Coproica hirticula Collin, 1956

Bergeron, Matthew D., Marshall, Stephen A. & Swann, John E., 2015, A review of the New World Coproica (Diptera: Sphaeroceridae) with a description of 8 new species, Zootaxa 3953 (1), pp. 1-157 : 31-32

publication ID

https://doi.org/10.11646/zootaxa.3953.1.1

publication LSID

lsid:zoobank.org:pub:C38905B9-01FC-4112-A759-50BE2B973BD7

DOI

https://doi.org/10.5281/zenodo.14951236

persistent identifier

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scientific name

Coproica hirticula Collin, 1956
status

 

Coproica hirticula Collin, 1956 View in CoL

Figs. 23 View FIGURES 21–28 , 145–155 View FIGURES 145–147 View FIGURES 148–151 View FIGURES 152–155

Coproica hirticula Collin, 1956: 178 View in CoL [both sexes]. Type locality: England, Cambs., Kirtling. STs (OXUM, see Pont 1995: 80).

Description. Body length 1.2–2.1 mm. Body brown to light brown. Face light brown, frons brown. Interfrontal bristles in 4 pairs, postocellar bristles slightly convergent, 1 pair of small postocellar setulae. Eye to gena ratio approximately 2:1; 3 genal setae in longitudinal row posterior to vibrissa.

Scutellum with approximately 15 discal setae in 2 rows between basal 1/4 and basal 1/2; 3 lateral setae between larger anterior and posterior marginal scutellar bristles. Katepisternum with 2 dorsal bristles, anterior and posterior bristles each approximately 1/5 and 2/3 as long as the distance between dorsal margin of katepisternum and wing base, respectively. Legs light brown, fore coxa light brown or yellow. Mid tibia without proximal posterodorsal bristles. Mid basitarsus with two stout anterior setae one subapical and one at 1/2, pair of ventral basal bristles, and pair of small ventral bristles at 1/2. Second costal sector 0.7–1.0 times as long as third costal sector. Costa extending 7 or more vein widths beyond R 4+5.

Male terminalia: Sternite 5 rounded rectangular, with paired posteromedial tabs, right tab larger, with posteromedial patch of dense microtrichia, discal bristles enlarged laterally. Sternite 6 straight medially. Epandrium with enlarged pair of dorsolateral bristles; subanal plate present, narrow, with medial suture; ventral margin of cercus convex. Surstylus curved along anteroventral margin, with row of small anteroventral bristles, and pointed posteroventral process. Postgonites straight, narrow, and unmodified. Basiphallus straight, tubular, posteriorly open dorsal to epiphallus; epiphallus with small, blunt process. Lateral sclerite of distiphallus with bifurcate proximal dorsal process extending laterally and medially, ventral apical portion cup shaped, and with additional, sharp ventral process; central sclerite short, fused to lateral sclerites beneath proximal dorsal process. Membranous sheets broad.

Female terminalia: Tergite 8 long laterally, short dorsally. Tergite 10 pollinose, narrowly fused to cerci laterally; cerci conical, pollinose, apical bristles long and sinuate. Sternite 7 broad, with large triangular point on posterior margin. Sternite 8 small, with 4 minute setae along posterior margin. Sternite 10 with 1 pair of enlarged medial bristles, posterior margin not curved dorsally, lateral margins curved dorsally. Spermathecae spherical, with conical base; duct length 7–8 times as long as spermathecal bulb.

Type material. Syntypes: ENGLAND: Cambs., Kirtling , v–vii, x.1951, (♂ ♀, OXUM, not examined) .

Material examined. See Appendix A.

Distribution: Afrotropical: South Africa; Australasian/Oceanian: Australia (ACT, NSW, QLD), New Zealand; Nearctic: Canada (AB, BC, NB, ON, SK) , USA (AK, AR, AZ, CA, FL, GA, IL, IN, KY, MA, MI, MN, MO, NC, NM, OK, SC, TN, TX, UT, WY); Neotropical: Argentina, Barbados, Bolivia, Brazil, Chile, Colombia, Costa Rica, Ecuador, Guatemala, Jamaica, Mexico (CHI, COA, HID, ROO, VRC), Peru, Trinidad; Oriental: China (HKG), Taiwan; Palaearctic: Belgium, Bulgaria, Canary Is. (Spain), Cyprus, Czech Republic, Denmark, Finland, France, Germany, Great Britain, Greece (incl. Crete), Hungary, Israel, Italy, Japan, Latvia, Madeira (Portugal), Malta, Norway, Poland, Romania, Slovakia, Spain (incl. Balearic Is.), Sweden, Switzerland, United Arab Emirates.

Comments. Coproica hirticula has a cosmopolitan distribution and is among the most common species of Coproica in the New World. It is easily distinguished from other Coproica by the absence of proximal posterodorsal bristles on the mid tibia ( Fig. 18 View FIGURES 13–20 ), a short anterior and long posterior katepisternal bristle ( Fig. 5 View FIGURES 3–5 ), and the unique arrangement of ventral setae on the mid basitarsus ( Fig. 23 View FIGURES 21–28 ). Coproica hirticula is the sister species to a clade including C. novacula , and the C. vagans and C. urbana species groups. Coproica hirticula is most morphologically similar to C. novacula ; both species are realtively plesiomorphic and unplaced to species group.

We were unable to obtain the type specimens in order to confirm the concept of C. hirticula used here. We have used the concept of C. hirticula from Richards (1960).

Collin, J. E. (1956) Some new British Borboridae (Diptera). Journal of the Society for British Entomology, 5, 172-178.

Richards, O. W. (1960) On two N. American species of Leptocera Olivier, subgenus Coproica Rondani, with a review of the subgenus (Diptera, Sphaeroceridae). Annals and Magazine of Natural History, Series. 13, 2 (16), 199-208. http://dx.doi.org/10.1080/00222935908651023

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FIGURES 21–28. left mid basitarsus, ventral. 21. Coproica bifurcata sp. nov. 22. Coproica brachystyla sp. nov. 23. Coproica hirticula Collin. 24. Coproica lugubris (Haliday). 25. Coproica mitchelli (Malloch). 26. Coproica novacula sp. nov. 27. Coproica rufifrons (Hayashi). 28. Coproica setulosa (Duda). Abbreviations: ARS = anterior row of setae.

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FIGURES 145–147. Coproica hirticula Collin. 145. Male left wing, dorsal. 146. Male terminalia, left lateral. 147. Male terminalia, posterior (setae on right omitted).

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FIGURES 148–151. Coproica hirticula Collin. 148. Spermathecae. 149. Aedeagus and associated structures with postgonites removed. 150. Male sternite 5 and synsternite 6–8. 151. Left postgonite. Abbreviations: AT = asymmetrical tab, BP = bifurcate process, LS = lateral sclerite, MS = membranous sheet, VP = ventral process.

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FIGURES 152–155. Coproica hirticula Collin. 152. Female terminalia, dorsal. 153. Female terminalia, ventral. 154. Female terminalia, left lateral. 155. Female sternite 8.

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FIGURES 13–20. Figs 13–15: Coproica acutangula (Zetterstedt). 13. Male left fore basitarsus, posterior. 14. Male left hind basitarsus, anterior. 15. left mid tibia, dorsal. Figs 16–20: left mid tibia, dorsal. 16. Coproica bifurcata sp. nov. 17. Coproica ferruginata (Stenhammar). 18. Coproica lugubris (Haliday). 19. Coproica novacula sp. nov. 20. Coproica rufifrons (Hayashi).

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FIGURES 3–5. left katepisternum. 3. Coproica diabolica sp. nov. 4. Coproica ferruginata (Stenhammar). 5. Coproica novacula sp. nov.

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Diptera

Family

Sphaeroceridae

Genus

Coproica