Homoneura beckeri

Shi, Li & Yang, Ding, 2009, Notes on the Homoneura (Homoneura) beckeri group from the Oriental Region, with descriptions of ten new species from China (Diptera: Lauxaniidae), Zootaxa 2325, pp. 1-28 : 2-3

publication ID	https://doi.org/ 10.5281/zenodo.192037
DOI	https://doi.org/10.5281/zenodo.6223903
persistent identifier	https://treatment.plazi.org/id/038A8794-2246-7B54-FF6B-FDFCFA23FCD6
treatment provided by	Plazi
scientific name	Homoneura beckeri
status

Treatment

Key to all species of the beckeri View in CoL group in Oriental Region

(females of H. (H.) columnaria , H. (H.) convergens , H. (H.) nigritarsis , H. (H.) yinggeling , and H. (H.) zhangae are unknown)

1. Abdominal tergites 1–5 entirely black or at least tergites 1–2 entirely black; tarsomere 5 not black .......................... 2

- Abdominal tergites 1–5 mostly yellow each with a brown triangular median spot and a narrow transverse band on hind margin; tarsomere 5 at least black or blackish brown on apical 1/3 .................................................................... 4

2. Upper half of occiput with a large brown spot on each side, extending to top margin of eye; abdominal tergites 1–2 entirely black and tergite 3 mostly black except narrow yellow anterior margin, tergites 4–5 black on posterior 1/2 and yellow on anterior 1/2; aedeagus with a pair of falciform dorsal sclerites in ventral view ( Fig. 45 View FIGURES 42 – 46 ) ...................... ............................................................................................................................................. H. View in CoL (H.) occipitalis Malloch

- Upper half of occiput usually without large brown spot on each side, but if brown spot present it does not extend to top margin of eye; abdominal tergites 1–5 entirely deep black; aedeagus not as above .............................................. 3

3. Face mostly black except a yellow spot near ventral margin; abdominal tergite 6 black without spot; aedeagus acuate apically without lateral processes in ventral view ......................................................... H. View in CoL (H.) bomiensis Gao & Yang

- Face yellow with a pair of black spots; abdominal tergite 6 yellow with distinct brown median spot; aedeagus blunt with a pair of acuate lateral processes in ventral view................................................ H. (H.) immaculata View in CoL (de Meijere)

4. Face with brown stripe or spot...................................................................................................................................... 5

- Face without stripe or spot............................................................................................................................................ 8

5. Face with one transverse band or two horizontal spots on ventral 1/3 or near ventral margin ....................................6

- Face with a pair of large brown elliptical median spots (sometimes two spots slightly confluent), occupying majority of face............................................................................................................................................................................ 7

6. Antennal 1st flagellomere yellow except dark at tip; hypandrium slightly concave at middle; aedeagus with many apical teeth and a pair of acuate lateral processes in ventral view.......................................... H. (H.) beckeri (Kertész) View in CoL

- Antennal 1st flagellomere entirely black; hypandrium projecting at middle; aedeagus with a few apical teeth but no acuate lateral processes in ventral view ........................................................... H. (H.) nigrantennata Yang, Hu & Zhu

7. Abdominal tergite 5 with a triangular median spot confluent with a blackish brown transverse posterior band; protandrium without ventral process ( Fig. 19 View FIGURES 17 – 21 ); aedeagus with a pair of acuate lateral processes and a pair of furcated dorsoapical processes constricted subapically in ventral view ( Fig. 20 View FIGURES 17 – 21 ) .......................... H. View in CoL (H.) columnaria sp. nov.

- Abdominal tergite 5 with a triangular median spot but no blackish brown transverse band; protandrium with a ventral process ( Fig. 24 View FIGURES 22 – 26 ); aedeagus consisting of a basal anterior sclerite with a pair of triangular lateral processes, a pair of basally furcated dorsal sclerites convergent at middle and a pair of hammer-like dorsal processes in ventral view ( Fig. 25 View FIGURES 22 – 26 )............................................................................................................................... H. (H.) convergens View in CoL sp. nov.

8. Antennal 1st flagellomere unicolor............................................................................................................................... 9

- Antennal 1st flagellomere bicolor............................................................................................................................... 11

9. Antennal 1st flagellomere brownish yellow; aedeagus with a pair of acuate lateral processes, a pair of incurved horn–like apical processes in ventral view ( Fig. 60 View FIGURES 57 – 61 ) ...................................................... H. View in CoL (H.) yinggelingica sp. nov.

- Antennal 1st flagellomere entirely black .................................................................................................................... 10

10. Mesoscutum with brown stripes; tergite 6 with a small brown median spot; surstylus with a knife-like process in lateral view ( Fig. 27 View FIGURES 27 – 31 ); aedeagus with a pair of acuate triangular ventral processes and a pair of subuliform dorsal processes in ventral view ( Fig. 30 View FIGURES 27 – 31 ) .................................................................................................. H. View in CoL (H.) flavida sp. nov.

- Mesoscutum without brown stripes; tergite 6 without spot; surstylus consisting of a broad hairy digitiform process and a narrow curved bar-like process in lateral view ( Fig. 52 View FIGURES 52 – 56 ); aedeagus with a pair of acuate lateral processes and a pair of wide S–shaped and furcated dorsoapical processes in ventral view ( Fig. 55 View FIGURES 52 – 56 ) ....... H. View in CoL (H.) tortifurcata sp. nov.

11. Wing with a narrow brown stripe or spot on dm-cu ................................................................................................... 12

- Wing clear, without stripes or spots............................................................................................................................ 16

12. Abdominal tergite 6 without brown spot; hypandrium with a pair of short bar–like ventral process, aedeagus with a pair of dorsoapical teeth ( Figs. 15–16 View FIGURES 12 – 16 ) ................................................................................ H. (H.) apicomata View in CoL sp. nov.

- Abdominal tergite 6 with a brown median spot; hypandrium and aedeagus not as above......................................... 13

13. Tarsomere 5 entirely yellow to brownish yellow; surstylus undulating, incurved claviform ( Figs. 47–48 View FIGURES 47 – 51 ) ................. ....................................................................................................................................... H. View in CoL (H.) semicircularis sp. nov.

- Tarsomere 5 partly or entirely brown to black; surstylus not undulating ................................................................... 14

14. Tarsomere 5 brownish black; surstylus only with a stick–like process, slightly curved and narrow apically ( Fig. 37 View FIGURES 37 – 41 ); aedeagus with a pair of curved subuliform processes, a pair of acuate lateral processes and a pair of dorsal teeth on dorsal sclerites ( Figs. 40–41 View FIGURES 37 – 41 ) ............................................................................................... H. View in CoL (H.) nigritarsis sp. nov.

- Tarsomere 5 brown on apical 1/2; surstylus consisting of a small subuliform process and a digitifom process; aedeagus with pairs of small acuate processes or teeth on dorsal sclerites ......................................................................... 15

15. Protandrium with a long T–shaped ventral process ( Fig. 34 View FIGURES 32 – 36 ); aedeagal with dorsal sclerites contorted in lateral view ( Fig. 36 View FIGURES 32 – 36 ) .................................................................................................................................. H. View in CoL (H.) latissima sp. nov.

- Protandrium without long T–shaped ventral process; aedeagal with dorsal sclerites straight in lateral view .............. ................................................................................................................................ H. View in CoL (H.) yunnanensis Li, Li & Yang

16. Abdominal tergites 3–5 each with an entirely blackish brown posterior band and triangular median spots, and a triangular median spot confluent with a blackish brown posterior band; surstylus short claw–like, not projecting beyond tip of epandrium in lateral view; hypandrium without lobate ventral sclerites in ventral view; aedeagus with bifurcated sclerite ventrally and two pairs of spine–like processes on dorsal and ventral membranous parts (see Sasakawa, 1992: 173, fig. 19) ............................................................................................ H. View in CoL (H.) fasciventris Malloch

- Abdominal tergites 3–5 each with a black median spot and a pair of black lateral bands; surstylus long knife–like, projecting beyond tip of epandrium in lateral view ( Fig. 62 View FIGURES 62 – 66 ); hypandrium with a pair of lobate ventral sclerites in ventral view; aedeagus obliquely truncate at tip with a small acuate process ( Figs. 65–66 View FIGURES 62 – 66 ). H. (H.) zhangae View in CoL sp. nov.