Helina annosa (Zetterstedt, 1837)

Li, Siyu, Xue, Wanqi, Nihei, Silvio S. & Zhang, Chuntian, 2025, Four new species of Helina Robineau-Desvoidy from far northwestern China (Diptera: Muscidae), Zootaxa 5719 (2), pp. 207-227 : 209-211

Key to Chinese species of Helina annosa -group (males)

(modified from Xue & Sun 2014)

1 Apex of scutellum with hairs ventrally.................................................................... 2

- Apex of scutellum bare ventrally......................................................................... 8

2 Sternite 1 bare........................................................................................ 3

- Sternite 1 setulose..................................................................................... 5

3 r-m crossvein without dark spot............................................................. H. luyashanensis

- r-m crossvein with dark spot............................................................................ 4

4 Mid femur with complete pv row in entire length, 5 strong in basal half; hind femur with strong av on apical half.................................................................................................... H. subevecta

- Mid femur with developed pv on basal half only; hind femur with row of 8 or 9 long av distributed evenly as a comb along the apical half...................................................................... H. ctenophora Xue sp. nov

5 Arista short plumose, the longest hairs about 1/2 the width of postpedicel................................ H. nemorum

- Arista plumose or long plumose, the longest hairs longer than the width of postpedicel.............................. 6

6 r-m crossvein with dark spot; fore tibia without p.................................................. H. mimevecta

- r-m crossvein without dark spot; fore tibia with p............................................................ 7

7 Lateral surface of scutellum setulose............................................................... H. evecta

- Lateral surface of scutellum bare.............................................................. H. pleuranthus

8 Sternite 1 bare........................................................................................ 9

- Sternite 1 setulose.................................................................................... 19

9 Arista short plumose, the longest hairs about 2/3 width of postpedicel.............................. H. brachytophalla

- Arista plumose or long plumose, the longest hairs not shorter than the width of postpedicel.......................... 10

10 Inner vitta of scutum (between acr rows) reaching scutellum.................................................. 11

- Inner vitta of scutum (between acr rows) not reaching scutellum............................................... 14

11 pra absent.......................................................................................... 12

- pra distinct......................................................................................... 13

12 Frons about 2.5x as wide as anterior ocellus; fore tibia without p, mid femur with 4–5 strong pv, mid tibia with 3 p............................................................................................. H. jiaodingshanica

- Frons about 1.3x as wide as anterior ocellus; fore tibia with 3 p, mid femur with 8–10 pv, mid tibia with 6– 7 p... H. dianica

13 Postpedicel about 3.5x as long as broad; r-m crossvein without dark spot; fore tibia with 2– 3 p............. H. nigriannosa

- Postpedicel about 2.5x as long as broad; r-m crossvein with dark spot; fore tibia without median p........... H. barkamica

14 Lateral surface of scutellum bare................................................................ H. shuensis

- Lateral surface of scutellum setulose..................................................................... 15

15 Ventrolateral margin of scutellum setulose................................................................ 16

- Ventrolateral margin of scutellum bare................................................................... 17

16 r-m crossvein without dark spot; hind femur with 4 av on apical 1/4; hind tibia with 4 av.................... H. azaleella

- r-m crossvein with dark spot; hind femur with 9–10 av on apical 3/4; hind tibia with 9–10 av on basal 2/3................................................................................................... H. subnigriabdomilis

17 pra absent................................................................................. H. sichuanica

- pra distinct......................................................................................... 18

18 Fore tibia with 1 p; lateral lobe of sternite 5 verrucose............................................... H. mimifica

- Fore tibia with 2–3 p on apical half; lateral lobe of sternite 5 broad, not verrucose...................... H. emeishanana

19 pra absent.......................................................................................... 20

- pra distinct......................................................................................... 23

20 Fore tibia with 2–4 median p................................................................. H. saementosa

- Fore tibia without median p............................................................................ 21

21 Inner vitta of scutum (between acr rows) not reaching scutellum...................................... H. maquensis

- Inner vitta of scutum (between acr rows) reaching scutellum.................................................. 22

22 R 4 +5 and M diverging in distal part; mid femur with complete pv row, strong on basal half; mid tibia with 6–7 p; hind tibia with 5 ad........................................................................................ H. floscula

- R 4 +5 and M straight; mid femur with pv row on basal 2/5; mid tibia with 4–5 p on apical 2/3; hind tibia with 3– 4 ad................................................................................................ H. subfloscula

23 Lateral surface of scutellum bare........................................................................ 24

- Lateral surface of scutellum setulose..................................................................... 26

24 Costal spine absent...................................................................... H. plumipostitibia

- Costal spine distinct.................................................................................. 25

25 r-m crossvein with dark spot; mid femur with 3–4 pv on basal half; hind tibia with 3–4 av and 4 ad........... H. xinanana

- r-m crossvein without dark spot; mid femur with complete pv row; hind tibia with 4–5 av and 2– 3 ad...... H. midmargiseta

26 Ventral lateral margin of scutellum bare................................................................... 27

- Ventral lateral margin of scutellum setulose................................................................ 30

27 Katepisternal setae 1+2................................................................... H. quadratisterna

- Katepisternal setae 2+2................................................................................ 28

28 Mid tibia with 7–9 strong p................................................................. H. setipostitibia

- Mid tibia at most with 5 p.............................................................................. 29

29 Arista plumose, the longest hairs subequal to width of postpedicel; mid tibia with 5 p; hind femur with 5–6 av on apical 1/3........................................................................................ H. nigriabdomilis

- Arista plumose, the longest hairs longer than width of postpedicel; mid tibia with 3 to 4 p; hind femur with 3–4 strong av............................................................................................... H. annosa

30 Fore tibia without median p............................................................................ 31

- Fore tibia with median p............................................................................... 39

31 Inner vitta of scutum (between acr rows) reaching scutellum.................................................. 32

- Inner vitta of scutum (between acr rows) not reaching scutellum............................................... 33

32 Postpedicel about 4.0x as long as broad; hind tibia with 7– 8 ad..................................... H. setosiannosa

- Postpedicel about 3.0x as long as broad; hind tibia with 2 ad......................................... H. tuomuerra

33 Mid femur with pv only on basal half.................................................................... 34

- Mid femur with complete pv row........................................................................ 35

34 Frons subequal to anterior ocellus; hind tibia with 5–6 av; anterior margin of surstylus cavate............ H. sinoccidentala

- Frons about 1.5 x as long as anterior ocellus; hind tibia with 7–8 av; anterior margin of surstylus curved forward............................................................................................... H. gongshanensis

35 r-m crossvein with dark spot...................................................................... H. maae

- r-m crossvein without dark spot......................................................................... 36

36 Costal spine absent................................................................................... 37

- Costal spine distinct.................................................................................. 38

37 The longest hairs about 2x width of postpedicel; hind tibia with 5 av on apical half; tergite 5 with median orange vitta; length of surstylus at most 1.3x as long as broad..................................................... H. baoshanensis

- The longest hairs about 2.5x width of postpedicel; hind tibia with 5 av on middle part; each tergite without vitta; length of surstylus at least 2x as long as broad.......................................................... H. longisurstyla

38 Hind femur without pv, hind tibia with 13 strong p on middle part................................. H. septentrionalis

- Hind femur with 2–3 pv only on apical half, hind tibia with a row of setiform setae on basal half........... H. longievecta

39 Inner vitta of scutum (between acr rows) reaching scutellum.............................................. H. fica

- Inner vitta of scutum (between acr rows) not reaching scutellum............................................... 40

40 r-m crossvein with dark spot........................................................................... 41

- r-m crossvein without dark spot......................................................................... 43

41 Frontal setae 5 pairs............................................................................... H. shii

- Frontal setae 7–8 pairs................................................................................ 42

42 Arista plumose, the longest hairs subequal to postpedicel, costal spine absent............................... H. pedana

- Arista long plumose, the longest hairs about 1.7x width of postpedicel, costal spine distinct.............. H. platycephala

43 Arista plumose, the longest hairs subequal to postpedicel; hind femur with 4–5 av on apical 1/3; outer protuberance of cercus distinct................................................................................... H. nivimonta

- Arista long plumose, the longest hairs longer than postpedicel; hind femur with 5–6 av on apical half; cercus without outer protuberance.................................................................................. H. subfica