Bromus dolichocarpus Wagnon (1950: 65)

Saarela, Jeffery M., Peterson, Paul M. & Valdés-Reyna, Jesus, 2014, A taxonomic revision of Bromus (Poaceae: Pooideae: Bromeae) in México and Central America, Phytotaxa 185 (1), pp. 1-147 : 77-83

9. Bromus dolichocarpus Wagnon (1950: 65) . Figs. 32 , 33 .

Bromopsis dolichocarpa (Wagnon) Holub (1973: 167) . Type:— MÉXICO. Michoacán: in fir forest, ca. 10 mi NW of Ciudad Hidalgo, 18 March 1949, R. McVaugh 9887 (holotype MICH-1108613 !, isotype US-2012739 !) .

Plants perennial, not rhizomatous. Culms up to 192 cm tall, 2–4 mm wide at base, erect, the bases ± decumbent, pubescent below inflorescences; nodes 2–6, glabrous, puberulent or pubescent. Leaf sheaths sparsely to moderately pilose, hairs up to 1.5 mm long; auricles usually present; ligules 0.6–1.5(–3) mm long, glabrous; blades up to 39 cm × 3–16 mm, flat, adaxial surfaces pilose, hairs up to 1.5 mm long, abaxial surfaces glabrous, sometimes with scattered hairs adjacent to the midnerve, or sparsely pubescent, margins serrulate. Panicles up to 33 cm × 3–11 cm, open, nodding, branches ascending, 0.7–12 cm long, shorter or longer than spikelets, scabrous to densely pubescent, hairs up to 0.6 mm long, 1–2(–3) spikelets per branch. Spikelets 2.5–3.6(–4) cm long, 5–8-flowered, elliptic to lanceolate, terete to moderately laterally compressed, rachillas sometimes visible at maturity; glumes glabrous, sometimes a few hairs proximally and along the margins, or pubescent proximally, margins hyaline, midnerves scabrous, sometimes pubescent proximally, hairs up to 0.6 mm long; lower glumes 10–13(–14) mm × 0.3–0.7(–1) mm, narrowly lanceolate, 1(–3)-nerved, green along the nerve(s), apices acute, rarely acuminate; upper glumes 12–19 mm × 0.7–1.3 mm, lanceolate, 3-nerved, green along the nerves, green to light green–translucent between the nerves, apices acute, rarely acuminate; lemmas (13–) 14–20 mm × 0.9–1.5 mm, linear-lanceolate, rounded over the backs, apices acute, 3(–5)-nerved, the nerves strong, backs puberulent proximally, glabrous distally, pubescent to pilose between the lateral nerves and the margins, hairs up to 0.5 mm long, light greentranslucent between the nerves, margins hyaline, nerves scabrous or pilose, hairs up to 0.6 mm long; awns 7–13 mm long, attached up to 0.5 mm below lemma apex, straight; paleas shorter and narrower than the lemmas, backs

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Phytotaxa 185 (1) © 2014 Magnolia Press 77

scabridulous to pubescent, keels ciliate, cilia up to 0.1 mm long; anthers 1.1–2.5 mm long; caryopses 10−15 mm long, light brown. 2 n = 28 ( Wagnon 1950).

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Distribution: ―Native. Bromus dolichocarpus is distributed from central México (Chiapas, Distrito Federal, Durango, Guerrero, Hidalgo, Jalisco, Estado de México, Michoácan, Morelos, Oaxaca, Puebla, Veracruz) to Guatemala (El Progreso, Quezaltenango, San Marcos) ( Fig. 34 ). It was reported from Colima and Jalisco by Beetle (1977), but we have not seen specimens from these states. This is the first report of B. dolichocarpus from Durango, as the species was not reported for the state by Herrera Arrieta (2001, 2014).

Ecology: — Bromus dolichocarpus is found on gentle slopes and flats that are often steep and rocky and along small draws; associated with Pinus spp. , Pinus hartwegii Lindley (1839: 62) , Abies religiosa , Quercus spp. , Juniperus deppeana , Arbutus xalapensis , Gaultheria Linnaeus (1753: 395) , Pteridium Gled. ex Scopoli (1760: 169) , Tagetes Linnaeus (1753: 887) , Symphoricarpos Duhamel du Monceau (1755: 295) , Arctostaphylos pungens , Alnus , Ribes Linnaeus (1753: 200) , Prunus serotina Ehrhart (1783: 285) , Rhamnus Linnaeus (1753: 193) , Crataegus Linnaeus (1753: 475) , Toxicodendron Miller (1754a) , Geranium ( Linnaeus 1753: 676) and Baccharis . Elevation: 2500–3800 m ( Guatemala), (1900–) 2200–3920 m ( México).

Common Names: ―Unknown.

Comments: ― Bromus dolichocarpus is one of three Mexican species with 3-nerved lemmas, which are newly recognized here in sect. Mexibromus . It is easily distinguished from B. attenuatus by its pubescent lemmas, longer awns and leaf blades that do not distinctly narrow towards their bases, and from B. densus by its longer awns, shorter anthers and wider leaf blades (see taxonomic key).

Specimens Examined:― GUATEMALA. El Progreso: Sierra de Las Minas, between Finca Piamonte and top of Montaña Piamonte , along Joya Pacayal , [15.13°N, 89.75°W], 2500–3000 m, 7 February 1942, J. A . Steyermark 43619 (F-1202053, US-2436906 ). Quezaltenango: Cerro Quemado , [14.8°N, 91.52°W], 2600 m, September 1954, M . de Koninck 210 ( US-2153572 ); mountains SE of Palestina, on old road to San Juan Ostuncalco , [14.87°N, 91.62°W], 2550–2850 m, 21 January 1941, P. C . Standley 84221 (F-1202285). San Marcos: along road between San Sebastián at km 21 and km 8, 8– 18 mi NW of San Marcos, [15.06°N, 91.83°W], 2700–3800 m, 15 February 1940, J. A . Steyermark 35769 (F-1052081) & 35771 (F-1052024); Sierra Madre Mountains , ca. 6 km (airline) Nof San Marcos, [14.96°N, 91.79°W], 2700 m, 13 December 1963, L. O Williams , A . Molina R. & T.P. Williams 25876 (F-1656824, US-2537602 ) & 25885 ( NY, mixed sheet with B. carinatus var. marginatus ) . MÉXICO. Chiapas: Mpio. Zinacantán, near the summit of Muk'tavits , [16.3333°N, 92.75°W], 2745 m, 16 November 1981, D. E . Breedlove & B . Bartholomew 55564 ( CAS, GH, MO) . Distrito Federal: cerca de Eslava, 19.42°N, 99.17°W, 19 October 1952, J . Rzedowski 2005 ( MEXU-100903 , MICH-1119221 ); Cumbre de Estepa, 18.05°N, 97.7°W, April 1842, Liebmann 487 ( US-2473643 ); in imperio Mexicano , from Zuccarini , Karwinsky s.n. ( US-1009477 ); San Rafael , 19.25°N, 98.79°W, 2600 m, 18 September 1932, M . St. Pierre 938 ( MICH-1119222 , P-03629787 , US); San Roman, Liebmann 501 ( MO, US-2473641 ); 10 mi Eof Amecameca on road towards Paso de Cortés , in Parque Nacional Izta-Popo, 19.0764°N, 98.71°W, 2840 m, 12 October 2001, P. M . Peterson & Y . Herrera-Arrieta 16160 ( CAN, MO, US); 1er Dínamo, Delegacíon Magdalena Contreras, [19.42°N, 99.13°W], 2580 m, 17 October 1985, A . Miranda & P . Guerrero 141 ( MEXU-1110774 ); Desierto de Los Leones , 19.2531°N, 99.3286°W, 19 October 1960, T . Tateoka 389 ( MEXU-151424 ) & 1151 ( US-2380596 ); Desierto de Los Leones, Delegación de Guajimalpa , [19.2533°N, 99.3308°W], 25 January 1976, M . González E . 82 ( ANSM); Mpio. Cuajimalpa, Desierto de Los Leones , [19.2533°N, 99.3308°W], 2 November 1979, 2700 m, F . Santoyo 178 ( ANSM); Río Grande , July 1928, E . Lyonnet 266 ( MEXU-258403 ); Vallede México, [19.42°N, 99.13°W], 3500 m, 9 December 1951, E . Matuda 25754 ( MEXU-499579 , MEXU-499578 ); Vallede México, Lago Zempoala , Hdo. de Morelos, 19.0494°N, 99.3175°W, 3000 m, 7 October 1951, E . Matuda 25587 ( MEXU, US-2079183 ); Vallede México, San Rafael , 19.2333°N, 98.75°W, 2600 m, 19 November 1950, E . Matuda 18722 ( MEXU, US-2040890 ) .

Durango: Mpio. Súchil, Reserva de la Biosfera La Michilia, Mesa El Burro, 23.4108°N, 104.3108°W, 2650 m, 16 September 1982, Y. Herrera 261 ( ANSM). Guerrero: Mpio. Chilpancingo de los Bravos, Omiltemi , [17.55°N, 99.51°W], 1900 m , 15 November 1991, V. Nandi 134 ( MEXU-1089776 ); Mpio. Chilpancingo de los Bravos, al NW de Omiltemi en direccion a la toma de Agua de la Cañada de la Perra, [17.55°N, 99.5°W], 2120 m , 10 October 1985, R. Antonio Ocampo 405 ( MEXU-502476 ). Hidalgo: El Chico, 20.2167°N, 98.7333°W , July 1929, P. Lyonnet 262 ( US-1034151 ); Las Ventanas, Mineral del Chico, [20.17°N, 98.73°W], 2970 m , 19 May 1993, J. Práxedes Pérez 23 ( MEXU); Pachuca Dist. , below Pueblo Nuevo on road from Real del Monte to El Chico, 20.1667°N, 98.7333°W, 2850 m , 17 October 1946, H. E. Moore , Jr. 1531 ( GH, US-1963095 ). Jalisco: on the divide above headwaters of Río Mascota (25–30 km, airline, SE of Talpa de Allende), steep mountainsides and barrancas 1–2 mi Nof the sawmill " La Cumbre", and 11–12 road mi Sof El Rincón, 20.0403°N, 103.8753°W, 2000–3000 m ,

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Phytotaxa 185 (1) © 2014 Magnolia Press 79 29 November 1960, R . McVaugh 21533 ( NY); steep mountainsides above Amacueca, near the summit of the plateau, road to Tapalpa , [19.1556°N, 102.9889°W], 2100–2250 m, 2 November 1960, R . McVaugh 20654 ( NY). Estado de México: 8 km de la desviación al Nevado de Toluca, [19.1083°N, 99.7583°W], 3260 m, 29 November 1983, Manrique, Jaramilo & Núñez 558 ( MEXU-1089780 ); a 6 km sobre la desviación a Tequesquiapan, carretera Toluca-Temascaltepec, 19.31°N, 98.97°W, 2580 m, 25 October 1983, Manrique, Jaramillo & Nuñez 445 ( MEXU- 1098071 , MEXU-1033954 ); al SE de Zinacantepec, [19.14°N, 99.71°W], 3920 m, 8 April 1992, A . Vega, A . Ramírez & R . Mendoza 2 ( MEXU-1089771 ); camino de terracería Malinalco-San José Tenería, [18.93°N, 99.49°W], 27 October 1988, J. G . Aldape et al. 34 ( MEXU); carretera Toluca-Temascaltepec, [19.12°N, 99.88°W], 2530 m, Manrique, Guerrero, Guzmán & Jaramillo 140 ( MEXU); Cieneguillas de Cabra , entre la carretera Sultepec-La Puerta, al Sdel Nevado de Toluca, [19.1083°N, 99.7583°W], 2600 m, 1 August 1981, R. Guzmán 4021 ( MEXU); entre Texcaltitlán y Las Juntas , al Sdel Nevado de Toluca, [18.93°N, 99.95°W], 2660 m, 1 August 1981, R. Guzmán 4015 ( MEXU-1110805 ); Estación experimental San Cayetano, 2500 m, 2 August 1978, F. M . Fuchs 97 ( MEXU-231170 ); Mpio. Amecameca, 1 km al noroeste de San Antonio, 2550 m, 11 December 1977, N . Valentín M . 74 ( ANSM); Mpio. Amecameca, km 15 carretera Amecameca-Tlamacas , [19.08°N, 98.67°W], 2 October 1992, A . Miranda & G . Villegas 636 ( MEXU); Mpio. Atlautla de Victoria, Volcán Popocatépetl, ladera W por Atlautla, [19.05°N, 98.64°W], 2725 m, 24 October 1987, J . Hernández 230 ( MEXU-565906 ); Mpio. México, 3 km al Ede San Rafael , [19.21°N, 98.74°W], 2700 m, 19 September 1976, C . Flores G . s.n. ( MEXU-292304 ); Mpio. Morelos, Valle del Tepeite, [19.27°N, 99.18°W], 16 October 1937, E . Lyonnet 18112 ( MEXU-278684 ); Mpio. Ocuilan de Arteaga , 2 km al NW de Sta. Mónica de Ocuilan, 18.9833°N, 99.4417°W, 2350 m, 4 February 1990, González & O . Miranda J . 328 ( MEXU-1031709 ); Mpio. Tepotzotlán, 2400 m, 26 October 1975, J. R . Torres V. ( ANSM); Mpio. Texcoco, 2 km al SE de San Pablo Ixayoc, sobre el camino hacia el Aguaje, 12.5 km al SE de Texcoco , [19.41°N, 98.86°W], 2670 m, 21 October 1976, S . D. Koch 76210 ( MEXU-265511 , MO); Mpio. Tlalmanalco , 3 km al Ede San Rafael, [19.21°N, 98.76°W], 2700 m, 27 May 1965, J . Rzedowski 19863 ( MEXU- 187911 ); Mpio. Tlalmanalco, a 3 km al Ede San Rafael , [19.21°N, 98.76°W], 2700 m, L. M . García R. 73 ( ANSM, MEXU-220002 ); Mpio. Villa de Allende, San Cayetano (estación experimental) al Nde Agua Escondida y a 26 km al NE de Valle de Bravo, [19.37°N, 100.15°W], 2490 m, August–December 1974, Ma. E . Maury Hdez., V . Serrano Cardenas & S . Gallina Tessaro 5 ( MEXU-200087 ); Mpio. Villa de Allende, San Cayetano (Estación experimental), al Nde Agua Escondida y a 26 km NE de Vallede Bravo, [19.33°N, 100°W], 2490 m, August–December 1974, S . Gallina Tessaro s.n. ( MEXU); Mpio. Villa Guerrero, carretera de cuota de Tenango de Arista a Ixtapan de la Sal , [18.84°N, 99.68°W], 2200 m, 30 September 1995, H . Vibrans 5564 ( MEXU-895630 ); Mpio. Zimatlán, Cañada El Frijolón, 8 km al NW de La Cofradía , comunidad de San Pedro El Alto, [19.61°N, 99.9°W], 2790 m, 8 October 1998, A. G . Miranda M. & O. L . Hernández Martínez 248 ( MEXU-907764 ); 1 km Wof Amecameca along road to Ayapango, 19.15°N, 98.7833°W, ca. 2500 m, 29 September 1978, H. H . Iltis, J . Doebley & A . Lasseigne 764 (F-1919412, MO, MSC-277227 , RSA-POM-320576 , US); 1.4 mi Nof Tequexquipan , 2490 m, 19.071°N, 99.9388°W, 10 October 2001, P. M . Peterson & Y . Herrera-Arrieta 16139 ( CAN, MO, P-03216909 , US); 1.9 mi Wof Chiltepec on Hwy. 12, 18.9268°N, 99.8442°W, 2596 m, 10 October 2007, P. M . Peterson, J. M . Saarela & M. J . Flores Villegas 21364 ( CAN, MO, P-03631106 , US); 12.1 mi NE of San Miguel, SW of Volcán Toluca , 19.0317°N, 99.9102°W, 2826 m, 10 October 2001, P. M . Peterson & Y . Herrera-Arrieta 16143 ( CAN, MO, P- 03216911 , US); 5 km al NW de Amecameca, laderas vecinas al Cerro Tenayo , [19.12°N, 98.76°W], 2550 m, 17 November 1968, A . Pineda R. 603 ( MICH-1119227 , MSC-223104 ); 6.7 mi Eof Amecameca towards Paso de Cortés, 19.0706°N, 98.6923°W, 2488 m, 11 October 2007, P. M . Peterson, J. M . Saarela & M. J . Flores Villegas 21382 ( CAN, MO, US); 8.7 mi NE of La Comumidad on Hwy. 134 toward Toluca , 19.1858°N, 99.86°W, 3190 m, 9 October 2001, P. M . Peterson & Y . Herrera-Arrieta 16132 ( US); at La Comunidad, along stream just below puebla ( La Comunidad), 19.1518°N, 99.9118°W, 2607 m, 9 October 2001, P. M . Peterson & Y . Herrera-Arrieta 16136 ( CAN, MO, P-03216910 , US); Coulter 1651 ( GH); Mpio. Amecameca , 1 km al NE de San Antonio, 19.12°N, 98.76°W, 2550 m, 21 October 1968, J. J . Rzedowski 26431 ( MICH-1119223 , MSC-233266 ); Mpio. Amecameca , cerca de San Pedro Nexapa, 19.08°N, 98.74°W, 9 October 1966, 2550 m, J. J . Rzedowski 23285 ( MSC-217972 ); Mpio. Amecameca, Zumpango, carretera al Paso de Cortés, 16 km al Edel entronque con la carretera Amecameca-Cuautla, Mor., 19.06°N, 98.7°W, 3300 m, 20 November 1976, S. D. Koch 76275 B ( US- 2824581 ); Mpio. Huixquilucan, 5 km al Ede México, carretera de la Marquesa a Naucalpan , 19.3833°N, 99.3333°W, 2600 m, 29 July 1976, S . D. Koch 76107 ( MEXU-270647 , CHAPA, MO, US); Mpio. Texcoco, 2 km al SE de San Pablo Ixayoc, 12.5 km al SE de Texcoco , 19.42°N, 98.83°W, 2670 m, 21 October 1976, S .D. Koch

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Phytotaxa 185 (1) © 2014 Magnolia Press 81 76210 (US-2824630); Sslopes of Nevado de Toluca, 35 km (road) SW of Toluca on Hwy. 130, 19.0833°N, 99.8333°W, ca. 3000 m, 29 August 1965, K. Roe, E. Roe & S. Mori 149 1 ( MSC-229117 ); Temascaltepec Dist. , Comunidad, 19.1292°N, 99.9347°W , 2600 m, 25 November 1932, G. B. Hinton 2459 ( NY, US-1867841 ); Temascaltepec, Crucero, 19.04°N, 100.23°W , 28 October 1935, G. B. Hinton 8395 ( MO, NY, 2 sheets, RSA-POM- 364158 ); Tlalmanalco, 3.0 km al Ede San Rafael, [19.21°N, 98.74°W] , 2700 m, 19 September 1976, C. Flores G. s.n. ( MO, SD-110652 ); Volcán de Toluca , 3760 m, 19.1014°N, 99.7694°W, 18 October 1953, E. R. Sohns & E. Matuda 1004 ( US-2118910 ); Wslopes of Nevado de Toluca , 35 km (road) SW of Toluca on Hwy. 130, 19.0833°N, 99.8333°W , 3000 m, 29 August 1965, K. Roe, E. Roe & S. Mori 1491 ( US-3115415 ); Wslopes of Nevado de Toluca , 35 km (road) SW of Toluca on Hwy. 130, [19.11°N, 99.79°W], ca . 3000 m, 29 August 1965, K. Roe, E. Roe & S. Mori ( GH). Michoácan: Cerro Tancítaro, 27 km al Wde Uruapan en linea recta, 1 km al oeste de El Tepetate, [19.41°N, 102.3°W] , 2850 m, 29 August 1996, I. Garcia Ruíz, J.A. Machuca & M. Cházaro 4153 ( MICH- 1119229 ); 11 km al Nde Indaparapeo , sobre el camino a Las Peras, [19.78°N, 100.97°W] , 2200 m, 8 November 1987, J. Rzedowski 45692 ( MEXU-809934 ); 37 mi Eof Morelia on MEX 15, 19.67°N, 100.73°W , 8000 ft [2438 m], 20 October 1972, L. H. Harvey & J. T. Witherspoon 9419 ( MICH-1119244 ); 5.7 mi NW of Huajumbaro on road towards Santa Clara, 19.7364°N, 100.78°W , 2430 m, 8 October 2001, P. M. Peterson 16129 ( CAN, MO, US); a 12 km al Nde Uruapan , carretera a Carapan, [19.41°N, 102.06°W] , 2075 m, 22 August 1980, J. C. Soto Núñez 2393 ( MEXU); above Villa Escalante on road to Ario de Rosales , 19.4°N, 101.65°W , 2340 m, 11 November 1949, H. E. Moore, Jr., E. Hernández-X. & H. Porras-H. 5634 ( US-1983697 ); ca. 10 mi NW of C. Hidalgo, 19.66°N, 100.71°W , 4 July 1950, H. K. Wagnon 1615 ( MEXU-772510 , US-2154832 ); ca. 18 mi Sof Pátzcuaro , 19.31°N, 101.66°W , 2713–2743 m, 20–25 November 1961, R. M. King & T. R. Soderstrom 5178 ( MICH-1119226 , MEXU- 55441 , NY, mixed sheet with B. carinatus var. carinatus ), US-2378973 ); canyon just NW of Congoon on Hwy. 120 towards Pátzcuaro , 19.3806°N, 101.65°W , 2350 m, 8 October 2001, P. M. Peterson 16128 ( CAN, MO, P- 03216912 , US); Cerro Azul, vicinity of Morelia , 19.7167°N, 101.1833°W , 2300 m, 4 November 1909, G. Arséne 3273 ( MO, US-1002650 ); grown at the Botanical Gardens, University of Michigan, Ann Arbor, from seed collected with the type in fir forests ca. 10 mi NW of Ciudad Hidalgo , 4 July 1950, H. K. Wagnon 1615 ( MICH- 1119225 [chromosome number voucher specimen, Wagnon 1950], MO); Mesa Karitsho o del Padre, SE de San Francisco Pichataro, [19.57°N, 101.81°W] , 2625 m, 9 November 1978, J. Coballero & C. Mapes 644 ( MEXU); Mpio. Charo, El Salto de Agua , 19.65°N, 100.9444°W , 1 February 1994, J. A. Torres Estrada 35 ( MEXU-1009115 , MEXU-1089785 ); Mpio. Morelia , 3 km al Wde San Miguel del Monte, [19.7°N, 101.18°W] , 2300 m, 25 October 1986, J. Rzedowski 41312 ( MEXU-658295 ); Mpio. Pátzcuaro, vertiente Wdel Cerro del Burro , [19.415°N, 101.5006°W] , 2900 m, 23 November 1986, J. Rzedowski 41962 ( ANSM); Mpio. Queréndaro, 2 km al Nde Real de Otzumatlán , 2350 m, 20 November 1986, J. Rzedowski 41855 ( ANSM); Mpio. Senguio, Cerro Ross Azul, al Ede Chincus , 19.73°N, 100.35°W , 2800 m, 23 November 1986, J. Santos Martínez 1969 ( MICH-1119219 ); Mpio. Zacapu, ± 5 km de Zacapu, por la carretera a Zamora , 2000 m, 9 September 1988, A. Grimaldo Núñez 406 ( ANSM); Mpio. Zinapécuaro, 5 km al SSE de Jeráhuaro , 2550 m, 7 December 1987, J. Rzedowski 46150 ( ANSM, MEXU-915763 ); slopes of lava flow Eof San Juan Nuevo , ca. 8 km Sof Uruapan, 19.41°N, 102.06°W , 11–15 October 1961, R. M. King & T. R. Soderstrom 4760 ( MEXU-55372 ); vicinity of Morelia , Cerro Agul , [19.69°N, 101.16°W], 4 November 1909, G. Arséne 3293 [?] ( GH); Campo Experimental INIF, Uruapán , 2100 m, 9 October 1976, C. L. Días Luna 7504 ( ANSM); carretera Pátzcuaro-Uruapan km 25, [19.44°N, 101.84°W], 4 December 1986, J. J. Ortíz 1195 ( MEXU-638476 ); Mpio. Ocampo , 2 km al Ede Ocampo, camino al Rosario, [19.58°N, 100.34°W] , 2400 m, 2 November 1989, R. Torres & M. Ramírez 13553 ( MEXU-904824 ); Mpio. Pátzcuaro, 6 km al Sde Pátzcuaro , carretera a Opopeo, [19.5158°N, 101.6094°W] , 2400 m, 10 November 1985, J. Espinosa Garduño 1990 ( MEXU-993903 ); Mpio. Pátzcuaro, lado SW del Cerro El Fríjol, [19.5158°N, 101.6094°W] , 2700 m, 31 October 1985, H. Díaz Barriga 1652 ( MEXU-662626 ). Morelos: km 62 de la carretera Curenava-México, [19.27°N, 99.17°W] , 2460 m, 8 February 1984, Manrique, Beetle et al. 733 ( MEXU-1110847 ); Valle del Tepeite , [19.27°N, 99.18°W], 15 October 1937, E. Lyonnet 1812 ( MEXU-278689 ); Valle del Tepeite , 19.27°N, 99.25°W , 16 October 1937, E. Lyonnet & J. Elcoro 1811 ( MEXU, US-1746382 ). Oaxaca: 1.6 mi Nof San Miguel Suchixtepec along Hwy. 175, 16.1157°N, 96.4745°W , 2722 m, 23 September 2008, P. M. Peterson & J. M. Saarela 22378 ( CAN, US); 9.5 mi Nof San Cualimojoyas on road towards Santa María Yavesia, 17.1966°N, 96.4512°W , 2727 m, 20 September 2008, P. M. Peterson & J. M. Saarela 22307 ( CAN, US); Campamento Río de Molino , 4 km al SW de San Miguel Suchistepec, 16.09°N, 96.47°W , 2250 m, 21 September 1965, J. Rzedowski 20999 ( MICH-1119224 ), 21039 ( MICH-1119220 ) & 21082 ( MSC-210582 ); Cerro de San Felipe, Distrito del Centro , 17.1°N, 96.85°W ,

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2300 m, 12 October 1920, C. Conzatti 4069 ( MEXU-6111 , US-1014111 ); Mpio. Zimatlán, Comunidad El Tlacuache, San Pedro El Alto, 16.04°N, 96.47°W , 2420 m, 9 October 1998, A. G. Miranda & O. Hernández M. 306 ( MEXU-901092 ); 1.2 mi Eof San José del Pacifico on road towards San Sebastián, 16.1698°N, 96.4898°W , 2641 m, 22 September 2008, P. M. Peterson & J. M. Saarela 22353 ( CAN, US). Puebla: 23.4 Eof Amecameca on road towards Puebla in Parque Nacional Izta-Popo , 19.0897°N, 98.6064°W , 3330 m, 12 October 2001, P. M. Peterson & Y. Herrera-Arrieta 16170 ( US); 24.3 mi Eof Amecameca on road towards Puebla, in Parque Nacional Izta-Popo, 19.085°N, 98.6°W , 3316 m, 13 October 2001, P. M. Peterson & Y. Herrera-Arrieta 16175 ( CAN, US). Veracruz: Orizaba, 19.0167°N, 97.22°W , s.d., Liebmann 496 ( US-2473642 ) .

Figures

FIGURE 32. Illustration of Bromus dolichocarpus. A. Spikelet. B. Lemma C. Inflorescence. D. Ligule. Based on Breedlove 55564 (GH). Illustration by Paulette Dennis © Paulette Dennis.

FIGURE 33. Bromus dolichocarpus. Pineda R. 603 (MICH-1119227).

FIGURE 34. Geographical distribution of Bromus dolichocarpus in México and Central America.