Systaria cervina (Simon, 1897)

Dankittipakul, Pakawin & Singtripop, Tippawan, 2011, Seven new species of Systaria Simon, 1897 from Southeast Asia (Araneae, Clubionidae, Systariinae), Zootaxa 2905, pp. 16-32 : 20-21

Systaria cervina (Simon, 1897)

Figs 4–8

Syrisca cervina Simon, 1897b: 500 , description of female. Systaria cervina (Simon) . Deeleman-Reinhold, 2001: 203, figs 236– 237, transfer from Syrisca .

Type material. Holotype: Ƥ, PHILIPPINES: Rizal: Antipolo ( MNHN, AR 13751), examined. [Antipolo is a city locating in the Province of Rizal, ca. 25 km east of Manila. At the back side of Antipolo Church is a large mystical cave known as Inday Nelly Mystical Cave. It is likely that the female syntype was collected from this cave.]

Other material examined. PHILIPPINES: Quezon Province: 23, 1Ƥ, 2 juveniles (presumably female), Pagbilao, Cueva Balisen, 20 February 1975, leg. P. Strinati, ( MHNG, Syst-25449).

Diagnosis. Males of S. cervina can be distinguished from those of S. elberti (Strand, 1913) by the sickleshaped RTA provided with a small dorsal incision in lateral view ( Figs 5–6 ). Females can be distinguished from those of S. elberti by the anterior bursae abruptly narrowing, forming short, posterior stalks ( Fig. 8 ) instead of being elongate-oval, and by the much narrower insemination ducts connected between anterior bursae and posterior spermathecae. The male palpal structure of S. cervina also closely resembles S. insulana (Rainbow, 1902) but can be recognized by the more elongated RTA which is greatly narrower at base, and by the smaller conductor.

Description. Male (MHNG, Syst-25449): Total length 7.38. Prosoma 3.32 long, 2.28 wide. Opisthosoma 4.06 long, 1.92 wide. Leg formula: 4123; I 10.32 (2.78, 1.36, 2.66, 2.38, 1.14); II 8.66 (2.56, 1.18, 2.38, 1.62, 0.92); III 7.78 (2.08, 0.82, 1.76, 2.18, 0.94); IV 11.56 (3.24, 1.02, 3.08, 3.16, 1.06). Spination. Femora I d1-1-1, p1-1-1, r0-1- 1; II d1-1-1, p1-1-1, r0-1-0; III d1-1-1, p1-1-0, r1-1-1; IV d1-1-1, p1-0-1, r1-1-1. Tibiae: I v2-2 -1-1; II v2-2 -2; III d1-0-0, p1-0-0, r2-2-2, v2-2 -2; IV d0-1-0, p1-1-2, r1-1-0, v2-2 -2. Metatarsi: I v1 -0-0; II v1-1 -1; III p1-1-1, r1-1-2, v0-1-1; IV p1-1-1, r2-2-2, v1-1 -1. Eye size and interdistances: AME 0.12, ALE 0.08, PME 0.08, PLE 0.06; AME– AME 0.16, AME–ALE 0.14, PME–PME 0.22, PME–PLE 0.14, ALE–PLE 0.12; MOA 0.26 long, anterior width 0.28, posterior width 0.26.

Palp ( Figs 4–6 ). Ventral surface of tibia with elevated, distal swelling. RTA slender, sickle-shaped, with small dorsal incision ( Figs 5–6 ). Cymbium fold partially membranous, membrane situated basally. Cymbial ridge dark reddish-brown, slightly elevated, running obliquely. Tegulum oval, excavated baso-prolaterally. Embolus triangular, membranous at base. Conductor membranous, flange-like, situated apically.

Female (MHNG, Syst-25449): Total length 10.52. Prosoma 4.80 long, 3.32 wide. Opisthosoma 5.72 long, 3.06 wide. Leg formula: 4123; I 8.88 (2.60, 1.30, 2.52, 1.62, 0.84); II 8.30 (2.54, 1.22, 2.22, 1.58, 0.74); III 7.94 (2.84, 0.82, 1.70, 1.86, 0.72); IV 12.42 (3.70, 1.24, 2.76, 3.22, 1.50). Spination. Femora: I d1-1-1, p1-1-0, r0-1-1; II d1-1- 1, p1-1-0, r1-1-1; III d1-1-1, p0-1-1, r1-1-1; IV d1-1-1, p1-1-1, r0-1-0. Tibiae: I v2-1 -1; II v0-2-0; III p1-1-0, r1-0- 1, v2-2 -2; IV p1-1-0, r1-1-0, v2-2 -2. Metatarsi: II v1-1 -0; III p1-1-1, r1-1-2, v2-1 -1; IV p1-1-2, r1-1-2, v2-2 -2. Eye size and interdistances: AME 0.16, ALE 0.12, PME 0.12, PLE 0.10; AME–AME 0.18, AME–ALE 0.16, PME– PME 0.24, PME–PLE 0.18, ALE–PLE 0.14; MOA 0.26 long, anterior width 0.38, posterior width 0.36.

Genitalia ( Figs 7–8 ). Epigynal plate triangular, weakly sclerotized. Copulatory orifices teardrop-shaped, situated medially. Anterior bursae sclerotized, rounded anteriorly, abruptly narrowing posteriorly. Single pair of spherical spermathecae situated posteriorly. Fertilization ducts lanceolated.

Natural history. New specimens of S. cervina were collected from their retreats at the entrance of a huge limestone cave.

Distribution. The Provinces of Rizal and Quezon, Luzon Island, the Philippines (Fig. 43).

Figures

FIGURES 4 – 8. Systaria cervina, new material from Luzon. 4 – 5 Left male palp (4 ventral, 5 retrolateral); 6 Retrolateral tibial apophysis, retrolateral; 7 Epigyne, ventral; 8 Vulva, dorsal. Scale lines = 0.1 mm.