taxonID	type	description	language	source
03C95C3FF221074AFE98C1D2C962F9E9.taxon	type_taxon	Philoscia squamuligera Koelbel, 1895, here designated.	en	Gardini, Pietro, Sabatelli, Simone, Taiti, Stefano, Audisio, Paolo (2025): Hidden species’ diversitv and phvlogenetic relationships within the terrestrial isopod genus Tiroloscia (Crustacea: Isopoda: Oniscidea). Zoological Journal of the Linnean Society 203 (1): 1-17, DOI: 10.1093/zoolinnean/zlae173, URL: https://doi.org/10.1093/zoolinnean/zlae173
03C95C3FF221074BFF79C27EC974FA12.taxon	description	(Figs 2 A, 4 A – L)	en	Gardini, Pietro, Sabatelli, Simone, Taiti, Stefano, Audisio, Paolo (2025): Hidden species’ diversitv and phvlogenetic relationships within the terrestrial isopod genus Tiroloscia (Crustacea: Isopoda: Oniscidea). Zoological Journal of the Linnean Society 203 (1): 1-17, DOI: 10.1093/zoolinnean/zlae173, URL: https://doi.org/10.1093/zoolinnean/zlae173
03C95C3FF221074BFF79C27EC974FA12.taxon	materials_examined	Type locality: Artemare, Ain department, France (Fig. 1, site 13). Material examined: Lineage L 1. Italy — Piedmont: 4 ♂♂, Sampeyre (CN), loc. Dragoniere (site 6), 1100 m a. s. l., 03. I. 2020, P. Gardini leg.; 2 ♂♂, Sampeyre (CN), loc. Rore (site 7), beech forest, 1300 m a. s. l., G. and P. Gardini and S. Zoia leg. Lineage L 2. Italy — Piedmont: 1 ♂, Chiusa di Pesio (CN), Pian delle Gorre (site 1), 1050 m a. s. l., 11. VIII. 2020, P. Gardini leg. Lineage L 3. Italy — Piedmont: 1 ♂, Sampeyre (CN), Meire Biun (site 4), 1530 m a. s. l., 03. VIII. 2020, G. and P. Gardini leg.; 2 ♂♂, Sampeyre (CN), Pian Ciattiva (site 5), 1800 m a. s. l., 07. VIII. 2020, P. Gardini leg.; 1 ♂, Sampeyre (CN), loc. Dragoniere (site 6), 1100 m a. s. l., 03. I. 2020, P. Gardini leg. Lineage L 4. Italy — Piedmont: 2 ♂♂, Limone Piemonte (CN), S. Maurizio (site 2), 1020 m a. s. l., 05. VII. 2021, G. B. Delmastro leg.; 1 ♂, Trinità di Entracque (CN) (site 3), beech forest, 1300 m a. s. l., 17. VIII. 2020, P. Gardini leg.; 2 ♂♂, Barge (CN), loc. Lungaserra (site 8), chestnut forest, 630 m a. s. l., 01. VI. 2021, P. Gardini leg.; 3 ♂♂, Revello (CN), Staffarda (site 9), 265 m a. s. l., 27. V. 2021, G. B. Delmastro leg. France — Savoie: 7 ♂♂, Apremont, Col du Granier (site 12). Ain: 4 ♂♂, Chanay, Col de Richemond (site 14), beech forest, 1140 m a. s. l., 19. IX. 2020, G. and P. Gardini and C. Giusto leg.; 3 ♂♂, Valserhône (site 15), 19. IX. 2020, G. and P. Gardini and C. Giusto leg. Previous records: France — Ain: Artemare (Budde-Lund 1885); Mont Rond (Carl 1908); Labalme, near Poncin; Virieu-le-Grand (Vandel 1962). Alpes-Maritimes: Saint-Dalmas-de-Tende (Verhoeff 1931 a); Caussols; Fontan (Vandel 1962). Drôme: Die (Vandel 1962). Haute-Savoie: Chamonix (Dollfus 1897, 1899, Legrand 1950); Saint-Cergues (Carl 1908); Le Chatelard; Les Bossons; La Giettaz; La Clusaz; Col de Leschaux; Flumet; Mégevette (Legrand 1950). Hautes-Alpes: Gap; Château-Queyras. Isère: Dent-de-Crolles. Jura: Villards-d’Héria (Dollfus 1897, 1899); Viry; Choux; Vulvoz; Molinges (Vandel 1946). Rhône: Saint-André-la-Côte (Vandel 1962). Italy — Piedmont: Limone Piemonte; Ormea; Villanova Mondovì (Verhoeff 1931 a). Gran Paradiso National Park: Gran Piano di Noasca, Alpe Bruna, Vallone Noaschetta (Arcangeli 1932). Ala di Stura; Barge; Ceres;? Cossila; Crissolo; Pinerolo (Verhoeff 1936). Switzerland — Bois de Saint-Livres, near Bière; Bois-de-Bay, near Vernier; Prangins, near Nyon; Veyrier (Carl 1908); Salvan (Vandel 1962).	en	Gardini, Pietro, Sabatelli, Simone, Taiti, Stefano, Audisio, Paolo (2025): Hidden species’ diversitv and phvlogenetic relationships within the terrestrial isopod genus Tiroloscia (Crustacea: Isopoda: Oniscidea). Zoological Journal of the Linnean Society 203 (1): 1-17, DOI: 10.1093/zoolinnean/zlae173, URL: https://doi.org/10.1093/zoolinnean/zlae173
03C95C3FF221074BFF79C27EC974FA12.taxon	description	Description Male pleopod 1 Lineage L 1 (Fig. 4 A – C): Exopod cordiform, with well-developed but rather narrow posterior tip, rounded apex with fine setae; inner margin straight in distal half, outer margin concave with three setae in proximal part; width / length ratio 1.25. Endopod with sinuous distal part provided with row of short, strong setae; apical part distinctly bilobed with two triangular lobes protruding distally, large and broadly rounded inner lobe; distal part strongly bent outwards when observed on whole animal. Lineage L 2 (Fig. 4 D – F): Exopod cordiform, with well-developed posterior tip, rounded apex; inner margin slightly concave in distal half, outer margin concave with three setae in proximal part; width / length ratio 1.27. Endopod with parallel-sided distal part, with slightly sinusoidal row of short setae; apical part triangular, bilobed, with small, rounded subapical lobe on outer margin. Lineage L 3 (Fig. 4 G – I): Exopod cordiform, with broadly rounded posterior tip; inner margin convex, provided with fine setae in distal half, outer margin straight with three setae in proximal part; width / length ratio 1.25. Endopod with triangular distal part, with slightly sinusoidal row of short setae; apical part with two lobes, outer one much shorter than inner one, apex rounded, almost straight when observed on whole animal. Lineage L 4 (Fig. 4 J-L): Exopod cordiform, with well-developed, broadly rounded posterior tip; inner margin convex, outer margin slightly concave with three setae in proximal part; width / length ratio 1.20. Endopod with sinuous distal part provided with row of short, strong setae; apical part bilobed with large, broadly rounded inner lobe; distal part strongly bent outwards when observed on whole animal.	en	Gardini, Pietro, Sabatelli, Simone, Taiti, Stefano, Audisio, Paolo (2025): Hidden species’ diversitv and phvlogenetic relationships within the terrestrial isopod genus Tiroloscia (Crustacea: Isopoda: Oniscidea). Zoological Journal of the Linnean Society 203 (1): 1-17, DOI: 10.1093/zoolinnean/zlae173, URL: https://doi.org/10.1093/zoolinnean/zlae173
03C95C3FF221074BFF79C27EC974FA12.taxon	discussion	Remarks Specimens belonging to the four lineages show a certain degree of morphological differentiation and can be distinguished based on the fine structure of the male pleopod 1 endopod. Lineage L 1 is readily distinguished from the others by the distinctly bilobed apical part of the endopod, with two clearly visible triangular lobes, with the one on the outer face highly developed. It is similar to L 4 in having the distal part of the endopod distinctly sinuous, but the latter is readily distinguished by the much less developed apical lobes. Lineage L 2 is similar to L 1 in having a rounded subapical lobe on the outer margin of the male pleopod 1 endopod, although much less developed, but is easily distinguished from both L 1 and L 4 by having the distal part of the endopod almost straight with parallel sides and no broadly rounded inner lobe. Lineage L 3 is readily distinguished from L 1 and L 4 in having the male pleopod 1 endopod almost straight, with two less developed apical lobes, of which the outer one much shorter than the inner one. It differs from L 2 in having a rounded apical part rather than a triangular one, and in the absence of a subapical lobe.	en	Gardini, Pietro, Sabatelli, Simone, Taiti, Stefano, Audisio, Paolo (2025): Hidden species’ diversitv and phvlogenetic relationships within the terrestrial isopod genus Tiroloscia (Crustacea: Isopoda: Oniscidea). Zoological Journal of the Linnean Society 203 (1): 1-17, DOI: 10.1093/zoolinnean/zlae173, URL: https://doi.org/10.1093/zoolinnean/zlae173
03C95C3FF220074BFF61C1C4CCD8FA58.taxon	description	(Figs 2 B, 4 M – O)	en	Gardini, Pietro, Sabatelli, Simone, Taiti, Stefano, Audisio, Paolo (2025): Hidden species’ diversitv and phvlogenetic relationships within the terrestrial isopod genus Tiroloscia (Crustacea: Isopoda: Oniscidea). Zoological Journal of the Linnean Society 203 (1): 1-17, DOI: 10.1093/zoolinnean/zlae173, URL: https://doi.org/10.1093/zoolinnean/zlae173
03C95C3FF220074BFF61C1C4CCD8FA58.taxon	materials_examined	Type locality: Razzes, South Tyrol, Italy (Fig. 1, site 26). Material examined: Lineage L 5. Italy — Piedmont: 4 ♂♂, Nichelino (TO), Park of Stupinigi (site 10), 240 m a. s. l., 02. VI. 2021, G. B. Delmastro leg.; 3 ♂♂, Castell’Alfero (AT), loc. Callianetto (site 11), 180 m a. s. l., 28. IX. 2020, G. Gardini leg.; 6 ♂♂, Alice Castello (VC) (site 16), 290 m a. s. l., 29. V. 2021, G. and P. Gardini leg.; 1 ♂, Campiglia Cervo (BI) (site 17), beech forest, 860 m a. s. l., 28. V. 2021, G. and P. Gardini leg.; 1 ♂, Varallo (VC), loc. Crosa (site 18), 670 m a. s. l., 28. V. 2021, G. and P. Gardini leg.; 4 ♂♂, Gignese (VB) (site 19), chestnut forest, 680 m a. s. l., 27. V. 2021, G. and P. Gardini leg .. Lombardy: 1 ♂, Valganna (VA) (site 20), 460 m a. s. l., 26. V. 2021, G. and P. Gardini leg.; 1 ♂, S. Fedele Intelvi (CO), Monte Generoso (site 21), beech forest, 985 m a. s. l., 25. V. 2021, G. and P. Gardini leg.; 6 ♂♂, Schilpario (BG), loc. Ronco (site 22), 1100 m a. s. l., 05. VIII. 2021, P. Gardini leg. Trentino-South Tyrol: 1 ♂, Pellizzano (TN), loc. Claiano (site 23), 1100 m a. s. l., 07. VIII. 2021, P. Gardini leg.; 3 ♂♂, Ville d’Anaunia (TN), Lago di Tovel (site 24), 1050 m a. s. l., 07. VIII. 2021, P. Gardini leg.; 3 ♂♂, Soprabolzano (BZ) (site 25), beech forest, 1000 m a. s. l., 08. VIII. 2021, P. Gardini leg.; 1 ♂, Siusi (BZ), Razzes (site 26), 1130 m a. s. l., 08. VIII. 2021, P. Gardini leg. Previous records: Italy — Piedmont: Cossila (Verhoeff 1936). Lombardy: Luino; Laveno (Verhoeff 1928, 1931 a); Edolo (Verhoeff 1931 b). Trentino-South Tyrol: Bressanone (Verhoeff 1908 a, 1908 b, 1928); Vipiteno (Verhoeff 1934).	en	Gardini, Pietro, Sabatelli, Simone, Taiti, Stefano, Audisio, Paolo (2025): Hidden species’ diversitv and phvlogenetic relationships within the terrestrial isopod genus Tiroloscia (Crustacea: Isopoda: Oniscidea). Zoological Journal of the Linnean Society 203 (1): 1-17, DOI: 10.1093/zoolinnean/zlae173, URL: https://doi.org/10.1093/zoolinnean/zlae173
03C95C3FF220074BFF61C1C4CCD8FA58.taxon	description	Description Male pleopod 1. Lineage L 5 (Fig. 4 M – O): Exopod cordiform, with poorly developed, broadly rounded posterior tip; outer margin slightly concave with four setae in proximal part; width / length ratio 1.47. Endopod with triangular distal part, with row of short, slightly sinusoidal setae on inner side, smaller than in the other lineages of the T. exigua complex; outer margin straight, apical part without distinct lobes, apical part rounded.	en	Gardini, Pietro, Sabatelli, Simone, Taiti, Stefano, Audisio, Paolo (2025): Hidden species’ diversitv and phvlogenetic relationships within the terrestrial isopod genus Tiroloscia (Crustacea: Isopoda: Oniscidea). Zoological Journal of the Linnean Society 203 (1): 1-17, DOI: 10.1093/zoolinnean/zlae173, URL: https://doi.org/10.1093/zoolinnean/zlae173
03C95C3FF220074BFF61C1C4CCD8FA58.taxon	discussion	Remarks The material examined corresponds fairly well to the description and figures of the male pleopod 1 endopod of T. squamuligera provided by Verhoeff (1931 a). This species differs from the lineages of the T. exigua species complex in having the male pleopod 1 exopod with a much less developed posterior tip and a greater width / length ratio (1.47 vs. 1.20 – 1.27) and the endopod almost straight without distinct apical lobes, with a row of much more reduced setae. All observed characters were consistent among individuals across all sampled populations.	en	Gardini, Pietro, Sabatelli, Simone, Taiti, Stefano, Audisio, Paolo (2025): Hidden species’ diversitv and phvlogenetic relationships within the terrestrial isopod genus Tiroloscia (Crustacea: Isopoda: Oniscidea). Zoological Journal of the Linnean Society 203 (1): 1-17, DOI: 10.1093/zoolinnean/zlae173, URL: https://doi.org/10.1093/zoolinnean/zlae173
