Promitobates

Bragagnolo, Cibele & Pinto-Da-Rocha, Ricardo, 2012, Systematic review of Promitobates Roewer, 1913 and cladistic analysis of Mitobatinae Simon, 1879 (Arachnida: Opiliones: Gonyleptidae), Zootaxa 3308, pp. 1-48 : 12-13

Key to males of Promitobates species

1 External apical apophysis of coxa IV single-branched ( Fig. 11 A,B).............................................. 2

- External apical apophysis of coxa IV with two dorsal branches ( Figs. 4 , 5 )........................................ 3

2 Ocularium armed with a pair of low tubercles ( Fig. 8 C).............................................. P. a l e sp. n.

- Ocularium armed with a pair of high spiniform apophysis ( Fig. 8 D).................................. P. trapista sp. n.

3 Posterior margin of dorsal scutum armed with a pair of tubercles or with median spiniform apophysis ( Figs. 4 ; 5A,B)...... 4

- Posterior margin of dorsal scutum with a row of tubercles of the same size ( Fig. 5 C,D)................ P. viridigranulatus

4 Apex of coxa and trochanter IV and base of femur IV black, standing out from the rest of body coloration; area IV of dorsal scutum totally covered by tubercles (without unarmed region) ( Fig. 6 C)............................ P. weissbier sp. n.

- Apex of coxa and trochanter IV and base of femur IV of similar coloration to the rest of leg segments, or slightly blackish, but never standing out from the rest of body coloration; area IV with unarmed region, not entirely covered by tubercles ( Figs. 4 , 5 , 6 A,B,D)............................................................................................. 5

5 Area I of dorsal scutum with few tubercles (less than 10), scattered; ocularium sulphurous yellow, standing out from the rest of body ( Fig. 4 A, 17F)............................................................................. P. bellus

- Area I of dorsal scutum densely tuberculated or with tubercles along the groove II (roughly in rows); ocularium coloration not standing out from the rest of body (of same color or slightly yellowish) ( Figs. 4 B–D, 5; 6)........................... 6

6 Areas III and IV of dorsal scutum densely tuberculated, except around apophyses; prosoma with longitudinal row of tubercles at sides of ocularium ( Fig. 6 D)................................................................. P. lager sp. n.

- Areas III and IV with tubercles concentrated on median region (between the apophyses) and/or on the grooves III–V; prosoma without longitudinal row of tubercles at sides of ocularium ( Fig. 4 C, D).......................................... 7

7 Areas III and IV with a row of tubercles up to the posterior groove; tubercles of dorsal scutum yellowish, with circular patches of same color around them ( Fig. 4 C)........................................................... P. hatschbachi

- Areas III and IV with tubercles concentrated on median region; tubercles of dorsal scutum sulphurous yellow, without circular patches around them ( Figs. 4 D, 15B)..................................................................... 8

8 Dorsal scutum black tiger-like stripes; coxa IV densely tuberculated, and the tubercles as tall as wide ( Figs. 4 B, 15A).................................................................................................. P. difficilis

- Dorsal scutum not black tiger-like stripes; coxa IV with few tubercles (less than ten), the tubercles taller than wide ( Figs. 4 D, 5A,B)............................................................................................... 9

9 Free tergite III with median spiniform tubercle ( Fig. 5 A).............................................. P. nigripes

- Free tergite III unarmed on median region ( Fig. 5 B)......................................................... 10

10 Area IV of dorsal scutum with transverse row of tubercles up to groove V ( Fig. 4 D)..................... P. intermedius

- Area IV of dorsal scutum without transverse row of tubercles up to groove V ( Fig. 5 B)...................... P. o r n a t u s

Figures

FIGURE 4. Dorsal view of males of Promitobates. A. P. bellus, (IBSP 3306); B. P. difficilis (MNRJ 11395, syntype); C. P. hatschbachi (MHNCI 0 164, holotype; D. P. intermedius (MZSP 14410). Scale bar, 1 mm.

FIGURE 5. Dorsal view of males of Promitobates. A. P. n i gr i p e s, (MZSP 27425); B. P. o r n a t u s (MZSP 27417); C and D. P. viridigranulatus (MZSP 27401), intraspecific variation. Scale bar, 1 mm.

FIGURE 6. Dorsal view of males of Promitobates A. P. a l e sp. n. (MZSP 27404); B. P. trapista sp. n. (MZSP 27406); C. P. wiessbier sp. n. (MZSP 16171); D. lager sp. n. (MNRJ 4545). Scale bar, 1 mm.

FIGURE 8. Lateral view of males of Promitobates. A and B. P. viridigranulatus, (A. MZSP 27401; B. MHNCI 6527); C. P. ale sp. n. (MZSP 27404); D. P. trapista sp. n. (MZSP 27406); E. P. weissbier sp. n. (MZSP 16171); F. P. l a g e r sp. n. (MNRJ 4545). Scale bar, 1 mm.

FIGURE 11. Dorsal view of females of Promitobates. A. P. bellus (MZSP- 19509); B. P. dificillis (MNRJ- 11395, syntype); C. P. hatschbachi (MZSP 18811); D. P. intermedius (MZSP- 16764); E. P. n i g r i p e s (MZSP - 31029); F. P. ornatus (MZSP 27440). Scale bar = 1 mm.