Tropidolaemus laticinctus Kuch, Gumprecht & Melaun, 2007

Vogel, Gernot, David, Patrick, Lutz, Mario, Rooijen, Johan Van & Vidal, Nicolas, 2007, Revision of the Tropidolaemus wagleri-complex (Serpentes: Viperidae: Crotalinae). I. Definition of included taxa and redescription of Tropidolaemus wagleri (Boie, 1827), Zootaxa 1644 (1), pp. 1-40 : 11-12

Tropidolaemus laticinctus Kuch, Gumprecht & Melaun, 2007

Tropidolaemus laticinctus Kuch, Gumprecht & Melaun, 2007: 3 , Figs. 1-6 , 7A–B , 8A–B , 9–10 , 11A–B . — Type locality.

“ Between L. Posso and Tomini Bay , Celebes ”, now between Lake Poso and Tomini Bay, Province of Sulawesi Ten-

gah, Indonesia.— Holotype. BMNH 96.12 .9.80 (subadult or adult male). Coll. by P. & F. Sarasin.

Comments. This taxon, briefly described while the present paper was in preparation, was named to accommodate populations identified either as Lachesis wagleri variety E by Boulenger (1896), Tropidolaemus wagleri , “red form” by de Lang & Vogel (2005), or Tropidolaemus subannulatus ( celebensis morph 2) by Vogel (2006).

Status. A valid species, which includes populations from Sulawesi not conspecific with Tropidolaemus subannulatus .

Besides these specific names, several chresonyms resulting from misidentifications were applied to members of the Tropidolaemus wagleri complex:

Trimesurus sumatranus (nec Coluber sumatranus Raffles, 1822 , now Trimeresurus sumatranus or Parias

sumatranus, a valid species): Gray (1842: 48; 1849: 10).

Trigonocephalus formosus (nec Trigonocephalus formosus Müller & Schlegel, 1842 ; see Leviton [1964: 267]): Gray

(1849: 10).

To summarize, the available specific epithets currently available are as follows:

- Tropidolaemus wagleri ( Boie, 1827) : valid combination for western populations, namely Thailand, West Malaysia, Sumatra, Bangka and Mentawei Archipelago.

- Tropidolaemus subannulatus ( Gray, 1842) : valid combination for populations of the Philippine Islands and any, or all of the eastern populations which might be conspecific with Philippine populations, at the exception of Tropidolaemus philippensis and Tropidolaemus laticinctus Kuch, Gumprecht and Melaun, 2007 .

- Tropidolaemus philippensis ( Gray, 1842) : valid combination for some populations of southern and western Mindanao Island, Philippines.

- Tropidolaemus celebensis ( Gray, 1849) : available name for any population inhabiting Sulawesi which would prove to be distinct from Tropidolaemus subannulatus as defined here and from Tropidolaemus laticinctus .

- Tropidolaemus immaculatus (Peters, 1872) : available name for any population inhabiting Borneo which would prove to be distinct from Tropidolaemus subannulatus as defined here.

- Tropidolaemus alboviridis ( Taylor, 1917) : available name for any population inhabiting Negros and adjacent islands, which would prove to be distinct from Tropidolaemus subannulatus as defined here.

- Tropidolaemus laticinctus Kuch, Gumprecht & Melaun, 2007 : valid combination for some populations of Sulawesi, not conspecific with Tropidolaemus subannulatus as defined here.

The splitting of Tropidolaemus wagleri into five subspecies proposed by Iskandar & Collijn (2001) was both premature and unsupported, and partly based on unavailable names. The taxonomy of the Tropidolaemus wagleri complex is now addressed.

Figures

FIGURE 1. Ordination of the OTU’s along the first two principal components based on PCA of quantitative morphological characters.

FIGURE 2. Ordination of the OTU’s along the first two canonical variates based on nonlinear discriminant analysis of morphological and coloration characters.

FIGURE 3. Ordination of the three clusters along the first two canonical variates based on linear discriminant analysis of morphological characters.

FIGURE 4. Neotype of Tropidolaemus wagleri (MNHN 5784). Lateral view of the head, right side. Photograph by Gernot Vogel.

FIGURE 5. Neotype of Tropidolaemus wagleri (MNHN 5784). Lateral view of the head, left side. Photograph by Gernot Vogel.

FIGURE 6. Neotype of Tropidolaemus wagleri (MNHN 5784). Dorsal view of the head. Photograph by Gernot Vogel.

FIGURE 7. Neotype of Tropidolaemus wagleri (MNHN 5784). General view. Photograph by Gernot Vogel.

FIGURE 8. Tropidolaemus wagleri. Living adult female from Jambi Province, Sumatra, Indonesia. Photograph by Gernot Vogel.

FIGURE 9. Tropidolaemus wagleri. Living adult female from Pahang, West-Malaysia. Photograph by Dick Visser.

FIGURE 10. Tropidolaemus wagleri. Living adult male from South Thailand. Photograph by Gernot Vogel.

FIGURE 11. Holotype of Trimesurus maculatus (BMNH 1.2.5a = BMNH 1946.1.17.69). Adult male, Singapore. General view. Photograph by Gernot Vogel.