Paraglypturus Türkay and Sakai, 1995

Poore, Gary C. B., Dworschak, Peter C., Robles, Rafael, Mantelatto, Fernando L. & Felder, Darryl L., 2019, A new classification of Callianassidae and related families (Crustacea: Decapoda: Axiidea) derived from a molecular phylogeny with morphological support, Memoirs of Museum Victoria (Mem. Mus. Vic.) 78, pp. 73-146 : 128

Paraglypturus Türkay and Sakai, 1995

Paraglypturus Türkay and Sakai, 1995: 26–27 .— Sakai, 1999a: 122.— Sakai, 2005b: 204–205.— Sakai, 2011: 504–505.— Sakai, 2018: 744.

Type species. Paraglypturus calderus Türkay and Sakai, 1995 , by original designation and monotypy.

Diagnosis. Maxilliped 3 ischium and merus more than twice as long as wide at their articulation; ischium with strong proximal lobe on lower margin, wider proximally than distally. Male major cheliped with palm about twice as long as carpus, 1.5 times as long as wide, minor chela slightly smaller, with elongate fingers. Male pleopod 1 article 1 with distal long setae, article 2 blade like, with bifid apex, sometimes obscure; with appendix interna (at least with hooks). Male pleopod 2 appendix interna digitiform; appendix masculina a lobe fused with appendix interna, subdistal on endopodal margin. Telson widest anteriorly, more or less semicircular.

Remarks. Paraglypturus differs from other Eucalliacidae in having the pleopod 2 endopod almost as wide as that of pleopod 3 (cf. figs 19u, z’’). The second article of the pleopod 1 of the male has a minute appendix interna with hooks (fig. 19s). The species share with many eucalliacids two long setae distally on the upper margin of the propodus of pereopod 4 (fig. 18m) but differ in the arrangement of the appendices on the male pleopod 2. The chelipeds are swollen and similar, the minor slightly smaller (figs 18g, i).

Robles et al. (in press) found weak molecular support for one of its two species to be more closely related to Callianopsidae than to any other callianassoids despite the morphological evidence supporting eucalliacid affinities ( Sakai (2005b, 2011, 2018). Paraglypturus was said by Türkay and Sakai (1995) to be close to Glypturus but this was realised to be wrong when Sakai (1999a) noted its similarity to Calliax and placed both genera in Eucalliacinae . He differentiated the two on the possession in Paraglypturus (in which he included five species) and absence in Calliax (seven species) of an exopod on maxilliped 3. Later, Sakai (2005b, 2011) restricted the genus to its type species because it alone possesses the “yellow transparent circular structure on the uropodal endopod”. Türkay and Sakai (1995) described and figured a “rounded yellow-transparent circular structure centrally” on the upper surface of the uropodal exopod of P. calderus . This was confirmed by PCD on a paratype ( SMF 22951) but subsequent examination by GCBP in 2008 and in 2018 of another specimen ( MNHN Th1416) failed to distinguish the structure. Ahn et al. (2017) figured in colour a yellow patch on the upper surface of the uropodal endopod of P. tonganus .