Otitoma Jousseaume, 1898

Kilburn, Richard N., 2004, The identities of Otitoma and Antimitra (Mollusca: Gastropoda: Conidae and Buccinidae), African Invertebrates 45, pp. 263-270 : 264-265

Otitoma Jousseaume, 1898

Otitoma Jousseaume, 1898: 106 . Type species (virtual tautonomy) Otitoma ottitoma Jousseaume, 1898 .

Thelecytharella Shuto, 1969: 208 (as subgenus of Euclathurella ). Syn. n. Type species (o.d.) Agladrillia oyamai Shuto, 1965 , Miocene of Philippines (and Japan).

[Junior synonyms of Thelecytharella Shuto, 1969 , were listed by Kilburn (1995: 262).]

Remarks: Although Powell (1966: 126) regarded Otitoma as a nomen dubium, its type species can now be recognised and proves to be a synonym of the widely distributed Pleurotoma cyclophora Deshayes, 1863 . Powell (1966) and others have referred P. cyclophora to Daphnella Hinds, 1844 , subgenus Hemidaphne Hedley, 1918 . However, the three­whorled protoconch is domed, with protoconch II smooth, except for a few arcuate axial riblets near its termination and extremely minute granular sculpture, and totally lacks the diagonally reticulate sculpture, characterisitic of the genus Daphnella . In overall characters, P. cyclophora agrees well with members of the group previously called Thelecytharella (see Kilburn 1995), over which Otitoma clearly has priority. The radula of this genus remains unknown.

Daphnella crenulata Pease, 1868 ( Fig. 14 ), is based on a juvenile of Otitoma cyclophora . Although Powell (1966) referred D. crenulata to Antimitra Iredale, 1917 , that taxon has been much­misunderstood, and in fact its type species, Pleurotoma aegrota Reeve, 1845 , shows no characters indicative of a turrid genus. The two syntypes ( Fig. 1 ) appear to be based on juveniles of a species of Metula Adams & Adams, 1853 , family Buccinidae .

Of described turrids, I refer the following Recent species, which have been distributed in the published literature amongst a wide range of different genera, to Otitoma . I have examined types of all species, with the exception of Hemidaphne gouldi for which

SEM’s are available. Literature references to these taxa are omitted, as these can be found in Tucker (2004). carnicolor Hervier, 1896, Drillia crenulata Pease, 1868 , Daphnella [= Pleurotoma cyclophora ] crokerensis Shuto, 1983, Austropusilla (Metaclathurella) cyclophora Deshayes, 1863 , Pleurotoma gouldi Yen, 1944 , Hemidaphne [synonym Mangelia pura ] kecil Sysoev, 1997, Thelecytharella kwandangensis Schepman, 1913 , Drillia lirata Reeve, 1845 , Pleurotoma metuloides Kilburn, 1995 , Thelecytharella mitra Kilburn, 1986 , Mitrellatoma oneili Barnard, 1958 , Drillia ottitoma Jousseaume, 1898 , Otitoma [= Pleurotoma cyclophora ]

pura Gould, 1860 (non Reeve, 1846), Mangelia [= Hemidaphne gouldi ] rubiginosa Hinds, 1843, Clavatula timorensis Schepman, 1913 , Drillia vitrea Reeve, 1845 , Pleurotoma . Although Otitoma ottitoma is synonymised with O. cyclophora , the following description is based on the ottitoma morph, primarily on series from Mozambique, which agree well with the neotype.

Figures

Figs 9–14. Otitoma cyclophora (Deshayes, 1863). 9–10. Type figures of Pleurotoma cyclophora (after Deshayes, 1863: pl. 12, figs 19, 21), Réunion Is. 11–12. Topotypes from Réunion Is., M. Jay, 8.0 x 2.4 mm and 6.3 x 2.3 mm (ESEM). 13. Protoconch and early whorls, scale­line = 500 mm. 14. Lectotype of Daphnella crenulata (designated Johnston, 1994), ANSP 15694, Polynesia [Howland Island, fide Johnston]; length 6.5 mm.

Fig. 1. Antimitra [= Metula] aegrota (Reeve, 1845): Syntype of Pleurotoma aegrota, BMNH 1963924, Singapore, 7 fath. [= approx. 13 m]; 10.5 x 3.7 mm.