Metalycaeus heudei (Bavay & Dautzenberg, 1900)

Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander & Auffenberg, Kurt, 2021, Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea), Zootaxa 4973 (1), pp. 1-61 : 53

Metalycaeus heudei (Bavay & Dautzenberg, 1900)

Fig. 41

Alycaeus (Charax) heudei Bavay & Dautzenberg, 1900a: 121–122 .

Alycaeus (Charax) heudei — Bavay & Dautzenberg 1900b: 458–459, plate 11, figs 15–18.

Alycaeus paviei Bavay & Dautzenberg, 1912: 50–51 , pl. 4, figs 5–8.

Alycaeus paviei var. minor Bavay & Dautzenberg, 1912: 51 , pl. 4, fig. 9.

Chamalycaeus (Dicharax) compressicosta Zilch, 1957: 145–146 , fig. 33.

Chamalycaeus (Dicharax) fractus Varga, 1974: 165–167 , figs 1–5.

Alycaeus zhuangiyucuii Yang, Fan, Qiao & He, 2012: 32 , fig. 2.

Metalycaeus heudei — Páll-Gergely et al. 2017: 74–84, figs 49C–D, 50–52, 53C–D. ( paviei Bavay & Dautzenberg, 1912 , paviei var. minor Bavay & Dautzenberg, 1912 , compressicosta Zilch, 1957 , fractus Varga, 1974 and zhuangiyucuii Yang, Fan, Qiao & He, 2012 are synonyms; Inkhavilay et al. 2019: 16, fig. 6C.

Alycaeus kengtungensis Godwin-Austen, 1914: 409 , plate 139, figs 6, 6a. new synonym

Material examined. Holotype: Myanmar, Shan State, “Kengtung, Siam Frontier, leg. Woodthorpe ”, NHMUK 1903.7 .1.3037 (holotype of A. kengtungensis ). For the examined type specimens of other taxa see Páll-Gergely et al. (2017) . Other material: 4 shells, Myanmar, Kayah State, Hpruso Distr., Maw Thi Do Village, Bo Pha Gu Cave sink, 19°21.121’N, 97°3.759’E (locality code: JG10 ), 10 Feb. 2019, J. Grego leg., JG ; 2 shells, Myanmar, Kayah State, Hpruso Distr., Maw Thi Do, road towards Han Li Village , under bridge over Phruno River , 19°22.966’N, 97°02.153’E (locality code: JG100 ), 12 Dec. 2019, J. Grego leg., JG ; 1 shell, Myanmar, Kayah State, Hpruso Distr., Maw Thi Do, road towards Han Li Village , rocks above bridge over Phruno River , 19°23.011’N, 97°02.108’E (locality code: JG101 ), 12 Dec. 2019, J. Grego leg., JG ; 1 shell, Myanmar, Kayah State, Hpruso Distr., Maw Thi Do Village, Entrance of Phruno River Cave , 19°22.744’N, 97°02.570’E (locality code: JG102 ), 12 Dec. 2019, J. Grego leg., HNHM 104867 (photographed shell: 4.77 mm, H: 3.66 mm) ; 17 shells, same data as preceding, JG .

Diagnosis of specimens from Myanmar. A medium-sized to large species with very long R2 having typical Metalycaeus - type elevated, thin, widely spaced ribs, a narrow swelling on R3, and a rounded aperture.

Measurements. D: 4.65–5.05 mm, H: 3.66–3.71 mm (newly collected specimens).

Remarks. Metalycaeus heudei is a widely distributed, variable species reported from China, Vietnam and Laos. Alycaeus kengtungensis ( Fig. 41B ) from the eastern border of Shan State, Myanmar, and the newly collected shells from Kayah State ( Fig. 5 ) fall within the known morphological variation of Metalycaeus heudei . Therefore, we consider A. kengtungensis a new synonym of M. heudei . With this, M. heudei is the most widely distributed alycaeid species known.

Figures

FIGURE 5. Distribution of alycaeid species. Inverted empty triangle: Dicharax cyclostoma Páll-Gergely, n. sp., empty triangle: Chamalycaeus spiratus Páll-Gergely, n. sp., filled triangle: Metalycaeus bicarinatus Páll-Gergely & Hunyadi, n. sp., inverted filled triangles: Chamalycaeus erawan Páll-Gergely & Hunyadi n. sp., empty circles: Metalycaeus aries Páll-Gergely & Auffenberg, n. sp., filled circles: Metalycaeus heudei (Bavay & Dautzenberg, 1900) (the one in the north indicates the type locality of Alycaeus kengtungensis Godwin-Austen, 1914), square: Metalycaeus pygmachos Páll-Gergely & Hunyadi, n. sp.

FIGURE 41. Metalycaeus heudei (Bavay & Dautzenberg, 1900). A: from Kayah State, Myanmar (site JG102); B: holotype of Alycaeus kengtungensis Godwin-Austen, 1914 (NHMUK 1903.7.1.3037).