Madagaster procarina, Perkins, 2017

Perkins, Philip D., 2017, Hydraenidae of Madagascar (Insecta: Coleoptera), Zootaxa 4342 (1), pp. 1-264 : 188-191

Madagaster procarina , new species

Figs. 180 (habitus), 182 (aedeagus), 301 (map)

Type Material. Holotype (male): Fianarantsoa, 32 km S Ambositra , in seepage over exposed granite cliff face, elev. 1400 m, 20° 43' S, 47° 16' E, 25 iii 1990, W. E. Steiner, C. Kremen, V. Razafimahatratra ( USNM) . Paratypes (26): Same data as holotype (2 USNM). Antsiranana, Diana: Antsaba : Galoko mountains , 3.4 km NW from Anstaba, S 13.60931 E 48.72129, aspirator, forceps, sieves: hygropetric rocks and pools, elev. 296 m, 28 xi 2012, J. Bergsten, R. Bukontaite, J.H. Randriamihaja & T. Ranarilalatiana ( MAD12-31 ) (6 NHRS) ; Fianarantsoa, Amoron'i Mania, 3km south of Ambalamanakana next to RN7, Ankazomivady forest , 20.7722S 47.1809E, 1700m, hygropetric rocks and marsh with vegetation, elev. 1700 m, 1 xi 2014, J. Bergsten, T. Ranarilalatiana & S. Holmgren ( MAD14-02 ) (1 NHRS); Andringitra NP hygropetric cascade stream crossing way to cascades, 16.2°C, elev. 1722 m, 22° 8' S, 46° 53' E, 11 iv 2011, R. Gerecke (MD 198) (7 NMW); Isalo Canyon de Makis , sandy bottom of river, with side pools and hygropetric sections at sides, wood in water, N-22° 32.922, E45° 24.064, elev.

780 m, 22° 33' S, 45° 24' E, 11 v 2006, Bergsten et al. (P41E) (8 NHRS); Ihorombe , Isalo NP 50m before the entrance of Canyon de Makis, GB nets and sieves, canyon river with side pools, S22.48660, E45.37954, elev. 752 m, 22° 33' S, 45° 24' E, 13.xi.2012, R. Bukontaite & J. H. Randriamihaja ( MAD12-05 ) (1 NHRS) ; Antsaba , Galoko Mountains, hygropetric cascade, elev. 303 m, 13° 39' S, 48° 45' E, 1 xi–29 xii 2004, M. Balke (MD 016) (on female " MNCN AI546 DNA specimen").

Differential Diagnosis. Similar in habitus to M. barbata and M. bergsteni ( Figs. 179, 180, 183 ). Differentiated from both species by the low midlongitudinal carina on the prosternum, the straight meso- and metatibiae, and the densely setose medial margin of the metafemur. The very complex male genitalia of the three species share a hookshaped process on the ventral surface of the main piece, but the parameres and processes of the distal part of the main piece differ markedly ( Figs. 181, 182, 185).

Description. Size: holotype (length/width, mm): body (length to elytral apices) 2.23/1.02; head width 0.60; pronotum 0.55/0.86, PA 0.69, PB 0.68; elytra 1.38/1.02.

Dorsum and venter black, legs dark brown, ocelli light brown to testaceous.

Entire dorsum dull, with very small, closely spaced tubercles, each with a short recumbent seta. Labrum bilobed, anterior margin slightly reflexed, sparsely granulate and with sparse small setae except with longer setae along anterior margin. Labrum originating at anterior, slightly overhanging, margin of clypeus. Labroclypeal suture moderately deep. Short, deep sulcus in front of each ocellus. Frons with deep concavity between ocelli. Maxillary palpi (male) ca. as along as antenna, with last two palpomeres slightly enlarged. Antenna with penultimate antennomere larger than ultimate.

Pronotum with two narrowly separated midlongitudinal fovea, anterior ca. twice as long as posterior; each side with two admedian and two sublateral foveae; transverse fovea along base; lateral margin rounded.

Elytral suture costate in posterior. Each elytron with four non-interrupted costae; first costa (from suture) slightly curved in ca. middle ½ of length. Each costa with unilinear row of granules. Areas between costae with four irregular rows of granules. Elytral apices truncate.

Mentum and postmentum densely micropunctulate, sparsely pubescent; Genae micropunctulate, dull, concave behind cardo, with very strong, high posterior ridge, obliquely oriented on each side of midline. Subocular groove for antenna moderately deep and wide. Postocular margin not carinate.

Prosternum with midlongitudinal carina. Mesoventrite without tubercle on intercoxal process. Metaventrite densely micropunctulate and with very short and very dense vestiture; midlongitudinal depression moderately deep ( Fig. 180). First four abdominal ventrites and anterior margin of fifth with very dense, recumbent pubescence (plastron?); setae sparser in midline than laterally. Last ventrite (male only) oval, and with dense pubescence, setae longer at margin forming oval rim.

Pro- and metatibiae slightly arcuate, toward midline. All tibiae with brush of setae apically. Metafemur with brush of setae along lower margin in basal ½.

Female length (ca. 2.09–2.39 mm) sometimes greater than males, but form narrower (less robust).

Etymology. Named in reference to the midlongitudinal carina of the prosternum.

Figures

FIGURE 183. Madagaster barbata, holotype, frontal view, habitus dorsal, lateral, and ventral.