopappus sp. (Asteraceae) , 1103 (AMNH). Wickenburg, 6 mi W of, April 2, 1966, Joe Schuh, Astragalus (Fabaceae) , 13 (AMNH). Pima Co. : Santa Catalina Mountains , 2 mi N of Molino Canyon Campground, April 10, 1989, T. J. Henry and A. G. Wheeler, Jr.,? Hypericum sp. (Saxifragaceae) , 13, 3♀ (USNM). Tucson, Santa Catalina Mts., Bear Creek near Saddleback Drive, February 26, 1995, M. D. Schwartz, 33 (CNC). Tucson, wash nr. 8712 Saddleback Dr., February 26, 1995, M. D. Schwartz, Ambrosia ambrosioides (Asteraceae) , 103 (CNC). Pinal Co.: Oracle, 14 mi E of, July 27, 1924, E. P. Van Duzee, 213 (CAS). Superior, 4 mi E of, 4000 ft, May 5, 1979, E. M. Fisher ( Fagaceae ), 13 (UCR). Santa Cruz Co. : Nogales, October 15, 1906, Koebele, 13 (CAS). California: El Dorado Co. : Lake Tahoe, lakeside, July 27, 1930, H. H. Keifer, Chrysothamnus nauseosus (Asteraceae) , 23, 2♀ (CAFA, CAS, LACM). Fresno Co.: Coalinga, April 16, 1980, R. S. Miller, 13 (TAMU). Jacolitos Canyon, April 8, 1966, R. O. Schuster, 13 (UCB). Inyo Co. : Alabama Hills, May 12, 1979, L. D. French, 13 (UCD). Big Pine, 12 mi E of, June 9, 1966, W. Gagne, 63 (UCB). Darwin, May 12, 1969, J. Doyen, 13 (UCB). Mesquite Spring Campground, Death Valley National Monument, 1760 ft, March 24, 1982, L. Russell, Mentzelia sp. (Loasaceae) , 73 (OSU). Plauche, May 20, 1937, N. W. Frazier, 13 (UCB). Saline Valley Rd., 26 mi N of junction with Hwy. 190, April 21, 1975, J. D. Pinto, Amphipappas fremontii (Asteraceae) , 43 (UCR). Westgard Pass summit, White Mountains, 2250 m, July 2, 1980, R. T. Schuh, Chrysothamnus sp. (Asteraceae) , 123 (AMNH). Wildrose Station, May 14, 1969, J. Doyen, 43 (UCB). Kern Co.: Antelope Canyon, Tehachapi Mts. , 4800–5200 ft, May 15, 1976, J. D. Pinto, 13 (UCR). Freeman Junction on Rt 178 and 14, 21 mi N of Mojave, April 24, 1980, Russell and Schwartz ( Asteraceae ), 183 (OSU). McKittrick, April 9, 1966, L. and C. W. O’Brien, 23 (UCB). Mojave, May 3, 1918, 23, 5♀ (CNC). Mojave, May 3, 1919, 23 (CNC). Red Rock Canyon, May 2, 1968, J. Doyen, 263 (UCB). Red Rock Canyon, May 18, 1937, E. P. Van Duzee, 63 (CAS). Red Rock Canyon, 4 mi N of, May 1, 1968, J. Powell, 163 (UCB). Shadow Mountain, May 27, 1941, G. H. and J. L. Perry, 13 (AMNH). Lassen Co.: Madeline, July 24, 1954, B. P. Bliven, 13 (CAS). Sagehen, July 26, 1954, B. P. Bliven, 13 (CAS). Susanville, June 17, 1959, L. A. Kelton, R. Madge, 273, 17♀ (CNC). Los Angeles Co.: Lancaster, 3 mi S of, March 22, 1971, P. Oman, 153 (OSU). Lovejoy Butte, Mojave Desert, May 10, 1944, A. L. Melander, 33 (UCR). Mint Canyon, 5 mi E of Saugus, April 20, 1932, E. P. Van Duzee, 413 (CAS). Mint Canyon, May 16, 1937, E. P. Van Duzee, 213 (CAS). Neenach, May 17, 1928, J. O. Martin, 73 (CAS). Palmdale, June 17, 1937, E. P. Van Duzee, 83 (CAS). Palmdale, 5 mi W of, April 14, 1932, E. P. Van Duzee, 93 (CAS). Pearblossom, May 15, 1955, W. E. Simonds, 33 (CAFA). Solemint, April 28, 1955, W. R. M. Mason, 113, 5♀ (CNC). Valyermo, May 1, 1968, J. Powell, 33 (UCB). Modoc Co.: 2.5 mi S of Rt 139 towards Lookout, 1440 m, July 6, 1979, R. T. and J. Schuh, Chrysothamnus nauseosus (Asteraceae) , 153 (AMNH). 8 mi S of Rt 139 towards Lookout, 1420 m, July 6, 1979, R. T. and J. Schuh, Chrysothamnus nauseosus (Asteraceae) , 103 (AMNH). Altura­Likely, June 20, 1956, J. Schuh, 33, 2♀ (CNC). Canby, 24.7 mi NW of, 1375 m, July 1, 1979, R. T. Schuh and B. M. Massie, 13 (AMNH). Cedar Pass, W of Cedarville, 1850 m, July 1, 1979, R. T. Schuh and B. M. Massie, Chrysothamnus nauseosus (Asteraceae) , 303 (AMNH). Hackamore, July 12, 1947, R. L. Usinger, 163 (UCB). Mono Co.: Benton Hot Springs, June 8, 1966, W. Gagne, 63 (UCB). Coleville, 5200 ft, June 10, 1966, W. Gagne, 13 (UCB). Crooked Creek Naval Res. Station, White Mts. , 10,000 ft, August 8, 1966, L. and C. W. O’Brien, 13 (UCB). No specific locality, August 3, 1939, W. Dwight Pierce, 53 (LACM). Sardine Creek, 8500 ft, July 6, 1951, A. T. McClay, 13 (UCD). Nevada Co. : Boca, June 18, 1974, R. D. Moon, 23 (UCD). Boca, June 25, 1976, R. M. Giblin, 23 (UCD). Boca, July 23, 1970, R. M. Bohart, 13 (UCD). Sagehen Creek Station near Hobart Mills, August 20, 1969, W. Gagne, Artemisia sp. (Asteraceae) , 13 (UCB). Truckee, July 5, 1927, E. P. Van Duzee, 13 (CAS). Orange Co.: Cleveland National Forest, 1.5 mi E of San Juan Campground, May 12, 1978, J. D. Pinto and R. T. Schuh, 63 (AMNH). Irvine, April 5, 1972, Lilly and Enns ( Asteraceae ), 13 (CAFA). Plumas Co.: Chester, 8 mi NW of, August 17, 1968, J. Powell, 13 (UCB). Near Chester, July 7, 1932, R. L. Usinger, 473 (UCB). Spring Garden, June 23, 1937, B. P. Bliven, 13 (CAS). Riverside Co.: Coyote Canyon, 3.5 mi S of Palm Desert, Boyd Desert Research Center, May 17, 1975, J. B. Tucker, 13 (UCR). Lake Elsinore, April 21, 1920, E. P. Van Duzee, 13 (CAS). Maze Stone County Park near Hemet, April 10, 1974, J. D. Pinto, 23 (UCR). Menifee Valley, hills on W end, 1800 ft, May 11, 1978, J. D. Pinto and R. T. Schuh, Salvia apiana (Lamiaceae) , 13 (AMNH). Near Menifee Valley, April 19, 1974, J. D. Pinto, 13 (UCR). Quail Valley, 8 mi NE of Elsinore, April 11, 1965, J. Powell, 13 (UCB). Thousand Palms, April 27, 1955, W. R. Richards, 93, 6♀ (CNC). White Water , March 19, 1955, W. R. Richards, 23 (CNC). Wilson Valley Rd., 1 mi N of Hemet, April 21, 1980, Russell and Schwartz, Salvia sp. (Lamiaceae) , 13 (AMNH). San Bernardino Co.: Apple Valley, May 19, 1955, W. R. Richards, 163, 7♀ (CNC). Barstow, 26.7 mi S of, on St Rt 247, 1000 m, May 2, 1985, R. T. Schuh and B. M. Massie, Lycium cooperi (Solanaceae) , 93 (AMNH). Cajon Pass Summit, 1.5 mi N of, on I­ 15, 1310 m, May 2, 1985, R. T. Schuh and B. M. Massie, Haplopappus linearifolius (Asteraceae) , 323 (AMNH). Granite Pass, 3800–4320 ft, April 27, 1968, C. Beesley, Larrea divaricata (Zygophyllaceae) , 13 (UCR). Joshua Tree National Monument, entrance, April 20, 1960, Gertsch, Ivie, and Schrammel, 13 (AMNH). Kramer Hills, Boron, May 10, 1955, W. R. M. Mason, 33 (CNC). Kramer Junction, 1.5 mi W of, April 24, 1980, M. D. Schwartz and L. K. Russell, unidentified sp. ( Asteraceae ), 403 (AMNH). Lucerne Valley, 5.4 mi W of, 935 m, May 13, 1978, J. D. Pinto and R. T. Schuh, Chrysothamnus nauseosus (Asteraceae) , 63 (AMNH, UCR). March’s Ranch Road, 2 mi NE Baldwin Lake, May 25, 1968, S. Freeman, Haplopappus sp. (Asteraceae) , 13 (UCR). Old Woman Springs Rd, 6.3 mi N of Yucca Valley, May 13, 1978, J. D. Pinto, 63 (UCR). Piñon Hills T4N, R7W, Sec 7, May 22, 1979, H. L. Murray, R. A. Sherman, 133 (UCR). Rialto, May 13, 1956, W. E. Simonds, 13 (CAFA). Victorville, April 21, 1935, 23 (LACM, UCD). Yucca Valley, 6.3 mi N of, 1125 m, May 13, 1978, J. D. Pinto and R. T. Schuh, Dalea sp. (Fabaceae) , 103 (AMNH). Yucca Valley, 10 mi N of, May 28, 1975, J. D. Pinto, Acamptopappus sphaerocephalus (Asteraceae) , 13 (UCR). San Diego Co.: Carrista Creek, Morettis Junction, May 13, 1980, Brown and Faulkner, 23 (SDNH). Palm Canyon Campground, Anza­Borrego State Park, 592 ft, May 16, 1982, M. D. Schwartz, 13 (AMNH). No specific locality, April 22, 1913, E. P. Van Duzee, 23 (CAS). Warner Springs, 7 mi S of, Carrizo Creek intersection w/ Rt 79, May 16, 1982, M. D. Schwartz, Gutierrezia sarothrae (Asteraceae) , 143 (AMNH). Warner Springs, 20 mi S of, April 5, 1967, D. M. Wood, 33, 4♀ (CNC). San Luis Obispo Co.: near Cuyama Rd., 1 mi W of Cottonwood Cyn. on Rt 166, April 20, 1980, Russell and Schwartz, Chrysothamnus nauseosus (Asteraceae) , 353 (AMNH). Tembler Range, 12 mi E of Simmler, 3200 ft, April 25, 1964, W. Turner, 13 (UCB). Santa Cruz Co. : Mount Hermon, July 15, 1922, F. E. Blaisdon, 13 (CAS). Shasta Co.: Lake Britton, June 29, 1947, T. F. Leigh, 13 (UCB). Sierra Co. : S of Sierraville, June 18, 1966, L. and C. W. O’Brien, at night, 13 (UCB). Siskiyou Co.: 12.3 mi N of St. Hwy. 89 on Pow­ der Hill Road, July 19, 1985, G. M. Stonedahl and J. D. McIver, Chrysothamnus nauseosus (Asteraceae) , 253 (AMNH). Hornbrook, April 14, 1959, L. A. Kelton, R. Madge, Artemisia sp. (Asteraceae) , 1♀ (CNC). Lava Beds National Monument, August 13, 1961, Joe Schuh, 13 (OSU). Lava Beds National Monument, just S of, on Medicine Lake Road, 1625 m, June 26, 1979, R. T. and J. Schuh, Haplopappus sp. (Asteraceae) , 603 (AMNH). Mammoth Crater, 0.3 mi SE of, Klamath National Forest, 5200 ft, June 26, 1979, M. D. Schwartz, Haplopappus sp. (Asteraceae) , 263 (AMNH). McCloud, July 16, 1974, Shaw, Ruddiman, 93 (CAFA). Medicine Lake, 2.5 mi N of, on Medicine Lake Rd, July 18, 1985, G. M. Stonedahl and J. D. McIver, Ribes cereum (Grossulariaceae) , 43 (AMNH). Medicine Lake Road, 5200 ft, June 26, 1979, G. Stonedahl, J.D. Lattin, 403 (OSU). Mount Shasta City, June 30, 1937, B. P. Bliven, 13 (CAS). Shasta City exit on Interstate 5, May 22, 1981, J. D. Lattin, 113 (OSU). Stanislaus Co.: Patterson, 74E14­67, May 2, 1979, Bingham, Paddock, Carlson, Stenotopsis sp. (Asteraceae) 233 (CAFA). Tulare Co.: 21 mi W of Rt 395 toward Kennedy Meadows, 1875 m, July 1, 1980, R. T. Schuh, Chrysothamnus parryi (Asteraceae) , 173 (AMNH). Colorado: Jackson Co.: 3 mi N of Gould, August 13, 1968, P. Oman, 13 (OSU). Idaho: Caribou Co.: 11 mi E of Way­ an on Rt 34, Pine Bar Campground, 6000 ft, July 30, 1981, M. D. Schwartz, Purshia tridentata (Rosaceae) , 13 (AMNH). Oneida Co. : 6 mi NW of Holbrook, June 9, 1969, G. F. Knowlton, 23 (USU). Holbrook, June 27, 1969, G. F. Knowlton, 13 (USU). Montana: Glacier Co.: 15 mi W of Babb, Glacier National Park, August 8, 1969, Oman, 13 (OSU). Nevada: Carson City Co. : Carson City, June 20, 1929, E. P. Van Duzee, 23 (CAS). Elko Co.: Elko, 15 mi SE of, on Rt 227, 5600 ft, June 26, 1983, R. T. Schuh and M. D. Schwartz, Lupinus sp. (Fabaceae) , 23 (AMNH). Elko, 30 mi W of, June 18, 1982, T. J. Henry, Artemisia sp. (Asteraceae) , 23 (USNM). Esmeralda Co. : Goldfield, 6 mi N of, 5300 ft, June 8, 1966, W. Gagne, 113 (UCB). Lida, 2 mi W of, 6500 ft, June 8, 1966, W. Gagne, 63 (UCB). Lida, 3 mi W of, 6800 ft, June 7, 1966, W. Gagne, 63 (UCB). Eureka Co.: 28 mi W of Eureka on Rt 50, 6000 ft, June 27, 1983, R. T. Schuh and M. D. Schwartz, Gutierrezia sarothrae (Asteraceae) , 253 (AMNH). Lincoln Co.: 11.7 mi NW of Oak Spring Summit on Rt 93, 5000 ft, M. D. Schwartz, Tetradymia canescens (Asteraceae) , 13 (AMNH). Lyon Co.: 7 mi N of Sweetwater Summit on Rt 22, Toiyabe National Forest, 1865 m, July 11, 1980, R. T. Schuh and G. M. Stonedahl, Chrysothamnus nauseosus (Asteraceae) , 13 (AMNH). Mineral Co.: 27 mi SW of Hawthorne on Rt 359, 7400 ft, July 2, 1983, R. T. Schuh and M. D. Schwartz, Chrysothamnus viscidiflorus (Asteraceae) , 53 (AMNH). Nye Co.: Atomic Test Site, 1.5 mi W Area 12 camp on Stockade Wash Rd, June 7, 1983, Schuh, Schwartz, Stonedahl, Chrysothamnus nauseosus (Asteraceae) , 553 (AMNH). Atomic Test Site, 2.6 mi W of Mercury Hwy, Cane Springs Rd, 3400 ft, June 6, 1983, Schuh, Schwartz, Stonedahl, Chrysothamnus viscidiflorus (Asteraceae) , 103 (AMNH). Atomic Test Site, 4.5 mi S GS500 on Jackass Flats Rd, 3300 ft, June 6, 1983, Schuh, Schwartz, Stonedahl ( Brassicaceae ), 23 (AMNH). Belmont, 7 mi S of, T8N, R45E, 6800 ft, June 29, 1983, R. T. Schuh and M. D. Schwartz, Gutierrezia sarothrae (Asteraceae) , 253 (AMNH). Belmont, 11 mi N of, T10N, R46E, sec. 26, 7200 ft, June 29, 1983, R. T. Schuh and M. D. Schwartz, Gutierrezia sarothrae (Asteraceae) , 83 (AMNH). Mercury, 19 M, July 23, 1965, H. H. Knight and J. Merino, 23, 1♀ (CNC, USNM). Northumberland Canyon Rd., Toquima Mts., 6400 ft, June 28, 1983, R. T. Schuh and M. D. Schwartz, Gutierrezia sarothrae (Asteraceae) , 13 (AMNH). Oregon: Deschutes Co.: Brothers, 8 mi SE of, July 9, 1968, Lattin, 23 (OSU). Millican, 0.5 mi S of, June 29, 1978, J. D. Lattin, 403 (OSU). Millican, 1 mi S of, 4380 ft, June 21, 1979: G. M. Cooper, 193 (OSU); G. M. Stonedahl, 103 (OSU); M. D. Schwartz, Chrysothamnus nauseosus (Asteraceae) , 153 (AMNH); R. T. Schuh, Chrysothamnus viscidiflorus (Asteraceae) , 1303 (AMNH). Millican, 2.2 mi SW of, 4520 ft, June 29, 1978, Lattin and Oman, 13 (OSU). Millican Sand Dunes, 3 mi W of, July 6, 1977, J. D. Lattin, 123 (OSU). Harney Co.: Burns, June 24, 1954, G. F. Knowlton, 13 (KU). Jackson Co.: Mount Ashland, 1 mi below summit, 6500 ft, September 24, 1968, J. D. Lattin, 13 (OSU). Klamath Co.: Bly Mount Summit, 25 mi E of Klamath Falls, June 18, 1952, V. Roth, 13 (OSU). Chemult, 8 mi S of, August 7, 1941, J. Schuh, 1♀ (CNC). Hayden Mountain Summit, W of Keno, 1475 m, June 27, 1979, R. T. and J. Schuh, 23 (AMNH). Near Pothole, July 25, 1958, Joe Schuh, Haplopappus sp. (Asteraceae) , 13 (OSU). R8E T30S Section 9, July 13, 1978, P. Oman, 13 (OSU).

Chlamydatus montanus Knight

Figures 1 , 3D , 5

Figures

Fig. 1. Habitus views of Chlamydatus spp.: C. artemisiae–C. opacus. See appendix 1 for localities.

Fig. 3. Coloration patterns on legs of Chlamydatus spp.

Fig. 4. Male genitalia of Chlamydatus spp.: C. artemisiae–C. keltoni. See appendix 1 for localities.

Fig. 8. Scanning electron micrographs of Chlamydatus keltoni, 3, British Columbia: Yoho National Park. A. Head, lateral view. B. Setae on hemelytra. C. Pretarsus, apical ventral view. D. Pretarsus, lateral view. E. Mesothoracic spiracle and metaepisternal scent efferent system, lateral view. F. Pygophore, posterodorsal view, showing left paramere and phallotheca in repose. Scales as indicated.

Fig. 5. Male genitalia of Chlamydatus spp.: C. monilipes–C. pulicarius. See appendix 1 for localities.