Rafaelomyia uniamazonia Ramos-Pastrana & Soares, 2023

Soares, Matheus M. M., Capellari, Renato S. & Ramos-Pastrana, Yardany, 2023, Rafaelomyia, a remarkable new genus of South American long-legged flies (Diptera, Dolichopodidae), Zootaxa 5389 (2), pp. 151-172 : 163-166

publication ID	https://doi.org/ 10.11646/zootaxa.5389.2.1
publication LSID	lsid:zoobank.org:pub:CDC5E9AC-B029-4D3F-B4C9-75EC018BA9C3
DOI	https://doi.org/10.5281/zenodo.10419054
persistent identifier	https://treatment.plazi.org/id/038687B7-5E2A-6D32-FF33-4321FE8D0710
treatment provided by	Plazi
scientific name	Rafaelomyia uniamazonia Ramos-Pastrana & Soares
status	sp. nov.

Treatment

Rafaelomyia uniamazonia Ramos-Pastrana & Soares View in CoL sp. nov.

( Figs 7 View FIGURE 7 , 8 View FIGURE 8 )

Diagnosis (males). Body length 6 mm ( Fig. 7A View FIGURE 7 ); femora mostly yellow, except dorsal and ventral surfaces of femora I and II darkened and apical 1/3 of femur III brown to dark brown ( Fig. 7A View FIGURE 7 ); wing dark brown, vein R 2+3 not thickened ( Fig. 7F View FIGURE 7 ); outer branch of cercus with subtriangular, weakly sclerotized and bare projection near apex ( Fig. 8B, C View FIGURE 8 ); hypandrium with a short concavity and two very short dentiform projections at apex ( Fig. 8G, H View FIGURE 8 ); phallus abruptly narrowing at apex, with wide and lobulated preapical processes ( Fig. 8E, F View FIGURE 8 ).

Description. Male ( Fig. 7A View FIGURE 7 ). Body length: 6 mm; wing length: 6 mm, width: 2 mm (n = 1). Head ( Fig. 7B–D View FIGURE 7 ). Face as wide as ocellar tubercle, eyes almost parallel sided, only slightly narrowing at middle of face. Palpus oval, short, black, covered with short stout setae and yellowish pruinosity, with tuff of long brown setae at apical 1/2. Thorax ( Fig. 7A, B View FIGURE 7 ). Mesonotum mostly metallic dark green with weak bluish reflections. Wing ( Fig. 7F View FIGURE 7 ). Membrane and veins dark brown. R 4+5 nearly straight, slightly curved posteriorly near apex; M 1 nearly straight and almost parallel sided with R 4+5, slightly curved posteriorly at apex; CuA+CuP long, ending at basal 1/2 of cell bm+dm. Legs ( Fig. 7A View FIGURE 7 ). Mostly brown to dark brown, except apical 1/2 of lateral surface yellow of coxa I, all trochanter, anterior and posterior surfaces of femora I and II, basal 1/2 and basal 2/3 of anterior surface of femur III yellow. Leg I. Podomere ratios: 69, 65, 36/16/11/9/11. Tibia I with 3 strong anterodorsal setae at: 1.5/6, 3.5/6 and apex, anterodorsal row of short setae from basal 2/6 to apex, about as long as width of tibia, ventral row of long slender setae from basal 1.5/6 to apex, about 1.5X longer than width of tibia ( MSSC). Leg II. Podomere ratios: 79, 80, 50/34/23/12/12.Anterior surface of coxa II covered with vestiture of long setae and dark silvery pruinosity, outer edge with 3 long setae, 2 near middle and 1 near apex, inner edge with row of long setae, apical edge with long setae. Femur II with row of 6 long setae increasing in length at apex ventrally. Tibia II with 2 stout anterodorsal setae at: 1.5/6 and 3.5/6, 3 stout posterodorsal setae at: 1/6, 2/6 and 3/6, and crown of strong setae at apex: 1 antero- and 1 posterodorsal, 1 dorsal, 1 posteroventral and 1 ventral long seta, about 1/2 as long as IIt 1. Leg III. Podomere ratios: 105, 105, 30/45/28/18/15. Femur III with dorsal row of long erect setae at basal 1/2, slightly longer than width of femur at broadest point, 1 long dorsal seta at 4.5/6, 4–5 strong anteroventral setae at apical 1/3. Abdomen ( Fig. 7A View FIGURE 7 , not dissected). Mostly dark brown, with weak greenish reflections, tergites 6, 7 and sternite 8 darker. Hypopygium ( Figs 7E View FIGURE 7 , 8 View FIGURE 8 ). Pale brownish, somewhat triangular. Cercus mostly yellow, distinctly longer than epandrial capsule, divided into two branches ( Figs 7E View FIGURE 7 , 8A View FIGURE 8 ). Outer branch of cercus longer, somewhat digitiform, with rounded apex, slightly bent ventrally at apical 1/2 with inner wide, subtriangular, weakly sclerotized and bare projection near apex; outer surface covered with long strong setae, longer at apex of dorsal edge, apical 1/4 of ventral edge and inner surface covered with a few short, slender yellow setae ( Fig. 8B, C View FIGURE 8 ). Inner branch of cercus short, slender, somewhat digitiform, about 1/2 as long as outer branch, dorsal surface covered with short erect setae, longer at apex ( Fig. 8B, C View FIGURE 8 ). Surstylus somewhat triangular, with acute apex, about 1/4 as long as epandrial capsule, inner edge with 3 short slender setae, 1 near base, 2 near middle, and 1 stout seta dorsally at apex ( Fig. 8A, D View FIGURE 8 ). Hypandrium slightly curved ventrally at apex, about 2.5X longer than wide, with short concavity and two very short dentiform projections at apex ( Fig. 8G, H View FIGURE 8 ). Phallus slightly longer than epandrial capsule, widening near base, and abruptly narrowing at apex, with wide and lobulated preapical processes ( Fig. 8E, F View FIGURE 8 ). Postgonite noose-shaped, surrounding base of phallus, fused to hypandrial arms ( Fig. 8E View FIGURE 8 ). Ejaculatory apodeme short and somewhat digitiform, well sclerotized and narrowing towards apex ( Fig. 8E View FIGURE 8 ).

Female. Unknown.

Type examined material. HOLOTYPE ♂ ( Fig. 8A View FIGURE 8 ) labelled as: “ COLOMBIA: Huila, Garzón | Vda.[Vereda] Las Mercedes , Parque Regional Natural | Regional Cerro Páramo de Miraflores , | 02°08′06″N 75°30′04″W, 2.074 m | 24.xi–08.xii.2022 Malaise, Ramos-Pastrana, Y. (LEUA)” “HOLOTYPE | Ramos-Pastrana & Soares [red label]” Holotype condition: Good , terminalia dissected and stored in microvial, left wing glued to paper triangle. GoogleMaps PARATYPE: Same data as holotype except, 14.i.–2.ii.2023 (1 ♂, LEUA) .

Remarks. Rafaelomyia uniamazonia sp. nov. is remarkably similar to R. inpa sp. nov. as discussed above (see remarks under R. inpa sp. nov.), but can be differentiated by the larger size 6.0 mm ( Fig. 7A View FIGURE 7 ), wing membrane dark brown ( Fig. 7F View FIGURE 7 ) and apex of phallus with a wide preapical lobulated process ( Fig. 8E, F View FIGURE 8 ).

Etymology. Rafaelomyia uniamazonia sp. nov. is named after the shorthand name for the Universidade de la Amazonia in Valencia, Colombia.

Distribution. The new species is known to occur only from the type locality, at the district of Huila in Colombia ( Fig. 13B View FIGURE 13 ).