Chlaenius (Pseudochlaeniellus) puncticollis Dejean, 1826

Anichtchenko, Alexander & Kirschenhofer, Erich, 2017, To the knowledge of Oriental species of subgenus Pseudochlaeniellus Jeannel, 1949 (Coleoptera, Carabidae, Chlaenius), Zootaxa 4231 (2), pp. 187-202 : 192

Chlaenius (Pseudochlaeniellus) puncticollis Dejean, 1826

( Figs. 16–17 , 30 )

Chlaenius puncticollis Dejean, 1826: 315

Chlaenius jedlickai Mandl, 1991: 133 [HN]

Chlaenius persicus Jedlicka, 1968d: 989 [HM]

C. (Pseudochlaeniellus) panjabensis Kirschenhofer, 1998: 320 syn. n.

Material. INDIA: ♂ Type—“ Chlaenius puncticollis Dej . India Or. ”, “ Indien ” ( NMB); 1 ♂ — “N India, Uttarakhand, 22– 23.04.2012, 8 km NW Jaspur , lake shore, 29°21’10.54”, 78°48’30.99”, Anichtchenko A. leg.” ( NMW) ; 17 ♂, 10 ♀ —“N India, Uttarakhand, 22– 23.04.2012, 8 km NW Jaspur , lake shore, 29°21’10.54”, 78°48’30.99”, Anichtchenko A. leg.” (cAA) ; 2 ♂—“Madras” (NMW); 1 ♀ —“ India, Panjab , Chandigarh ”, red label “ Holotypus ”, white label “ Chlaenius panjabensis n. det. Kirschenhofer 96”, white label “ Chl. (Pseudochlaeniellus) panjabensis n . det. Kirschenhofer 96” (NMW); 1 ♀— “ India, W. Bengal, Nalbani, N. Salt Lake , leg. Gy. Topál, No. 20, trodden from mud, 2.XII.1966 ( NMW) . IRAN: 1 ♂ — “ Iran, Hormozgan prov., Geno, Mola Dam (A.13/1), 27°20’44”N, 56°08’3”E, 232m, 1.3.2015, S. Azadbakhsh leg.” (cAA) . PAKISTAN: 1 ♀ — “ West Pakistan, Rawalpindi Umg., Dhok Pathan (Sohan R.) b. Pindi Gheb 1.56, Chr. Lindemann leg.” ( NMW) . NEPAL: 1 ♂ —“ C. cf. nigrosuturatus det. Kirschenhofer, 07”, “ Nepal—Himalaya, Annapurna Mts., Pokhara Tal , 900m, 19.6.1994, lg. Schmidt ” (cJS) ; 1 ♀ —“ Ch. cf. nigrosuturatus det. Kirschenhofer ”, “ Nepal, 31.10.92, Pokhara—See, leg. J. Schmidt leg.” ( NMW).

Diagnosis. This species is more similar in overall appearance to the sympatric C. (Chlaeniellus) nigrosuturatus Mandl, 1978 comb. n. The two species can be diagnosed easily by the pubescence of tarsa and lateral margins of pronotum, i.e. C. nigrosuturatus has tarsa smooth, and yellow lateral margins. Species with constant elytral pattern ( Figs. 16–17 ).

Description. Body length 9–10 mm. Dorsal surface with dense yellow pubescence. Coloration of head and pronotum metallic green with golden overtones on the disc. Mouthparts, antennomeres and legs yellow. Antennomeres IV–XI slightly darker. Elytra metallic green, with constant elytral pattern. The yellow lateral band narrow, strictly limited to two external intervals ( Figs. 16–17 ); apical part of band narrow. Ventral surface black, iridescent, except wide lateral band on all abdominal segments.

Head moderately punctate, smooth and shiny on vertex. Pronotum dull, strongly microreticulate, very deeply and densely punctate at the base, less one diameter distance from each other. Anterior half of pronotum less densely punctate. Lateral margins narrow. Hind angles right. Elytral striae narrow, finely punctured. All intervals flat.

Aedeagus long, narrowed to pointed apex ( Fig. 30 ). Apice with hook.

Distribution. Afganistan, India (Himachal Pradesh; Uttarakhand), Iran, Nepal, Pakistan.

Figures

FIGURES 13 – 18. Habitus of C. (Pseudochlaeniellus): 13 — Holotype of C. (P.) viangchanensis syn. n. = lomsakensis (cDW); 14 — C. lomsakensis, Paratype (NMW); 15 — Holotype of C. (P.) farkaci syn. n. = lomsakensis (cJF); 16 — Holotype of C. (P.) panjabensis syn. n. = puncticollis (NMW); 17 — C. puncticollis. India (cAA); 18 — C. (P.) davidsoni (cAA).

FIGURES 28 – 36. Aedeagus: 28 — Holotype of C. chloodes = C. neochloodes (BMNH); 29 — C. caesitius, Loc. Typ. (NMW); 30 — C. puncticollis. India (cAA); 31 — Paratype of C. mesopotamicus = lucasi (NMW); 32 — C. germanus, Karnataka, India (cAA); 33 — C. germanus, Sylhet, Bangladesh. (BMNH); 34 - 35 variability of aedeagus of C. sobrinus, Rajaji, India (cAA); 36 — C. sobrinus, Tonkin, Vietnam (NMW).