Cyanus nigrofimbrius (K.Koch) Soják (1972: 131)

Negaresh, Kazem, 2022, A checklist of the genus Cyanus (Asteraceae, Cardueae) in Turkey, with notes on taxonomy and distribution, Phytotaxa 531 (1), pp. 18-40 : 26-27

10. Cyanus nigrofimbrius (K.Koch) Soják (1972: 131) .

Centaurea montana var. nigrofimbria K. Koch (1851: 426) . Centaurea nigrofimbria (K.Koch) Sosnovsky (1926: 77) . Centaurea atrata subsp. nigrofimbria (K.Koch) Dostál (1931: 60) . Type:— TURKEY. Kl. Asien, Pontus: Lasistan, Djimil, 20 July 1843, K.Koch s.n. (lectotype B 10 0250671! designated here, isolectotype GOET008603!).

Taxonomic notes: — Koch (1851) described Centaurea montana var. nigrofimbria ( Cyanus nigrofimbrius ) based on two collections of him, i.e. “im kaukasischen Hochgebirge auf Porphyr und Thonschiefer, ca. 5000 bis 8000 hoch” and “im Gaue Hemschin auf Urgestein, ca. 6000 bis 9000 hoch”.

The collection “im kaukasischen Hochgebirge auf Porphyr und Thonschiefer” is located in Caucasus, but no specimen with such collection data was found among the herbaria I visited.

Hemschin is a region on N side of Tatus Daghlari (incl. Kaçkar Dagh ) between Ikizdere and Camlihemshin [ Turkey A8 Rize] ( Lack 1978). This region appears as a large historical district within the bounds of Lazistan , on the oriental Pontic chain which reaches 3937 m at the Kaçkar Dagh (east of Trabzon). Hemschin is near to locality Dschimil (Cimil), which, following Koch’s travel planning (Edmondson & Lack 1977), seems to be close to the modern city of Ikizdere . Two sheets bearing this region data, kept at B and GOET, were investigated. The preference is given to the B one, because Koch’s autographs are present only on this sheet. The other sample is kept in GOET collection; its label is not from Koch’s hand. The specimen Koch s.n. deposited in B is designated here as the lectotype because it is the only original material that was annotated by Koch and the specimen shows all the relevant characters in regard to the protologue.

On the label of the lectotype is printed: “ Kl. Asien, Pontus: Lasistan, Djimil, 1843–1845, K.Koch ” and handwritten “Dschimil, 20.VII.1843, no. 780”. On the label of the isolectotype GOET008603 has handwritten “ Centaurea montana var. nigrofimbria, Dschimil, K.Koch ”.

Distribution: — Cyanus nigrofimbrius generally occurs in Giresun, Trabzon, Ardahan and Rize Provinces, N to NE Turkey ( Fig. 5 ). There are also a few populations in Azerbaijan.

Selected specimens examined:— TURKEY. A7 Giresun: below Tamdere , 1600 m, 9 August 1952, Davis et al. 20690 (E!). A8 Trabzon: Bayburt to of 3 km N of pass top, 2400 m, 11 July 1960, Stainton & Henderson 6199 (E!); Soganli Daglari, 30 km N of Bayburt, 2600 m, 8 August 1972, Edmondson 859 (E!); N side of Soganli dağ above Caykara, 4 August 1957, Davis & Hedge D. 32159 (E!). A8/9 Çoruh (Ardahan-Artvin): Otingol, 1800 m, 4 July 1960, Stainton & Henderson 6005 (E!); Of-Bayburt Rd. about 45 miles S from of, 7 July 1952, Apold et al. 186 (E!); Haldizan N of Baiburt, 26 July 1936, Balls 1851 (E!) . Rize: Cermanin Tepe above Cimil, 3100 m, 29 August 1952, Davis & Dodds 21081 (E!); Ikizdere , Vercinin Tepe, steep grassy scree, 3300–3400 m, 29 August 1952, Davis & Dodds 21153 (E!); Baskoy (Cimil)-Cermanin Yayla, 2300 m, 28 August 1952, Davis & Dodds 21028B (E!) .

Figures

FIGURE 5. Distribution of Cyanus lanigerus (circles), C. matthiolifolius (squares), and C. nigrofimbrius (triangles) in Turkey.