Scaphobaeocera data, Lobl, 2011

Löbl, Ivan, 2011, On the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of the Philippines, II, Revue suisse de Zoologie 118 (4), pp. 695-721 : 700-702

Scaphobaeocera data sp. n. Figs 7-10

HOLOTYPE: Ƌ, Luzon Mount Data Lodge 2200-2300m 22-23.XII.1979, L. Deharveng & J. Orousset #100 ( MHNG).

PARATYPES: 26 Ƌ, 15 ♀, with the same data as the holotype; 2 Ƌ, with the same data but #86 . – 1 ♀, 1 ♀, with the same data but #162; Mount Data , 9.I.80, J. Orousset #165. – 3 Ƌ , 5 ♀, Luzon, Mt. Data (2250m) Mountain Prov. , 13., 14., 25. and 26. VII.1985, M. Sakai leg. – 2 Ƌ, Luzon Mt. Fangao, (2350m) Mountain Prov. , 14.VII.1985, M. Sakai leg. – 2 Ƌ , 14 ♀, Pacay (2400m) nr. Sayangan Benguet Prov., 11. VII.1985, M. Sakai leg. (all MHNG) .

FIGS 7-10

(7-10) Scaphobaeocera data sp. n., aedeagus (7, 9) and internal sac (8, 10) in dorsal and lateral views.

DESCRIPTION: Length 1.15-1.30 mm, width 0.65-0.73 mm, dorsoventral diameter 0.70-0.78 mm. Body very dark reddish-brown to almost black, apices of elytra usually lighter, apical abdominal segments and appendages ochraceous. Thorax, elytra and abdomen microsculptured and iridescent. Pronotal and elytral punctation very fine, hardly visible at magnification 100x. Length ratio of antennomeres as: III 6: IV 8: V 10: VI 8: VII 10: VIII 7: IX 10: X 10: XI 16. Segments III and IV equally narrow, segment III about 3 times as long as wide. Segments V and VI wider than segment IV, each about 3 times as long as wide. Segment VII almost 3 times as long as wide. Segment VIII slightly narrower than segment VII, about twice as long as wide. Segments IX and X distinctly wider than segment VII, each about twice as long as wide. Segment XI about 2.5 times as long as wide. Scutellum entirely concealed. Elytra with very fine parasutural striae, sutural striae starting at margin of pronotal lobe, slightly curved at base. Hypomeron without stria. Middle part of metaventrite flat, with shallow stria, very dense and fine punctation, and very short pubescence. Sides of metaventrite sparsely and very finely punctate, with long pubescence. Mesocoxal lines with fine marginal punctures, not extending laterally along mesepimera; submesocoxal areas about 0.02-0.03 mm long. Metepisterna flat, with exposed portion about 0.04-0.06 mm wide, parallel-sided, with straight suture. Abdomen with distinct microsculpture consisting of transverse striae. Abdominal ventrite 1 very finely punctate, with basal punctures fine, not elongate. Tibiae straight.

Male characters. Protarsi with segments 1 to 3 distinctly widened, narrower than protibiae. Aedeagus (Figs 7-10) 0.36-0.40 mm long.

HABITAT: Mountain broad-leaf forest, under bark, in rotten wood and floor litter.

DISTRIBUTION: Philippines: Luzon.

COMMENTS: As in S. data , the Asian S. cyrta Löbl, 1980, S. discreta Löbl, 1980 and S. smetanai Löbl, 1981 possess flagellar basal hook and the median lobe of the aedeagus is lacking ventral processes. The new species may be distinguished from them by the convex apicoventral side of the basal bulb. Besides, S. smetanai may be separated by the robust basal part of the internal sac, S. discreta by the parameres gradually widened apically (in lateral view) and S. cyrta by the median lobe conspicuously arcuate, with tip reaching beyond parameres. The Papuan and Australian S. papuana Csiki, 1909, S. ornata (Pic, 1956) , S. piceoapicalis Löbl, 1977 and S. queenslandica Löbl, 1986 have also aedeagi comparatively similar to that in S. data . The first three of them may be readily distinguished by their distinctive body coloration, S. queenslandica differs drastically by the shape of the parameres. Other Asian species that possess an internal sac with a basal hook are those originally described as Baeotoxidium Löbl, 1971 , and since transferred to Scaphobaeocera : S. elegans (Löbl, 1971), S. gagata (Löbl, 1971), S. indica (Löbl, 1979), S. lanka (Löbl, 1971), S. siamensis (Löbl, 1990), and S. yeti ( Löbl, 1992). These species are linked by the internal sac of the aedeagus with a basal vesicle containing spine-like structures, and probably form a monophyletic group.