Eugaedioxenis Cerretti, 2015

Cerretti, Pierfilippo, O’Hara, James E., Winkler, Isaac S., Lo Giudice, Giuseppe & Stireman, John O., 2015, Two tribes hidden in one genus: the case of Agaedioxenis Villeneuve (Diptera: Tachinidae: Exoristinae), Organisms Diversity & Evolution (New York, N. Y.) 15 (3), pp. 489-512 : 494

Eugaedioxenis Cerretti , O’ Hara and Stireman, gen. nov.

( Figs. 1a–d ; 2e–h ; 4a–b, f ; 5a–b ; 6f ; 7a–d)

http://zoobank.org/ urn:lsid:zoobank.org:act:FE7A5AB9-74D6-4627-8696-39D2398432E2

Type species. Gaedioxenis haematodes Villeneuve, 1937 , by present designation.

Diagnosis. Medium to large size, strongly bristly, flies characterized by the following combination of character states: Compound eye covered with scattered, long ommatrichia; parafacial covered with fine setulae; facial ridge with decumbent setae on lower 1/4–1/2; lower facial margin at least slightly visible in lateral view; prementum long and slender; 5 postpronotal setae, the 3, strongest, basal setae arranged in a line; scutum with 3 postsutural dorsocentral setae; katepimeron bare; first postsutural supra-alar seta longer than notopleural setae; preapical anterodorsal setae of fore tibia at least as long and strong as preapical dorsal seta; hind tibia with 4 strong preapical setae; basicosta yellow; mid-dorsal depression on abdominal syntergite 1+2 not extended back to hind margin of syntergite; male with 2 strong proclinate orbital setae; female with tergite 5 long, conical, 1.1–1.3 times as long as tergite 4; oviscapt slightly elongated and telescopically retracted within segment 5 ( Fig. 7); egg macrotype, membranous, fully embryonated. No remarkable sexual dimorphism has been observed between male and female in head chaetotaxy, frons width, compound eye size or postpedicel length.

Etymology. The generic name is a composite word formed from eu, meaning true, and the tachinid genus-group name Gaedioxenis Townsend.

Hosts. Unknown.

Distribution. South Africa.

Figures

Fig. 1 a–b Eugaedioxenis gen. nov. haematodes (Villeneuve) habitus in lateral view: a male (holotype); b female. c–d Eugaedioxenis horridus gen. et sp. nov. habitus in lateral view: c male (holotype); d female

Fig. 2 a Agaedioxenis brevicornis (Villeneuve), male, habitus. b Agaedioxenis kirkspriggsi sp. nov., male (holotype) habitus. c Agaedioxenis succulentus sp. nov., male (holotype) habitus. d Agaedioxenis timidus sp. nov., male (holotype) habitus.

Fig. 4 a–c Female abdomen: a Eugaedioxenis gen. nov. haematodes (Villeneuve); b Eugaedioxenis horridus gen. et sp. nov. c Agaedioxenis succulentus sp. nov. d–e Male abdomen: d Agaedioxenis kirkspriggsi sp. nov. (holotype); e Agaedioxenis timidus sp. nov. (holotype). f–g Male

Fig. 5 a–b Eugaedioxenis gen. nov. haematodes (Villeneuve), epandrial complex in posterior (a) and lateral (b) view. c–d Agaedioxenis setifrons Villeneuve, epandrial complex in posterior (c) and lateral (d) view. e–f Agaedioxenis brevicornis (Villeneuve), epandrial complex in

Fig. 6 a–b Agaedioxenis succulentus sp. nov. (holotype), epandrial complex in posterior (a) and lateral (b) view. c–d Agaedioxenis kirkspriggsi sp. nov. (holotype), epandrial complex in posterior (c) and lateral (d) view. Scale bar 0.2 mm. e Agaedioxenis brevicornis (Villeneuve), male terminalia in ventral view. f Eugaedioxenis gen. nov. haematodes (Villeneuve), male terminalia in ventral view. g Agaedioxenis kirkspriggsi sp. nov. (holotype), phallus, phallapodeme and ejaculatory