Scleropactes colombiensis, (PEARSE, 1915)

Schmidt, Christian, 2007, Revision of the Neotropical Scleropactidae (Crustacea: Oniscidea), Zoological Journal of the Linnean Society 151, pp. 1-339 : 42-44

publication ID	https://doi.org/ 10.1111/j.1096-3642.2007.00286.x
persistent identifier	https://treatment.plazi.org/id/03858799-4215-FFC1-9817-7C8BABDDFB59
treatment provided by	Felipe
scientific name	Scleropactes colombiensis
status

Treatment

SCLEROPACTES COLOMBIENSIS ( PEARSE, 1915) View in CoL

Sphaeroniscus colombiensis Pearse, 1915 View in CoL . Scleropactes columbiensis – comb. Schultz (1970) (specific name misspelled).

Scleropactes colombiensis View in CoL – Schmalfuss (1980, 1986); Leistikow & Wägele (1999 *); Schmalfuss (2003 *).

Material examined

Types: One ♀ m, broken into pieces [holotype, Colombia, Sierra Nevada de Santa Marta, Cincinnati Coffee Plantation, altitude 4800 feet (11°02′ N, 74°02′ W, altitude c. 1500 m), leg. A. S. Pearse, 12 July 1913, USNM 98374 View Materials ] GoogleMaps .

Other samples: One ♂ ( Colombia, Sierra Nevada de Santa Marta, San Lorenzo , altitude 1600 m, cloud forest, leg. H. Schmalfuss, 8 December 1974, SMNS 10048 View Materials e + slides); one ♀ m ( Colombia, Magdalena, Sierra Grande de Santa Marta , near El Campano, KT- 12, altitude 1000 m, from leaf litter, leg. H. G. Müller, 20 April 1986, SMF 19377) .

Description ( Figs 106–111 View Figure 106 View Figure 107 View Figure 108 View Figure 109 View Figure 110 View Figure 111 )

Adult female 16 mm long, 5.5–5.8 mm wide, cephalothorax 2.92 mm wide; male 5.8 mm wide, cephalothorax 2.90 mm wide. Dark brown with pale muscle insertion spots, coxal plates lighter (in the female the colour was not preserved).

Eyes composed of 20–23 ommatidia. Cephalothorax with deep furrow behind frontal line. Frontal shield with shallow grooves for second antennae; a medial area is distinctly set off the grooves. Small lateral lobes projecting about half of length of the eyes. Tergites smooth. All coxal plates simple. Tergal scale setae small and inconspicuous. Noduli laterales small, on posterior margin of tergites, seemingly at equal distance from the lateral margin, but this could not be clearly seen in the entire specimen.

First antenna three-jointed, second joint shortest. Distal joint with a pair of apical aesthetascs, and about 12 slightly smaller, irregularly arranged subapical aesthaescs.

Second antenna approximately as long as pereiopod 7, but more slender. Flagellum consisting of three subequal articles. Second and third articles have a transverse row of two or three aesthetascs, the third bears a slender apical cone. Apical cone somewhat shorter than distal flagellar article, with a pair of small lateral sensilla at 0.25 from its base (the sensilla are broken off, but their insertions are visible).

Left mandible with long pars incisiva apically indistinctly four-cusped, lacinia mobilis with two cusps, larger than pars incisiva. Hairy lobe with two hairy setae, and one hairy seta between hairy lobe and pars molaris that is represented by a tuft of indistinctly separated hairy setae. Right mandible with short, apically indistinctly four-cusped pars incisiva, small, irregular conical lacinia mobilis, hairy lobe with one hairy seta, and one hairy seta between the hairy lobe and the pars molaris, which is represented by (probably) a single hairy seta (?). First maxilla lateral endite on apical margin with lateral group of four stout, simple tooth setae, a small triangular lobe and one slen- der seta, which is apically hirsute, and with mesal group of six more slender, simple tooth setae. A pair of very small setae beside the mesal group of tooth setae on the caudal face. Mesal endite with two stout penicils, the laterodistal corner rounded and covered with small hairs. The examined specimen has two penicils on mesal endite of one side, and two penicils of same size and one smaller penicil of about half the size on the other side. Second maxilla apically bilobate, both lobes hairy, the mesal lobe with a field of sensilla along its mesodistal margin. Two (or three?) small sensilla between both lobes. Maxilliped base with scales and scale-setae. Palp three-jointed, proximal article bearing one large and one smaller seta; second article one broad and several slender setae on lateral margin, and a tuft of setae on a long socket on distal end of mesal margin. There are two single setae beside the latter socket, and the proximal tuft of setae present in other Crinocheta here is represented by a single seta, located near the socket of the distal tuft. Distal article with apical tuft of setae, and several setae on the lateral margin. Maxilliped endite distally somewhat narrower than basally, and distally covered with hairs; one large penicil is located on the frontal face of the endite, near the distal margin.

Male pereiopods 1–3 with ventral scale-brushes on merus and carpus; on pereiopod 3 these are much less extended than on pereiopods 1 and 2. Pereiopod 1 with antennal cleaning brush consisting of a small field of scales (or spines) on propodus and a long field with acute, tongue-shaped scales on carpus. Distal margin of carpal field concave and provided with a fringe of long scales with rounded tips. A smaller field also present on pereiopod 2, and a small group of scales can also be seen on pereiopod 3 near the distal margin of the carpus. Surface of all pereiopods covered with scales, which appear to be more regularly arranged on basipodites than on more distal parts. Large setae are found on ventral faces of merus, carpus and propodus, and on the distal margins of joints; small scale setae scattered among the scales. Male pereiopod 7 with elongate ischium bearing a dorsodistal depression on the frontal face and having a very slightly concave ventral margin. On basipodite, a distinct longitudinal goove is seen on the frontal face. This is without doubt homologous with the groove that is part of the water-conducting system in some other species of Oniscidea, but no scale-rows can be seen (at least not in the specimen examined).

Male pleopods: pleopod 1 and 2 exopodites with weakly wrinkled respiratory fields. Exopodite 1 broader than long, distal margin concave, without marginal setae. Exopodites 2–5 with marginal setae. Exopodite 5 on dorsal (caudal) face with irregular transverse band of pectinate scales, and with a furrow along mesal margin. Endopodite 1 apically slightly curved in lateral direction. Row of 31 small setae along the spermatic furrow. End of this row exceeded by a narrow tip bearing short longitudinal sulci. Endopodite 2 slightly exceeding exopodite. Sympodites 3 and 5 with acute, sympodite 4 with less acute, mediodistal process.

Uropod sympodite more than four times as wide as exopodite. Sympodite as long as wide (at the base), with rounded outer corner. Endopodite slightly longer than sympodite, with dense fringe of scales (and/or setae) on the inner surface.

Geographical distribution (Map Fig. 105 View Figure 105 )

Colombia: Sierra Nevada de Santa Marta.

Habitat

Cloud forest, ‘under leaves and logs in forest, not common; females carrying young were collected on July 3 and 12’ ( Pearse, 1915).

Remark

The transfer of this species from Sphaeroniscus to Scleropactes by Schultz (1970) is accepted. It is justified by the presence of putative synapomorphies with other species of Scleropactes (see clade 21), and the absence of the apomorphies of Sphaeroniscus .