Psammostiba hilleri ( Weise, 1877 ) Weise, 1877

Gusarov, Vladimir I., 2003, A revision of Nearctic species of the genera Adota Casey, 1910 and Psammostiba Yosii & Sawada, 1976 (Coleoptera: Staphylinidae: Aleocharinae), Zootaxa 185, pp. 1-35 : 32

Psammostiba hilleri ( Weise, 1877) , comb. nov.

( Fig. 5 in Sawada 1971; Figs. 18­19 , 25 in Brundin 1943; Figs. 34 , A­E in Yosii & Sawada 1976)

Homalota hilleri Weise, 1877: 90 .

Atheta (Metaxya) hilleri: Fenyes, 1920: 198 .

Atheta (Metaxya) hilleri: Bernhauer & Scheerpeltz, 1926: 614 . Atheta (Panalota) hilleri: Brundin, 1943: 20 .

Ischnopoda (Chaetida) multipunctata Sawada, 1971: 301 . Atheta (Psammostiba) hilleri: Yosii & Sawada, 1976: 82 . Atheta (Psammostiba) multipunctata: Yosii & Sawada, 1976: 82 (as synonym of At. hilleri ). Atheta (Psammostiba) hilleri: Sawada, 1977: 171 .

Diagnosis. Psammostiba hilleri can be recognized by smooth posterior margin of male tergum 8, and by the distinct shape of the aedeagus ( Figs. 34 , A­E: Yosii & Sawada 1976) and spermatheca ( Fig. 25 : Brundin 1943).

Psammostiba hilleri differs from P. comparabilis and P. kenaii by lack of crenulation of the posterior margin of male tergum 8.

Discussion. Brundin (1943) based his redescription of P. h i l l e r i on a single female type from Hagi, Japan. His description of P. jessoensis ( Brundin 1943) was also based on a single female type from Hokkaido, Japan. Apparently, Yosii and Sawada (1976) did not examine the types of these two species and based their interpretation entirely on Brundin's descriptions and on assumption that P. hilleri is a southern species, while P. jessoensis occurs in Northern Honshu and Hokkaido. However, the female spermatheca of P. h i l l e r i illustrated by Sawada (1971: Fig. 5 , M) does not match Brundin's drawing (1943: Fig. 25 ). To confirm the status of P. h i l l e r i and P. jessoensis the types of both species and additional samples of Psammostiba from different localities in Japan need to be examined.

Distribution. Psammostiba hilleri is known from Japan (central and southern Honshu and Kyushu).

Figures

FIGURES 1 ­ 5. Mouthparts of Adota maritima (Mannerheim) (male from Homer, Alaska). 1 – labrum; 2 – epipharynx; 3 – left mandible, dorsal view; 4 – left mandible, ventral view; 5 – right mandible, dorsal view. Scale bar 0.1 mm.

FIGURES 13 ­ 18. Details of Adota maritima (Mannerheim) (males (13, 15 ­ 16, 18) and female (14) from Homer, Alaska) and Ad. gnypetoides Casey (17; male from Bodega Head, California). 13 – meso­ and metathorax, ventral view; 14 – pronotum; 15 – right metatarsus; 16 – metatarsal segment 5 and metapretarsus, dorsal view; 17 ­ 18 – right antenna. Scale bar 0.1 mm (16), 0.2 mm (15), 0.4 mm (13 ­ 14, 17 ­ 18).

FIGURES 19 ­ 22. Abdominal segment 8 of Adota maritima (Mannerheim) (males (19 ­ 20) and female (21 ­ 22) from Homer, Alaska). 19 – male tergum 8; 20 – male sternum 8; 21 – female tergum 8; 22 – female sternum 8. Scale bar 0.2 mm.

FIGURES 23 ­ 30. Genitalia of Adota maritima (Mannerheim) (males (23 ­ 28) and female (29 ­ 30) from Homer, Alaska). 23 – median lobe, parameral view; 24 – apex of median lobe, parameral view; 25 – median lobe, lateral view; 26 – apex of median lobe, lateral view; 27 – copulatory piece, parameral view; 28 – apex of left paramere; 29 ­ 30 – spermatheca. Scale bar 0.1 mm (24, 26 ­ 28), 0.2 mm (23, 25, 29 ­ 30).

FIGURES 31 ­ 34. Abdominal segment 8 of Adota gnypetoides Casey (male (31 ­ 32) and female (33 ­ 34) from Bodega Head, California). 31 – male tergum 8; 32 – male sternum 8; 33 – female tergum 8; 34 – female sternum 8. Scale bar 0.2 mm.