Agaedioxenis Villeneuve, 1939

Cerretti, Pierfilippo, O’Hara, James E., Winkler, Isaac S., Lo Giudice, Giuseppe & Stireman, John O., 2015, Two tribes hidden in one genus: the case of Agaedioxenis Villeneuve (Diptera: Tachinidae: Exoristinae), Organisms Diversity & Evolution (New York, N. Y.) 15 (3), pp. 489-512 : 502-504

publication ID	https://doi.org/ 10.1007/s13127-015-0211-0
publication LSID	lsid:zoobank.org:pub:0C53D317-5C05-45D7-A9E4-FB6E6D6F28DE
persistent identifier	https://treatment.plazi.org/id/038587E4-FFB6-2942-FCC1-FB76FDB8CE0F
treatment provided by	Felipe
scientific name	Agaedioxenis Villeneuve, 1939
status	stat. nov.

Treatment

Agaedioxenis Villeneuve, 1939 View in CoL View at ENA , stat. nov.

( Figs. 1e–f View Fig , 2a–d View Fig , 3a–k View Fig , 4c–e, g View Fig , 5c–f View Fig , 6a–e, g View Fig , 8a–d View Fig )

Gaedioxenis Villeneuve, 1937: 206 View in CoL . Nomen nudum (proposed after 1930 without designation of type species from two included species).

Gaedioxenis Villeneuve, 1939: 1 View in CoL . Nomen nudum (proposed after 1930 without designation of type species from two included species).

Agaedioxenis Villeneuve, 1939: 2 View in CoL (as subgenus of Gaedioxenis Villeneuve, 1937 View in CoL [nomen nudum]). Type species: Gaedioxenis (Agaedioxenis) brevicornis Villeneuve, 1939 View in CoL , by monotypy.

Gaedioxenis Townsend, 1943: 335 . Type species: Gaedioxenis setifrons Villeneuve, 1937 , by original designation.

Review of genus-group names. Villeneuve (1937) proposed the new genus Gaedioxenis for two new species, G. setifrons and G. haematodes . Later, Villeneuve (1939) added two new species to this genus, Gaedioxenis propinqua and G. brevicornis , placing the latter in a new subgenus, Agaedioxenis . The type species of the genus-group name Agaedioxenis was thus fixed by monotypy as G. (A.) brevicornis . In neither of Villeneuve’ s papers was a type species fixed for Gaedioxenis from among the two species described in 1937 or the two species available for type species fixation in 1939 ( G. setifrons and G. propinqua ; G. haematodes was not mentioned in this work and G. brevicornis was fixed as type species of Agaedioxenis ). Townsend (1943: 335) attempted to remedy this situation by designating G. setifrons as the type species of Gaedioxenis (by “designation herein”). Crosskey (1980: 878) followed this interpretation, attributing authorship of Gaedioxenis to Villeneuve, 1937 and the type species designation to Townsend (1943) [as 1941, treating the “Addenda and corrigenda” to Part XI of Townsend’ s (1941) Manual of Myiology as published at the same time as the rest of the volume].

Evenhuis et al. (2008) discovered that earlier interpretations of Gaedioxenis were contrary to the Code that required the description of a new genus after 1930 to “be accompanied by the fixation of a type species in the original publication” (Article 13.3 of ICZN 1999). Thus, Gaedioxenis was unavailable from Villeneuve (1937, 1939) and took authorship and date from Townsend (1943). This interpretation was discussed in more detail by Evenhuis et al. (2015).

We agree with Evenhuis et al. (2008) and Evenhuis et al. (2015) that Gaedioxenis is correctly attributed to Townsend (1943), but we do not treat this name as valid. The genus-group name Agaedioxenis Villeneuve, 1939 has priority over Gaedioxenis Townsend, 1943 and is the valid name of the genus when these names are treated as subjective synonyms, as they are herein.

Diagnosis. Medium to large size, moderately bristly, exoristine characterized by the following combination of character states: Compound eye bare; parafacial covered with fine setulae; facial ridge with decumbent setae approximately on lower 1/4; lower facial margin protruded and clearly visible in lateral view; prementum long and slender; 4 or 5 postpronotal setae, the 3, strongest, basal setae arranged in a line; scutum with 4 postsutural dorsocentral setae; katepimeron bare or at most with 1–2 hair-like setulae on anterior 1/4; first postsutural supra-alar seta longer than notopleural setae; preapical anterodorsal setae of fore tibia at least as long and strong as preapical dorsal seta; hind tibia with 3 strong dorsal preapical setae; basicosta brownish black; mid-dorsal depression on abdominal syntergite 1+2 extended back to hind margin of that segment; male without proclinate orbital setae; female with tergite 5 normally developed, sub-conical, 0.8–0.9 times as long as tergite 4; egg microtype, planoconvex, fully embryonated; convex side hard-shelled, blackish brown, with 1 subcircular aeropilar area attached to anterior margin. Sexual dimorphism involves head chaetotaxy and morphometric ratios of head; specifically females have proclinate and lateroclinate orbital setae, a wider frons and in some cases also a shorter postpedicel.

Hosts. Unknown.

Distribution. South Africa, Zimbabwe.