Hemicryphalus Schedl, 1963b: 264

Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You & Jordal, Bjarte H., 2020, Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae), Insect Systematics and Diversity 4 (3), No. 1, pp. 1-81 : 60

Hemicryphalus Schedl, 1963b: 264 ( Fig. 30 )

Type of genus

Eidophelus argutus Wood, 1960 .

Diagnosis

Hemicryphalus can be diagnosed by the broadly emarginated, slightly tapered eye shape, by the straight or weakly procurved sutures on the antennal club, by the lateral margin of the pronotum which has a fine carina, and by the proventriculus which has clusters of long, giant spines.

Female

Eye broadly emarginated. Frons with abundant plumose setae. Antennal funicle 4-segmented. Antennae with straight or slightly procurved sutures, with an incomplete septum on both sides of the frist suture. Pronotum glossy. Pronotum with distinct raised line marking the lateral edge. Pronotum width slightly less than the elytra. Elytra with sparse vestiture. Under the elytra, the cuticle is poorly sclerotized, almost transparent. Proventriculus with short apical plate. Crop spines in bundles, not distributed with radial symmetry and is a mixture of hair-like and large sclerotized setae with multiple spines on its tip.

Male

Externally similar to female except with much less elaborate vestiture on the frons. Penis apodemes much shorter than penis body, fused at apex. Tegmen open dorsally, and of a similar thickness to penis apodemes. A very small median tegminal apodeme is present. Spiculum gastrale thicker than penis apodemes, with a fork. Basal sclerites visible.

Distribution

Pacific islands.

Remarks

Three species known. Superficially similar to some Eidophelus formerly in Ernoporicus , and in Eidophelus sensu Wood and Bright 1992 , which can be distinguished by the antennae in Hemicryphalus having straight sutures with complete septa versus procurved sutures with or without a partial septum, by the eye shape being broadly emarginated instead of oval shaped, and most reliably by the proventriculus having clusters of large, rod-like crop spines instead of short, tooth-like crop spines.

Several species described in this genus from Borneo are transferred to Eidophelus , based on the procurved antennal sutures, overall shape and a close similarity to an undescribed species from montane Java, which is confirmed by molecular data and internal morphology to be nested among other Eidophelus .

Type material examined

Holotype Eidophelus atomus Wood 1960 ( USNM); Two paratypes Eidophelus argutus Wood 1960 ( USNM).

Included species

Hemicryphalus argutus ( Wood, 1960: 33) (Eidophelus) .

Hemicryphalus atomus ( Wood, 1960: 34) (Eidophelus) .

Hemicryphalus incomptus ( Wood, 1960: 32) (Eidophelus) .

Figures

Figure 30. Images of Hemicryphalus argutus: Dorsal and lateral photographs of A) male and B) female, eye,and antennae of A) male and B) female,E) aedeagus, and F) proventriculus.