Helina subpubiseta, Xue, 1986

Li, Siyu, Xue, Wanqi, Nihei, Silvio S. & Zhang, Chuntian, 2025, Four new species of Helina Robineau-Desvoidy from far northwestern China (Diptera: Muscidae), Zootaxa 5719 (2), pp. 207-227 : 218-219

Key to Chinese species of Helina subpubiseta -group (males)

(modified from Xue & Sun 2015)

1 pra slightly shorter to longer than posterior notopleural seta.................................................... 2

- pra absent or hair-like, at most half as long as posterior notopleural seta......................................... 30

2 dc 2+3 .............................................................................................. 3

- dc 2+4 .............................................................................................. 8

3 Katepisternal setae 1+2................................................................................. 4

- Katepisternal setae 2+2................................................................................. 5

4 Femora and tibiae all yellow.............................................................. H. dicrocercacma

- Legs all black......................................................................... H. compositimacula

5 Fore tibia without median p....................................................................... H. altica

- Fore tibia with 1 median p.............................................................................. 6

6 Mid femur with complete pv row, sub-middle 3 strong............................................. H. harbinensis

- Mid femur without complete pv row, only with 2–3 pv on apical 1/3–1/4......................................... 7

7 Hind femur without pv.......................................................................... H. betula

- Hind femur with long pv on apical 1/3....................................................... H. qilianshanensis

8 Arista pubescent or short plumose, the longest hairs generally 1/2–2/3 of postpedicel width.......................... 9

- Arista long plumose, the longest hairs about equal to postpedicel width......................................... 18

9 Mid femur without av................................................................................. 10

- Mid femur with complete av row........................................................................ 15

10 Eye bare or with scarce ciliae; fore tibia with 1–2 median p................................................... 11

- Eye with dense and long ciliae; fore tibia without median p................................................... 14

11 Mid tibia with 1 ad......................................................................... H. tigrivestigia

- Mid tibia without ad.................................................................................. 12

12 Hind tibia anterodorsal and posteroventral surfaces with long hair row.............................. H. angulisternita

- Hind tibia anterodorsal and posteroventral surfaces without long hair row........................................ 13

13 Frons width equal to anterior ocellus; arista pubescent.............................................. H. platycerca

- Frons about 1.5x as wide as anterior ocellus; arista short plumose, the longest hairs about 1/2 as long as postpedicel width................................................................................................ H. solata

14 Mid tibia without ad, with 6–7 p; hind tibia with 6–8 pectinate av row on apical 3/5................... H. deslargideleta

- Mid tibia with 1 sub-median ad and 3 p; hind tibia with complete av row.......................... H. xianggelilaensis

15 Katepisternal setae 1+2...................................................................... H. subpubiseta

- Katepisternal setae 2+2................................................................................ 16

16 Arista pubescent.......................................................................... H. phantodonta

- Arista short plumose, about 1/2–2/3 as long as postpedicel width............................................... 17

17 Hind tibia with 2 av and 2 ad in the middle; pra slightly shorter than posterior notopleural seta; hind femur with complete av row..................................................................................... H. conicocerca

- Hind tibia with 4 av and 1 ad; pra as long as posterior notopleural seta; hind femur with 4–5 av on apical 2/5 H. spilopardalis

18 Katepisternal setae 1+2................................................................................ 19

- Katepisternal setae 2+2................................................................................ 20

19 Fore tibia without median p; mid femur without complete av row; hind tibia with a median row of p.......... H. subdeleta

- Mid femur with complete av row; hind femur only with av on basal 3/5; hind tibia with a submedian row of p............................................................................................ H. moirigkawagarboensis

20 Postpedicel about 1.3x as long as broad.......................................................... H. guizhenae

- Postpedicel 2.5–3.5x as long as broad.................................................................... 21

21 Hind femur with complete av row, the basal half with short bristle and the apical half developed...................... 22

- Hind femur with av only on basal 1/2.................................................................... 25

22 Mid femur with av only on basal 1/2..................................................................... 23

- Mid femur with only basal av........................................................................... 24

23 pra as long as posterior notopleural seta; mid tibia with 1 p........................................ H. minutideleta

- pra about 1.5x posterior notopleural seta; mid tibia with 3 p............................................. H. callia

24 Scutum inner vitta (between acr rows) not reaching scutellum; surstylus slightly longer than cercus.............. H. hyeta

- Scutum inner vitta reaching scutellum; surstylus significantly shorter than cercus......................... H. yaanensis

25 Legs black, except tibiae slightly brown to yellow on apical 1/2........................................... H. alea

- Legs black.......................................................................................... 26

26 Fore tibia without median p............................................................................ 27

- Fore tibia with 1 median p............................................................................. 28

27 Parafacial at the middle about 1.5x as wide as postpedicel; tergites 3–5 with conspicuous shifting patches on both sides; tergites 3 and 4 with a pair of large square dark patches................................................... H. largideleta

- Parafacial at the middle about 1.2x as wide as postpedicel; tergites 3–5 without conspicuous shifting patch on both sides; tergite 4 trace-like shape............................................................................... H. deleta

28 Scutum inner vitta (between acr rows) reaching scutellum............................................ H. meiguica

- Scutum inner vitta (between acr rows) at most reaching the middle of subscutum.................................. 29

29 Tergites 3 and 4 without conspicuous lateral patches; tergite 5 with a pair of small lateral patches.......... H. ampyxocerca

- Tergites 3 and 4 with a pair of brown patches in the middle........................................ H. paomashana

30 Katepisternal setae 1+2................................................................................ 31

- Katepisternal setae 2+2................................................................................ 33

31 Hind tibia without median p.......................................................... H. xinyiliia Xue sp. nov.

- Hind tibia with 1 median p............................................................................. 32

32 Eye bare; arista short plumose, the longest hairs about 2/5x as long as postpedicel; legs black............ H. flavipulchella

- Eye with sparse and long cilia; arista long plumose, the longest hairs 1.3–1.5x as long as postpedicel; legs black, except femora yellow on apex and hind tibia yellow on apical 4/5................................................ H. biastocerca

33 Fore tibia with 1 median p.................................................................... H. raoheensis

- Fore tibia without median p............................................................................ 34

34 Mid and hind tibiae yellow; eye bare........................................................... H. cilitruncata

- Legs black; eye with ciliae............................................................................. 35

35 Frons about 2x as long as postpedicel; dc 2+4................................................... H. xingkaiensis

- Frons slightly narrower than postpedicel; dc 2+3, the distance from the first seta to the second one large, and the distance from the second to third small, the first distance about 2.2x the second one...................... H. umbratiseta Xue sp. nov.