Heterhelus, Jacquelin du Val, 1858

Hisamatsu, Sadatomo, 2011, A review of the Japanese Kateretidae fauna (Coleoptera: Cucujoidea), Acta Entomologica Musei Nationalis Pragae 51 (2), pp. 551-585 : 563

Key to subgenera and species of Japanese Heterhelus

1 Labrum shallowly notched. Antenna with distinct 3-segmented club ( Fig. 8E ). Pronotum strongly convex. Parameres of male genitalia abruptly bent inward at mid-length ( Fig. 8A ). Head dark brown, always darker than elytra. Subgenus Boreades Parsons, 1943 . .. ............................................................................................... H. (B.) solani ( Heer, 1841)

– Labrum deeply notched. Antenna with indistinct 3-segmented club ( Figs. 6O , 7D ). Pronotum feebly convex. Parameres of male genitalia regularly arcuate inwards ( Figs. 6A, 6D , 7A, 7E ). Head always the same color as elytra. Subgenus Heterhelus Jacquelin du Val, 1858 ......................................................................................................................... 2

2 Body uniformly black; sides of pronotum regularly arcuate, never angulate; parameres of male genitalia short and wide (LP/WP = 2.1 (n = 1)) ( Figs. 7A, 7E ); apex of median lobe bluntly rounded in lateral aspect ( Figs. 7C, 7F ); base of ovipositor pigmented ( Fig. 7G ). ........................................................................................... H. (H.) morio ( Reitter, 1878)

– Body reddish-yellow to blackish; sides of pronotum regularly arcuate or strongly angulate; parameres of male genitalia long and slender (LP/WP = 3.1 (n = 1)) ( Figs. 6A, 6D ); apex of median lobe sharply acuminate in lateral aspect ( Figs. 6C, 6E ); base of ovipositor not pigmented ( Fig. 6F ). ...................................................... H. (H.) scutellaris ( Heer, 1841)

Figures

Fig. 8. Heterhelus (Boreades) solani (Heer, 1841). A – tegmen (ventral view); B – median lobe (ventral view); C – median lobe (lateral view); D – ovipositor; E – right antenna, male; F – pronotum; G – left maxillary palpus.A–G = Hokkaido, Kawayu.

Fig. 6. Heterhelus (Heterhelus) scutellaris (Heer, 1841).A – tegmen (ventral view); B – median lobe (ventral view); C – median lobe (lateral view); D – parameres of the lectotype of H. japonicus; E – median lobe of the lectotype of H. japonicus (lateral view); F – ovipositor; G – male abdominal sternite VII; H–N – variation of pronotum (H – Hokkaido; I – Aomori; J – Niigata; K – Tokyo; L – Yamaguchi; M – Ehime; N – Fukuoka); O – male right antenna. A, B, C, G, O = Hokkaido, Hitsujigaoka; F = Ehime Pref., Omogokei.

Fig. 7. Heterhelus (Heterhelus) morio (Reitter, 1878). A – tegmen (ventral view); B – median lobe (ventral view); C – median lobe (lateral view); D – male right antenna; E – parameres of the lectotype of H. morio; F – median lobe of the lectotype of H. morio (lateral view); G – ovipositor; H – male abdominal sternite VII; I – pronotum. A, B, C, D, G, H, I = Ehime Pref., Omogokei.