Dryinus janzeni Olmi, 2000

Perkovsky, Evgeny E. & Olmi, Massimo, 2018, Discovery of the first pincer wasp (Hymenoptera, Dryinidae) from Rovno amber, Zootaxa 4457 (2), pp. 296-304 : 297-298

Dryinus janzeni Olmi, 2000

( Figs 2–5 )

Dryinus janzeni Olmi 2000: 58 (♀); Olmi & Bechly 2001: 49; Olmi et al. 2010: 35; OlmI et al. 2014: 711.

Diagnosis. A fossil female of Dryinus with incomplete notauli and enlarged claw much longer than arolium and with two subapical teeth.

Redescription, ♀. Macropterous ( Figs 2, 3 ); length 4.6–6.2 mm (holotype 5.7 mm; specimen from RoVno amber 5.1 mm). Colour of holotype apparently black, except antenna, legs and anterior half of face testaceousbrown. Colour of specimen from RoVno amber black, except antenna brown and distal extremity of metasoma brown-testaceous. Antenna with eight flagellomeres, long and slender, coVered with dense and short hairs, slightly thickened distally. Antennal ADO’s not distinctly Visible in holotype, apparently present on antennomeres 7–10; not Visible in specimen from RoVno amber. Antennomeres in following proportions: 12:8:29:16:15:12:10:9:8:10. Antenna much shorter than body, approximately 4 as long as head (length of head measured dorsally from occipital carina behind ocelli to distal apex of mandibles): 129:32. Head slightly excaVated, dull, punctate and granulate; clypeus rounded, with anterior margin not emarginate; mandible with four teeth becoming regularly progressiVely larger from upper to lower tooth; occipital carina complete, laterally not reaching eye; occiput excaVated; eye bulging; POL = 2; OL = 6; OOL = 11; OPL = 1; temple absent; frontal line incomplete, not present in anterior third of face. Apparent palpal formula 6:3. Pronotum crossed by anterior strong transVerse impression between anterior collar and disc; disc slightly humped; posterior collar Very short; pronotum apparently shiny, punctate, unsculptured among punctures, hairy, slightly shorter than head (28:32); pronotal tubercle reaching tegula. Mesoscutum apparently shiny, with sparse hairs, punctate and slightly granulated, much shorter than pronotum (15:28). Notauli incomplete, reaching approximately 0.66 length of scutum. Mesoscutellum shorter than mesoscutum (13:15), shiny, punctate and slightly granulated, with sparse hairs. Metanotum shorter than mesoscutellum (4:13), as sculptured as mesoscutellum. Metapectal-propodeal complex longer than mesoscutum (30:15), reticulate rugose, setose; metapectal-propodeal areolae Very large, distinctly Visible on metapleuron, hardly Visible on dorsal surface; first abdominal tergum inclined, with two lateral complete longitudinal keels and median area with sculpture not distinctly Visible (median area apparently smooth); first abdominal tergum longer than metapostnotum (20:14). Shape of pronotum, mesoscutum, mesoscutellum, metanotum and metapectalpropodeal complex as usual for Dryininae . Forewing apparently completely darkened, with usual Venation of Dryininae ; pterostigma narrow, much longer than broad (35:4,5); marginal cell open (but apparently almost closed); distal part of stigmal Vein longer than proximal part (length of distal part not distinctly Visible, because full of air; distal part apparently much longer than proximal part (39:9); stigmal Vein not S-shaped, angled between proximal and distal parts; forewing with costal, median and submedian cells clearly enclosed by pigmented Veins. Hind wing apparently hyaline, without dark transVerse bands; hind wing shape usual for Dryininae . Proleg ratio: 28 (coxa): 31 (trochanter): 49 (femur): 40 (tibia): 25 (protarsomere 1): 4 (protarsomere 2): 8 (protarsomere 3): 15 (protarsomere 4): 34 (protarsomere 5). Enlarged claw slightly shorter than protarsomere 5 (30:34). Protrochanter slender and long, Very curVed, with long proximal stalk, broadening distally; protrochanter approximately 8 as long as broad (30:6) (maximum breadth measured on distal club). Protarsomeres 2 and 3 produced into hooks. Rudimentary claw present. Arolium much shorter than enlarged claw (6:30). Enlarged claw ( Figs 4 , 5 ) with two small subapical teeth and one row of three lamellae and four bristles (in holotype; seVen lamellae in specimen from RoVno amber); subapical teeth of enlarged claw slightly pointed; distal apex of enlarged claw apparently slightly spatulate. Protarsomere 5 apparently with ten lamellae (in holotype and in specimen in CHWH (N. 1138–1) it is impossible to see if there are 1 or 2 rows of lamellae; howeVer, in specimens in CGCG (N. 3010) and from RoVno amber it is possible to see that there is only one row of eight lamellae ( Figs 4 , 5 ); distal apex of enlarged claw apparently slightly spatulate. Distal apex of protarsomere 5 with group of approximately 21 lamellae. Mesoleg ratio: 15 (coxa): 6 (trochanter): 35 (femur): 38 (tibia): 24 (mesotarsomere 1): 10 (mesotarsomere 2): 7 (mesotarsomere 3): 5 (mesotarsomere 4): 7 (mesotarsomere 5). Metaleg ratio: 26 (coxa): 8 (trochanter): 55 (femur): 54 (tibia): 24 (metatarsomere 1): 11 (metatarsomere 2): 10 (metatarsomere 3): 5 (metatarsomere 4): 7 (metatarsomere 5). Petiole shape and length usual for Dryininae . Shape, length and breadth of wings as usual for Dryininae . Shape and morphology of body as usual for Dryininae . Tibial spurs 1:1:2.

Male. Unknown.

Material examined. Type: ♀ holotype in Late Eocene Baltic amber ( Succinite ) originally in JWJC, now deposited in AMNH . Other specimens: in Baltic amber: 1 ♀ ( No. 1138–1) in CHWH (to be donated to DEI) ; 1 ♀ ( No. 3010) in CGCG ; 1 ♀ (SMNS); in Rovno amber: 1 ♀ (L-29, RoVno Region, Vladimirets District , 10 km N of Voronki Village) ( SIZK) .

Locality and age. Eocene Baltic and RoVno amber (35–37 Ma).

Figures

FIGURE 5. Dryinus janzeni OlmI, ROvNO AmbEr: ChElA ANd prOtArSOmErES.

FIGURES 2–4. Dryinus janzeni OlmI, ROvNO AmbEr: 2–3. HAbItUS IN lAtErAl vIEW; 4. ChElA ANd prOtArSOmErES.