Novispathodus ex gr. waageni ( Sweet, 1970 ), 2022

Leu, Marc, Bucher, Hugo, Vennemann, Torsten, Bagherpour, Borhan, Ji, Cheng, Brosse, Morgane & Goudemand, Nicolas, 2022, A Unitary Association-based conodont biozonation of the Smithian-Spathian boundary (Early Triassic) and associated biotic crisis from South China, Swiss Journal of Palaeontology (19) 141 (1), pp. 1-61 : 37-38

Novispathodus ex gr. waageni ( Sweet, 1970)

Fig. 14C, D, F, G

*1970 Neospathodus waageni n. sp.; Sweet, pp. 260–261, pl. 1, figs. 11, 12.

1977 Neospathodus waageni Sweet ; Goel, p. 1094, pl. 2, figs. 1–4.

1978 Neospathodus waageni Sweet ; Weitschat & Lehmann, pl. 14, figs. 11–12.

1979 Neospathodus waageni Sweet ; Solien, p. 292, pl. 3, fig. 9.

1980 Neospathodus waageni Sweet ; Chhabra & Sahni, pl. 1, figs. 9–10, 14?, 16, 20?.

1982 Neospathodus waageni Sweet ; Koike, p. 39, pl. 6, figs. 24–27.

1983 Neospathodus waageni Sweet ; Matsuda, p. 88–91, pl. 1, figs. 6–10.

1984 Neospathodus waageni Sweet ; Berry et al., p. 133, pl. 1, figs. 1–4.

1984 Neospathodus waageni Sweet ; Dagis, p. 24, pl. 7, figs. 2–5, 7–10, pl. 8. fig. 7 (only).

2004 Neospathodus waageni Sweet ; Zhao et al., figs. 1, 3.

2007 Neospathodus waageni Sweet ; Zhao & Orchard in Zhao et al., pp. 36–37, pl. 1, figs. 5A, B, 10A, B.

2007 Neospathodus ex gr. waageni Sweet ; Orchard & Krystyn, plate, figs. 8–18.

2008 Neospathodus waageni Sweet ; Nakrem et al., figs. 5.7, 5.8, 5.11, 5.14.

2008 Neospathodus waageni Sweet ; Orchard, p. 406, pl. 8, figs. 8.1, 8.2, 8.8, 8.9.

2009 Novispathodus waageni (Sweet) ; Orchard & Zonneveld, p. 785, figs. 13.1–13.10, 14, 15.

2009 Neospathodus ex gr. waageni Sweet ; Igo in Shigeta et al., p. 194, figs. 152.1, 152.3, 152.14–15?, 152.1618, 152.19?, 153.8–9, 156.9, 156.14–19.

2010 Novispathodus waageni (Sweet) ; Beranek et al., figs. 6.22–23.

2012a Novispathodus waageni (Sweet) ; Goudemand & Orchard in Goudemand et al., p. 1031, figs. 3C?, D, E, H, N, S.

2013 Novispathodus waageni (Sweet) ; Zhao et al., figs. 9CC, 10A-H, 11F?, 11P-R?.

2014 Novispathodus waageni n. subsp. A; Goudemand, figs. 1A–1D

2014 Novispathodus ex gr. waageni (Sweet) ; Maekawa & Igo in Shigeta et al., p. 244, figs. 174.31–174.57, 175– 178, 179.1–179.3, 179.7–179.12, 179.16–179.30, 180, 181.1–181.27.

2015 Novispathodus waageni (Sweet) ; Chen et al., figs. 6.23, 7.11, 8.3, 8.8, 8.9, 8.11, 8.14.

2016 Novispathodus waageni (Sweet) ; Liang et al., fig. 4.8.

2018 Novispathodus ex gr. waageni (Sweet) ; Maekawa in Maekawa et al., figs. 22.3, 22.10–11, 22.13?, 23.14– 15, 23.18?.

2018 Novispathodus waageni (Sweet) ; Lyu et al., figs. 5, 6.

2019 Novispathodus waageni (Sweet) ; Lyu et al., figs. 7.8–10.

2019 Novispathodus waageni (Sweet) ; Souquet & Goudemand, figs. 3, 4a, 4g- 4t.

2019 Novispathodus aff. waageni (Sweet) ; Souquet & Goudemand, figs. 4b– 4f.

Number of specimens.> 50.

Diagnosis. See Sweet (1970).

Remarks. Despite the exclusion of several former morphotypes which have been formally described as separate species, this common species still encompasses a lot of variation in the P 1 element and may deserve further differentiation in the future. Up to six morphotypes have been recognized by distinct authors, e.g. Zhao et al. (2004) or Orchard and Krystyn (2007), and several others later by Goudemand (unpublished Ph.D. thesis, 2011). It is not yet clear whether these various morphotypes may be of any utility and hence whether some of those may deserve assignment to separate species (see for instance Lyu et al., 2018 for a discussion on the value of N. waageni eowaageni ).

Occurrence. N. waageni is a very common species in Smithian rocks worldwide. Its FAD has been proposed as a proxy for defining the base of the Olenekian (see discussions in e.g. Goudemand 2014; Lyu et al., 2018; Orchard, 2007, 2010; Orchard & Krystyn, 2007; Zhao et al., 2007; Shigeta, 2009).

Figures

Fig. 14 Neogondolellinae, Novispathodinae and uncertain from Shanggang. Magnification is × 80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise.A, Eurygnathodus costatus (Staesche); SHA325, PIMUZ 39130. B Eurygnathodus hamadai (Koike); SHA326, PIMUZ 39131. C, D, F, G Novispathodus ex gr. waageni (Sweet); C SHA328,PIMUZ 39238; D SHA328, PIMUZ 39239; F SHA325, PIMUZ 39240;G SHA326, PIMUZ 39241.E Neospathodus ex gr. cristagalli (Huckriede); SHA330, PIMUZ 39182.H Neospathodus dieneri (Sweet); SHA338, PIMUZ 39181