Andrena (Suandrena) maderensis COCKERELL , 1922

Arncke, 2021, Taxonomic re-evaluation of Andrena cyanomicans PÉREZ, 1895, A. fratella WARNCKE, 1968, A. maderensis WARNCKE, 1969, A. mirna WARNCKE, 1969, A. notata WARNCKE, 1968, and A. portosanctana WARNCKE, 1969 (Hymenoptera, Anthophila), Linzer biologische Beiträge 53 (2), pp. 631-698 : 638-641

3. Andrena (Suandrena) maderensis COCKERELL, 1922

Andrena bimaculata KIRBY ?, var.

Andrena maderensis maderensis COCKERELL, 1922

S t a t u s: Andrena maderensis was described by COCKERELL (1922) on the basis of four females and two males from Madeira Island, kept in the T. V. Wollaston collection (NHMUK). He referred to SAUNDERS (1903), who described these specimens as ‘ A. bimaculata var.?’. SAUNDERS (1903) also wrote, ‘males and females. Madeira. In too bad condition to determine for certain’. According to FELLENDORF et al. (1999), the types of A. maderensis (four females and two males) are deposited in the University Museum, Hope Entomological Collections, Oxford, but J. E. Hogan, Oxford, confirmed that the types are not in Oxford. They are deposited in the NHMUK (GUSENLEITNER & SCHWARZ 2002; also pers. comm. J. Monks, NHMUK). All specimens are pinned and in good condition. Five of the specimens have a syntype label on them. There is one female without a syntype label; presumably this specimen has lost its label. All six specimens have a small printed label reading ‘ Madeira 58-21’ and a second handwritten label reading ‘ Andrena maderensis , Ckll’. The exceptions are a female labelled by Cockerell as the type and a male, labelled as the cotype. WARNCKE (1968) defined the nominate species A. m. maderensis (Madeira), based on one female and one male (type and cotype of A. maderensis, NHMUK ), as well as one male collected on 13.06.1957 by Lindberg (Valparaíso west of Camacha, Madeira), which was deposited in the collection of Grünwaldt. Today, this last specimen is no longer part of the collection of Grünwaldt (ZMS), but the accompanying female is still there.

Type ( Fig. 6 ): The female NHMUK 014026851 ( NATURAL HISTORY MUSEUM 2014) is kept separately in the bee type collection (syntype: primary type number 17a.2820). The labels are attached in the following order: printed register label reading ‘ Madeira 58-21’; handwritten type label in ink by Cockerell reading ‘ Andrena maderensis Ckll type’; round printed label with a blue margin reading ‘Syntype’; printed label reading ‘B.M. TYPE HYM.’; and handwritten reading ‘17a2820’ (the registration number of Hymenoptera types at the NHMUK). The labelling is similar to the NHMUK’s lectotype and paralectotypes of A. wollastoni , which were analysed by KRATOCHWIL (2018). MICHENER (cited in ZUPARKO 2008) has pointed out that Cockerell, for a species newly described by him, marked one specimen with ‘type’ (in the sense of holotype) for every new species he described.

The female of A. maderensis with the barcode label reading ‘ NHMUK 014026851’ will be designated as the lectotype. The label of the type specimen reading ‘ Madeira 58-21’ is an administrative entry label for the museum’s acquisition register. The number 58 refers to the year 1858 (the entry in the museum’s collections). As in the case of the lectotype of A. wollastoni , the origin of this female A. maderensis specimen is still an open question (KRATOCHWIL 2014). The exact time of transfer to the NHMUK is not clear, but the reference to the specimen in COCKERELL’ s (1922) published description is evident. A comparison with typical Cockerell labels confirms the authenticity of these labels (KRATOCHWIL 2014). The syntypes were deposited sometime between the years

1858 and 1922. The number 21 in the label reading ‘ Madeira 58-21’ refers to the twentyfirst register of specimens, a specific group of specimens. MACHADO (2006) described the chronology of the distribution of the Wollaston collections. At the NHMUK, there is no record for ‘ Madeira 58-21’ ( MACHADO 2006). Some collections from other museums were sold or just transferred to the NHMUK and had own registration labels (further information in KRATOCHWIL 2014). This may be one of them.

Cotype ( Fig. 7 ): The labels are in the following order: printed register label reading ‘ Madeira 58-21’; handwritten type label in ink by Cockerell reading ‘ Andrena maderensis Ckll cotype’; round printed label with blue margin reading ‘Syntype’. This male, along with all other types of the series, will be designated as a paralectotype.

Figures

Fig. 6a-b: Lectotype of the female of Andrena maderensis COCKERELL, 1922 (NHMUK). (a) dorsal view, (b) lateral view, labels. Photos: J. Monks, copyright NHMUK.

Fig. 7a-b: Paralectotype of the male of Andrena maderensis COCKERELL, 1922 (NHMUK). (a) dorsal view, (b) lateral view, labels. Photos: J. Monks, copyright NHMUK.