Agathidium tenangoense Miller and Wheeler, 2005

MILLER, KELLY B. & WHEELER, QUENTIN D., 2005, Slime-Mold Beetles Of The Genus Agathidium Panzer In North And Central America, Part Ii. Coleoptera: Leiodidae, Bulletin of the American Museum of Natural History 2005 (291), pp. 1-167 : 121-123

Agathidium tenangoense Miller and Wheeler , new species Figures 149 , 277–279 , 367

TYPE MATERIAL: Holotype, Ƌ in CMNC labeled ‘‘GUAT: HUEHUETENANAGO 10 kmW La Capellania 3.viii.78. Helava. /HO­ LOTYPE Agathidium tenangoense Miller

and Wheeler, 2003 [red label with black line border]’’.

TYPE LOCALITY: Guatemala, Huehuetenango, 10 km W La Capellania.

DIAGNOSIS: This species is similar to other members of the A. aztec subgroup with a small male metafemoral tooth (fig. 149). The eyes are relatively prominent, the metasternum is flat with a small, medial fovea in the male and has indistinct oblique carinae, the gula is unmodified, and the elytra are conspicuously iridescent. The male genitalia are diagnostic with the apical portion of the median lobe broad basally and strongly constricted to a stout, short apically truncate apical process (fig. 277). The operculum is broad, long and apically broadly subtruncate with a small apical emargination (fig. 277).

DESCRIPTION: Body moderate in size (TBL = 3.72 mm), broad, robust (PNW/TBL = 0.48), strongly contractile.

Head piceous; pronotum piceous, dark red around margins; elytra piceous, red around margins, prominently iridescent laterally; venter dark red­brown, piceous on metasternum, antennae and palpi dark red­brown; legs red­brown.

Head broad (MDL/OHW = 0.63), dorsal surface flattened, dorsoventrally compressed; with very fine punctures, each with a short, very fine seta, surface between punctures shiny, smooth; frontoclypeal suture obsolete medially; eyes moderately large, not dorsoventrally compressed; gula slightly concave; antennomere ratios: length I:II:III = 1.7:1.0: 1.8, width VII:VIII:IX = 1.0:1.0:2.1. Pronotum very large, broad (PNL/PNW = 0.73), strongly convex, anterolateral lobes strongly produced, lateral margin broadly curved, not angulate; with very fine, sparse punctures, each with a short, very fine seta, surface between punctures smooth. Elytra broad, lateral margins strongly rounded, apically rounded (SEL/ELW = 1.08); punctation similar to pronotum; sutural stria present in apical onethird of elytron. Flight wings fully developed. Mesosternum narrow medially (MTL/ MTW = 0.18), broadly rounded; medial carina well developed. Metasternum moderately broad, flat, sloping dorsad anteriorly, smooth and relatively glabrous; oblique femoral carinae indistinct, medially meeting in very low carina.

Male tarsi 5­5­4; pro­ and mesobasotarsomeres somewhat laterally expanded, with small field of ventral spatulate setae; mandibles not modified; metafemur moderately broad with small, indistinct tooth subapically on posterior margin (fig. 149); metasternal fovea small, slightly transverse with small pencil of dense, fine setae. Median lobe in lateral aspect slender and strongly curved basally, slightly expanded submedially, with prominent lateral carinae with lateral sulcus in which fits the lateral lobe, apical portion slightly angled dorsad, stout, slender, apically slightly expanded (fig. 278); in ventral aspect slender, long, slightly constricted submedially, laterally expanded at base of apical portion by lateral carinae, apical portion broad basally, abruptly narrowed, with apex consisting of stout, truncated process (fig. 277); operculum long, flat, broad, stout, apically broadly subtruncate with small medial emargination (fig. 277); lateral lobes slender basally, curved basally, apically distinctly sinuate and expanded submedially and apically, apex rounded with 2 stout setae (fig. 279).

Female not examined.

ETYMOLOGY: This species is named after an abbreviated form of the name of the type locality in Guatemala.

DISTRIBUTION: This species is known only from the type locality in Guatemala (fig. 367).

PARATYPE: GUATEMALA: Huehuetenango: 10 km W LaCapellania, 3 Aug 1978, Helava (1, CNCI).

Figures

Figs. 145–172. Agathidium oniscoides­group species, left male metafemur, ventral aspect: 145, A. oculeum. 146, A. recurvatum. 147, A. impensum. 148, A. cheneyi. 149, A. tenangoense. 150, A. rumsfeldi. 151, A. hirsutum. 152, A. cortezi. 153, A. tumidiventre. 154, A. aztec. 155, A. iridescens. 156, A. dentigerum. 157, A. stephani. 158, A. akrogeneios. 159, A. appalachium. 160, A. pocahontasae. 161, A. carolinense. 162, A. gallititillo. 163, A. divaricatum. 164, A. bushi. 165, A. georgiaense. 166, A. chauliodoum. 167, A. microphthalmum. 168, A. nimbosilva. 169, A. oniscoides. 170, A. rubellum. 171, A.

Figs. 261–287. Agathidium oniscoides­group species, aedeagus: 261–263, A. oaxacaense: 261, ventral; 262, median lobe, lateral; 263, left lateral lobe, lateral. 264–266, A. oculeum: 264, ventral; 265, median lobe, lateral; 266, left lateral lobe, lateral. 267–269, A. recurvatum: 267, ventral; 268, median lobe, lateral; 269, left lateral lobe, lateral. 270–272, A. impensum: 270, ventral; 271, median lobe, lateral; 272, left lateral lobe, lateral. 273–276, A. cheneyi: 273, ventral; 274, median lobe, apex, ventral; 275, median lobe, lateral; 276, left lateral lobe, lateral. 277–279, A. tenangoense: 277, ventral; 278, median lobe, lateral; 279, left lateral lobe, lateral. 280–283, A. rumsfeldi: 280, ventral; 281, median lobe, apex, ventral; 282, median lobe, lateral; 283, left lateral lobe, lateral. 284–287, A. hirsutum: 284, ventral; 285, median lobe, apex, ventral; 286, median lobe, lateral; 287, left lateral lobe, lateral.

Figs. 367, 368. Geographic distribution of Agathidium oniscoides­group species: 367, A. hirsutum = -; A. tenangoense = •; A. iridescens = ^; A. rumsfeldi = *. 368, A. aztec = ^; A. cortezi = *; A. tumidiventre = •.