Paradrilus marginatus, Pic, 1918

Kundrata, Robin & Bocak, Ladislav, 2011, Redescription and relationships of Pseudothilmanus Pic (Coleoptera: Rhagophthalmidae) — a long-term neglected glow-worm beetle genus from the Himalayas, Zootaxa 2794, pp. 57-62 : 60-62

Pseudothilmanus marginatus Pic, 1918

( Figs 2, 4, 6, 11 , 14–15 )

Pseudothilmanus (Drilothilmanus) marginatus Pic, 1918: 3 .

Type material. Holotype, 1 male, India, Darjeeling, June, Fruhstorfer leg. ( MHNP).

Diagnosis. The diagnosis is given under P. alatus above.

Redescription. Male. Body small, slender, dorsoventrally flattened ( Fig. 2 ). Head, elytra and antennomeres 3– 12 dark brown, basal antennomeres and legs slightly lighter, mouthparts yellow to light brown; pronotum dark in the middle of disc, margins lightly colored, yellow to light brown. Entire body covered by sparse, yellow pubescence.

Head slightly hypognathous, partly hidden by pronotum, including eyes 1.06 times wider than anterior margin of pronotum. Cranium transverse, shiny, with fine microstructure, covered with long sparse erected pubescence. Eyes medium-sized, hemispherically prominent, eye diameter 1.13 times their frontal distance. Labrum small, retracted in cranium, with simple anterior margin. Mandibles slender, long, with simple incisor, curved in right angle apically, shiny. Maxillary palpi 4-segmented, basal palpomere small, triangular, palpomeres 2 and 3 subequal, their combined length less than those of palpomere 4, both as long as width, apical palpomere twice longer than width, gradually narrowed to apex, sharply pointed. Labial palpi tiny, apical palpomere pointed. Labium small, feebly sclerotized. Gula small, transverse, membranous anteriorly. Antennae robust, serrate, reaching first third of elytral length, scapus robust, longer than pedicel, pedicel small, ratio of antennomeres 1–3 1.56:1:1.89, antennomeres 3 to 9 slightly serrate, equal in length, moderately flattened (only 9 antennomeres preserved in the type specimen); all antennomeres with small tubercles and long, erected pubescence ( Fig. 4 ).

Pronotum with bulge in the middle, widest at basal margin, 1.60 times wider posteriorly than length at midline. Anterior margin almost straight, lateral margins convex, posterior margin bisinuate, with median notch. Anterior angles rounded, posterior angles prominent, acute ( Fig. 11 ); disc surface coarsely punctured, with sparse, erected, long setae, denser at margins. Prosternum transverse, forming slender bridge-shaped structure, prosternal process plate-like, about as long as width, emarginate apically. Scutellum small, flat, lateral margins parallel-sided in posterior half, posterior margin straight, widely emarginate at middle. Mesosternum transverse, anterior margin vshaped. Elytra slightly divergent, narrower to apex, longer than abdomen, humeri prominent, with strong costa running from humeri to interior margin of elytral apex, weaker to inconspicuous in apical third of elytra ( Fig. 2 ), elytra widest at humeri, punctured, 2.60 times longer than width at humeri, with strengthened edges, apex with strongly strengthened margin, covered with sparse, erected pubescence. Abdomen short, reaching over three fifths of elytral length. Legs very slender, covered by dense, long, erected hairs, coxae long, slender, trochanters small, obliquely attached to femora, only tibiae and femora slightly compressed, tarsi slender, without pulvillae, tarsomeres 1–3 subequal, tarsomere 4 about half length of tarsomere 3, tarsomere 5 slightly longer than tarsomere 3, claws simple, slender, slightly curved ( Fig. 6 ).

Male genitalia robust, apical third of phallus slender, parameres robust and short, phallobase transverse ( Figs 14–15 ).

Measurements. BL 4.85 mm, EL 3.75 mm, WH 1.44 mm, PL 0.63 mm, PW 1.01 mm, Edist 0.31 mm, Ediam 0.35 mm.

Distribution. India: West Bengal, Darjeeling.

Figures

FIGURES 1 – 11. 1 – Pseudothilmanus alatus, general appearance; 2 – P. m a rg i n a t u s, general appearance; 3 – P. alatus, antenna; 4 – P. m a rg i n a t u s, antenna; 5 – P. alatus, leg; 6 – P. m a rg i n a t u s, leg; 7 – P. alatus, detail of elytral apex; 8,9 – P. alatus, terminal abdominal segments; 10 – P. alatus, pronotum; 11 – P. m a rgi n at u s, pronotum. Scale 2.0 mm (Figs 1 – 2), 0.5 mm (Figs 3 – 11).

FIGURES 12 – 15. Male genitalia. Pseudothilmanus alatus 12 – laterally; 13 – ventrally; Pseudothilmanus marginatus 14 – laterally; 15 – ventrally. Scale 0.25 mm.