taxonID	type	description	language	source
03C8F225B26C9C3FFF07F892FDEDFE49.taxon	description	(Figs 1, 2 A – F, SM 1)	en	Santa-Rita, José Vicente Pérez, Dombroskie, Jason J., Ledezma, Julieta, Baixeras, Joaquín (2025): Description of Enicotaenia, a new genus of Archipini from Bolivia, with comments of male-female coupling adaptation (Lepidoptera: Tortricidae). Zootaxa 5613 (2): 361-370, DOI: 10.11646/zootaxa.5613.2.10, URL: https://doi.org/10.11646/zootaxa.5613.2.10
03C8F225B26C9C3FFF07F892FDEDFE49.taxon	type_taxon	Type species. — Enicotaenia interandina Pérez Santa-Rita, Dombroskie, Ledezma & Baixeras, new species	en	Santa-Rita, José Vicente Pérez, Dombroskie, Jason J., Ledezma, Julieta, Baixeras, Joaquín (2025): Description of Enicotaenia, a new genus of Archipini from Bolivia, with comments of male-female coupling adaptation (Lepidoptera: Tortricidae). Zootaxa 5613 (2): 361-370, DOI: 10.11646/zootaxa.5613.2.10, URL: https://doi.org/10.11646/zootaxa.5613.2.10
03C8F225B26C9C3FFF07F892FDEDFE49.taxon	diagnosis	Diagnosis. The forewing pattern is superficially similar to many other Neotropical Archipini, especially in the genera Argyrotaenia and Furcataenia, but is not as sexually dimorphic. The venation shows similarities with Exorstaenia, Ochrotaenia, Saetotaenia and Argyrotaenia, but the combination of R 4 approximated to R 5 at base (but not stalked) in forewing, with Rs and M 1 connate, and M 3 and CuA 1 stalked in hindwing is unique. Overall morphology of the male genitalia shows affinities with Furcataenia and Aphelia (s. s.), but the combination of characters delimits this genus as distinctive. Uncus is bifid in many Claduncaria, Cornuclepsis, some Exorstaenia, Farragona, Tacertaenia and Aphelia (s. s.), but entire in Enicotaenia. A continuous undivided transtilla is only known in Argyrotaenia group. However, the transtilla is relatively simple in most of these genera (Argyrotaenia, Ceritaenia, Exorstaenia, Farragona, Ochrotaenia, Raisapoana and Tacertaenia). A complex transtilla with a medial process is only known in Furcataenia. Variably developed lateral processes from the transtilla are found in some species of Claduncaria, Spinotaenia and Aphelia (s. s.); Enicotaenia has two dorso-lateral horn-shaped processes, which are strongly developed and serrated. Well-defined processes of the sacculus are only reported in Furcataenia and Aphelia (s. s.), but in Enicotaenia the process is smooth, single, and ventrally extended, not serrated. The female genitalia of Enicotaenia are less informative, and no character except the presence of two lateral subconical pockets on the 8 th sternite is autapomorphic. The two depressions present in the sclerotized antrum of Aphelia (see Fig. SM 1) may resemble these pockets but lack apodem-like projections. Enicotaenia does not present a cestum in the ductus bursae, as in other genera such as Clepsis, Sychnovalva, Saetotaenia or Spinotaenia. The antrum of Enicotaenia is rather simple without vestiture, similar to some species of Argyrotaenia, while it is strongly spinulated in other genera such as Furcataenia or Aphelia (s. s.). Enicotaenia has a well-developed signum with capitulum, typically Archipini, as in other related genera such as Argyrotaenia, Farragona, Furcataenia, Idolatteria and Raisapoana. Some genera present a signum as a microthorny sclerite (Chamaepsichia, Rubropsichia or Aphelia (s. s. )), or directly the signum is absent (Claduncaria, Ochrotaenia or Tacertaenia).	en	Santa-Rita, José Vicente Pérez, Dombroskie, Jason J., Ledezma, Julieta, Baixeras, Joaquín (2025): Description of Enicotaenia, a new genus of Archipini from Bolivia, with comments of male-female coupling adaptation (Lepidoptera: Tortricidae). Zootaxa 5613 (2): 361-370, DOI: 10.11646/zootaxa.5613.2.10, URL: https://doi.org/10.11646/zootaxa.5613.2.10
03C8F225B26C9C3FFF07F892FDEDFE49.taxon	description	Description. Head: Typically tortricoid. Vertex with long scales. Frons slightly convex and covered by scales in upper part. Antennae length ca. 0.5 as long as forewing costa, dorsally scaled, ventrally ciliated, cilia less 0.5 times width of flagellomere, males with more numerous cilia than females, two rows of scales per flagellomere. Maxillary palpus undetectable. Labial palpus porrect, length (all three segments combined) same as diameter of compound eye, uniformly scaled. Proboscis well developed, naked. Ocelli and chaetosemata well developed, evident. Thorax: Smooth-scaled including tegulae, with no tufts. Legs unmodified, male foreleg hairpencil absent. Wings with a venation typically for Archipini (Fig. 1). Forewing with a complete venation (based on one slide); M-stem absent, chorda and discal cross-vein obsolescent; discal cell ca. 0.7 times length of wing; all veins present and well defined except CuP reduced, retained only in terminal part; R 4 to costa near apex, R 5 to termen; distance between pairs of veins on termen between R 5 and CuA 1 relatively constant but distance between CuA 1 and CuA 2 slightly larger; anal veins bifurcate in basal area of wing, anal loop ca. 0.4 times length of 1 A + 2 A; costal fold absent. Hindwing with no tubular M-stem in discal cell; vein Sc + R 1 somewhat parallel to Rs basally, length ca. 0.7 times length of wing; Rs and M 1 stalked, parallel basally, ca. 0.4 times length of wing; M 3 and CuA 1 connate; CuA 2 well developed; CuP reduced, present only in distal portion; 1 A + 2 A sinuous, anal loop ca. 0.2 times length of 1 A + 2 A; 3 A developed. Frenulum in females with three bristles and males with one. Male Abdomen: Segment 8 in males with lateral long brush of scales projected caudad covering genitalia. Male genitalia (Fig. 2 C – D) (based on three preparations) with tegumen strongly sclerotized and developed; uncus long (length ca. 0.5 as long as valva) simple clearly differentiated from tegumen; socii weakly developed, sclerotized, with some few long seta on each lobe; arms of gnathos sclerotized, mesally fused, with short dorsal serrations; transtilla continuous, well sclerotized, ventrolaterally with paired elongate serrated processes projecting caudad; valva regular, slightly sclerotized except for the sacculus, costa concave, cucullus rounded, hairy; sacculus well sclerotized, with presence of a fine ventral tooth (digitus) ca. 2 / 3 length of costa; juxta fused with caulis forming a subrectangular plate; phallus with coecum penis straight, distal portion strongly curved down, dorsally minutely serrated; vesica simple, globous, with no cornuti (no cornuti sockets; no deciduous cornuti detected in female genitalia corpus bursae). Female Abdomen: Segment 7 in females without modified scaling (corethrogyne); Female genitalia (Fig. 2 E – F) (based on two preparations) typically Archipini; ostium ringlike; subrectangular sterigma, concave in middle anteriorly, slightly sclerotized on posterior edge, lamella antevaginalis with two differentiated ventral pockets and two lateral depressions (sternite 8) each internally projected in a short apodeme. Antrum short; ductus bursae approximately the length of the corpus bursae, with distinct junction between corpus bursae and ductus bursae; ductus seminalis postmedial; bulla seminalis well-defined; corpus bursae ovoid; signum well developed, elongated with the presence of interior dagger-shaped spine, length more than 0.5 width of the corpus bursae, an external capitulum well developed (one or two globular spermatophores were extracted from the corpus bursae). Anal papillae large, well developed, mesally constricted and covered by sparse long setae, posterior apophysis shorter than anterior apophysis.	en	Santa-Rita, José Vicente Pérez, Dombroskie, Jason J., Ledezma, Julieta, Baixeras, Joaquín (2025): Description of Enicotaenia, a new genus of Archipini from Bolivia, with comments of male-female coupling adaptation (Lepidoptera: Tortricidae). Zootaxa 5613 (2): 361-370, DOI: 10.11646/zootaxa.5613.2.10, URL: https://doi.org/10.11646/zootaxa.5613.2.10
03C8F225B26C9C3FFF07F892FDEDFE49.taxon	etymology	Etymology. The genus name combines the related genus name Argyrotaenia with the classic Greek adjective enikós (ενικός), meaning “ singular ”, in reference to the unusual combination of characters present in the new genus. It is to be considered as feminine in gender.	en	Santa-Rita, José Vicente Pérez, Dombroskie, Jason J., Ledezma, Julieta, Baixeras, Joaquín (2025): Description of Enicotaenia, a new genus of Archipini from Bolivia, with comments of male-female coupling adaptation (Lepidoptera: Tortricidae). Zootaxa 5613 (2): 361-370, DOI: 10.11646/zootaxa.5613.2.10, URL: https://doi.org/10.11646/zootaxa.5613.2.10
03C8F225B26E9C39FF07FB48FC09FCF1.taxon	description	urn: lsid: zoobank. org: pub: 0 EB 9272 D- 4 A 57 - 48 DA-AD 47 - 972 D 519 B 9876 (Fig 2 A – F)	en	Santa-Rita, José Vicente Pérez, Dombroskie, Jason J., Ledezma, Julieta, Baixeras, Joaquín (2025): Description of Enicotaenia, a new genus of Archipini from Bolivia, with comments of male-female coupling adaptation (Lepidoptera: Tortricidae). Zootaxa 5613 (2): 361-370, DOI: 10.11646/zootaxa.5613.2.10, URL: https://doi.org/10.11646/zootaxa.5613.2.10
03C8F225B26E9C39FF07FB48FC09FCF1.taxon	materials_examined	Type material. Holotype. [BOLIVIA] ♂, Santa Cruz Department, Florida Province, Pampa Grande Municipality, locality of El Millu, 1534 m, S 17 º 59 ’ 21.6 ”; W 64 º 03 ’ 14.5 ”, 28 Jan 2011, J. Baixeras, A. Valdivia and G. Fernández (GS 20681) - coll. MNKM. Paratypes. [BOLIVA] 3 ♂, 2 ♀, Santa Cruz Department, Florida Province, Pampa Grande Municipality, locality of El Millu, 1534 m, S 17 º 59 ’ 21.6 ”; W 64 º 03 ’ 14.5 ”, 28 Jan 2011, J. Baixeras, A. Valdivia and G. Fernández (GS 20680, 20717, 20720) (DNA collection codes: DNA 2023006, DNA 2023033, DNA 2023038). 1 male and 1 female deposited in MNKM, 2 males and 1 female deposited in ICBiBE.	en	Santa-Rita, José Vicente Pérez, Dombroskie, Jason J., Ledezma, Julieta, Baixeras, Joaquín (2025): Description of Enicotaenia, a new genus of Archipini from Bolivia, with comments of male-female coupling adaptation (Lepidoptera: Tortricidae). Zootaxa 5613 (2): 361-370, DOI: 10.11646/zootaxa.5613.2.10, URL: https://doi.org/10.11646/zootaxa.5613.2.10
03C8F225B26E9C39FF07FB48FC09FCF1.taxon	diagnosis	Diagnosis. This species may be distinguished from other Archipini by the characters discussed in the diagnosis of the genus. Only two species are known in the genus Enicotaenia, Enicotaenia marabana (Razowski & Becker, 2000) comb. n. and E. interandina here described, both extremely similar. The male of E. marabana is unknown, and the female is only known from the holotype, a single worn specimen from Brazil. The ductus bursae is rather broad in E. interandina and appears narrower in E. marabana. The anterior hypophysis is twice as long in E. interandina compared to E. marabana.	en	Santa-Rita, José Vicente Pérez, Dombroskie, Jason J., Ledezma, Julieta, Baixeras, Joaquín (2025): Description of Enicotaenia, a new genus of Archipini from Bolivia, with comments of male-female coupling adaptation (Lepidoptera: Tortricidae). Zootaxa 5613 (2): 361-370, DOI: 10.11646/zootaxa.5613.2.10, URL: https://doi.org/10.11646/zootaxa.5613.2.10
03C8F225B26E9C39FF07FB48FC09FCF1.taxon	description	Description. Head: Frons, vertex, labial palpus and antenna orangish; presence of dark scales dorsally on labial palpus and antenna. Thorax: Dorsum orangish with the presence of groups of dark scales mixed; forewing length 5.7 – 6.6 mm (x ̄ = 6.2; n = 4) in males, 7.0 – 7.4 mm (x ̄ = 7.2; n = 2) in females. Forewing pattern (Fig. 2 A – B) not sexually dimorphic. Forewing upperside general background colour from ochreous to orangish with scattered dark brown marks, without distinct system of strigulae; system of fasciae incomplete; basal and subbasal fasciae only expressed on dorsum as a dark brown transverse suffused area; median fascia brown, broader on costa, confluent to termen, narrowly fragmented across the wing; postmedian and preterminal fasciae fused as a preapical triangular spot; fringe dark in apical area, lighter in termen and tornus; interfascial areas light brown. Forewing underside uniformly creme, no system of strigulae. Hindwing upperside and underside fairly concolorous, uniformly light orangish brown. Female wing pattern similar to male, no obvious dimorphism. Abdomen: Dorsad orangish, brownish cephalad, paler toward the caudal segments. Male and female genitalia as described for the genus. Molecular characterization. We were able to obtain sequences for small fragments of COI from three specimens. Two identical small fragments of 164 bp were obtained from one male and one female specimens using the primer pair combination C _ TypeF 1 and C _ TypeR 1. Another small fragment of 307 bp was obtained from one female specimen using the pair combination LepF 1 and MLepR 1 primers. The sequences are publicly available through GenBank accession numbers PV 105885, PV 241496 and PV 241497.	en	Santa-Rita, José Vicente Pérez, Dombroskie, Jason J., Ledezma, Julieta, Baixeras, Joaquín (2025): Description of Enicotaenia, a new genus of Archipini from Bolivia, with comments of male-female coupling adaptation (Lepidoptera: Tortricidae). Zootaxa 5613 (2): 361-370, DOI: 10.11646/zootaxa.5613.2.10, URL: https://doi.org/10.11646/zootaxa.5613.2.10
03C8F225B26E9C39FF07FB48FC09FCF1.taxon	biology_ecology	Biology and Distribution. Early stages are unknown; adults have been collected in January (n = 5). Adults were collected at middle elevation (1534 m) in Bolivia, Santa Cruz Department, Florida Province, in municipality of Pampa Grande, and locality of El Millu. The collecting sites include transitions between Bolivian montane dry forest and the slopes of the northern cloud forests of Yungas ecoregion.	en	Santa-Rita, José Vicente Pérez, Dombroskie, Jason J., Ledezma, Julieta, Baixeras, Joaquín (2025): Description of Enicotaenia, a new genus of Archipini from Bolivia, with comments of male-female coupling adaptation (Lepidoptera: Tortricidae). Zootaxa 5613 (2): 361-370, DOI: 10.11646/zootaxa.5613.2.10, URL: https://doi.org/10.11646/zootaxa.5613.2.10
03C8F225B26E9C39FF07FB48FC09FCF1.taxon	etymology	Etymology. The specific epithet refers to the location of the collecting place, an area between inter-Andean dry valleys of the Bolivian montane dry forest and the slopes of the northern Bolivian Andes, a northern Yungas forest. This location has enormous interest from a conservation perspective.	en	Santa-Rita, José Vicente Pérez, Dombroskie, Jason J., Ledezma, Julieta, Baixeras, Joaquín (2025): Description of Enicotaenia, a new genus of Archipini from Bolivia, with comments of male-female coupling adaptation (Lepidoptera: Tortricidae). Zootaxa 5613 (2): 361-370, DOI: 10.11646/zootaxa.5613.2.10, URL: https://doi.org/10.11646/zootaxa.5613.2.10
03C8F225B2689C39FF07FCA2FC88FAB4.taxon	discussion	Razowski & Becker (2000) described the genus Furcataenia to include a series of species from Brazil with bizarre male genitalia. They also described F. marabana, a species based on a single worn female. In the female genitalia drawing by Razowski & Becker (2000) the ductus bursae is presumably rotated. The signum appears on the left part of the corpus bursae and should be on the right side. Anyway, the female genitalia are similar in both species, and they both appear to have the A 8 pockets. It seems reasonable to assume that such a structure is an apomorphy of the genus, suggesting monophyly. The discovery of E. interandina relates both species through the female genitalia, but the male of E. interandina is not assignable to Furcataenia. We propose the description of a separate genus, Enicotaenia, to include the new species together with E. marabana, closely related species. Because the information on E. interandina is more complete, we propose it as the type species for the genus.	en	Santa-Rita, José Vicente Pérez, Dombroskie, Jason J., Ledezma, Julieta, Baixeras, Joaquín (2025): Description of Enicotaenia, a new genus of Archipini from Bolivia, with comments of male-female coupling adaptation (Lepidoptera: Tortricidae). Zootaxa 5613 (2): 361-370, DOI: 10.11646/zootaxa.5613.2.10, URL: https://doi.org/10.11646/zootaxa.5613.2.10
