Macrocryphalus Nobuchi, 1981: 14

Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You & Jordal, Bjarte H., 2020, Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae), Insect Systematics and Diversity 4 (3), No. 1, pp. 1-81 : 39

Macrocryphalus Nobuchi, 1981: 14 stat. res.

( Fig. 19 )

Type of genus

Macrocryphalus oblongus Schedl, 1965 .

Diagnosis

The teardrop-shaped antennae, rounded lateral margins of the pronotum, separated mesocoxae, and absence of a sclerotized proventriculus diagnose members of this genus.

Female

Eye acutely emarginated. Frons with an abrupt concavity lined with long, golden setae which converge toward its median. Antennae with five funicle segments. Antennal club flattened, teardrop-shaped (narrower at base). Antennae with a single procurved suture. Pronotum armed with five to seven marginal asperities, and more than 30 asperities on the pronotal declivity, leading to a slightly elevated summit.

Lateral margins of pronotum rounded. Vestiture of pronotum is hair-like to slightly flattened; setae on the hypomeron simple. Scutellum visible. Elytra simple and convex. Striae not visible. Interstrial bristles slightly flattened, and ground vestiture and strial setae presumably intermixed, confused and hair-like. No sclerotized structure (crop and proventriculus) visible in the upper gut.

Male

Similar except the vestiture of the frons is much shorter. Most males are of a similar size to females and have functional wings. Penis apodemes of a similar length or shorter than penis body, separated at apex.Tegmen without apodemes, open dorsally. Spiculum gastrale of a similar diameter to penis apodemes. Basal sclerites complex and developed.

Distribution

East Asia, East Africa.

Remarks

Three species known. The phylogenetic placement of this genus remains unknown. Molecular data, obtained after the analyses, confirm the placement in Trypophloeini. The transfer of two African

species from Afromicracis Schedl, 1959 to this genus is dubious and based on external appearance only.

Type material examined

Holotype of Miocryphalus elongatus Schedl, 1965 ( NHMW); holotype of Macrocryphalus oblongus Nobuchi, 1981 ( ITLJ); holotype of Miocryphalus punctipennis Schedl, 1965 ( NHMW).

Included species

Macrocryphalus elongatus ( Schedl, 1965c: 366) (Miocryphalus) comb. nov. [ Afromicracis ].

Macrocryphalus oblongus Nobuchi, 1981: 14 stat. res. [ Hypothenemus ].

= Hypothenemus nobuchii Knížek, 2011: 86 syn. nov. (unnecessary replacement name)

Macrocryphalus punctipennis ( Schedl, 1965c: 367) (Miocryphalus) comb. nov. [ Afromicracis ].

Figures

Figure 19. Images of Macrocryphalus oblongus: Dorsal and lateral photographs of A) male and B) female, D) Antennae of female, and D) aedeagus.