taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03D2A373FFDAFFAFFF26F9ADFD822CD2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17870967/files/figure.png	https://doi.org/10.5281/zenodo.17870967	FIGURE 16. Phylogenetic placement of Triozidus in family Triozidae is indicated by a black square representing a subgroup within Group B (see Percy et al. 2018), the related genera are expanded on in the text. An asterix indicates the placement of the type species of Heterotrioza, H. chenopodii in Group D, and Trioza, T. urticae in Group M, and Dyspersa sensu Cho et al. (2022) in Group A. Configuration of the three Triozidus taxa to one another based on the 714 bp COI fragment data (unique haplotypes only) is shown, with T. burckhardti placement indicated based on the backbone analysis (see text); taxon placements within Triozidus are uncertain in both analyses due to short internal branches and low to moderate bootstrap support, with a slightly different taxon configuration recovered in the ML backbone constraint analysis (see text). Illustrations of forewing, paramere and aedeagus are shown for all six Triozidus species to indicate comparative sizes; an asterisk by T. ceratophorus and T. eleutherococci indicates the illustrations are redrawn with reference to Li (2011) and Konovalova (1980, 1988) respectively, and in these cases paramere and aedeagus may not be to scale, plus the apex of the aedeagus is partly obscured in the illustration for T. eleutherococci. The annotation of the mitogenome of T. yangorum is graphically represented at top right.	FIGURE 16. Phylogenetic placement of Triozidus in family Triozidae is indicated by a black square representing a subgroup within Group B (see Percy et al. 2018), the related genera are expanded on in the text. An asterix indicates the placement of the type species of Heterotrioza, H. chenopodii in Group D, and Trioza, T. urticae in Group M, and Dyspersa sensu Cho et al. (2022) in Group A. Configuration of the three Triozidus taxa to one another based on the 714 bp COI fragment data (unique haplotypes only) is shown, with T. burckhardti placement indicated based on the backbone analysis (see text); taxon placements within Triozidus are uncertain in both analyses due to short internal branches and low to moderate bootstrap support, with a slightly different taxon configuration recovered in the ML backbone constraint analysis (see text). Illustrations of forewing, paramere and aedeagus are shown for all six Triozidus species to indicate comparative sizes; an asterisk by T. ceratophorus and T. eleutherococci indicates the illustrations are redrawn with reference to Li (2011) and Konovalova (1980, 1988) respectively, and in these cases paramere and aedeagus may not be to scale, plus the apex of the aedeagus is partly obscured in the illustration for T. eleutherococci. The annotation of the mitogenome of T. yangorum is graphically represented at top right.	2025-12-08	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.		Zenodo	biologists	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.			
03D2A373FFDCFFAFFF26FDB8FA142A30.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17870945/files/figure.png	https://doi.org/10.5281/zenodo.17870945	FIGURE 9. Adult Triozidus ukogi comb. nov. A, head and antennae; B, forewing; C, hindwing; D, hind leg. Scale bars: 0.1 mm (A, D); 1 mm (B, C).	FIGURE 9. Adult Triozidus ukogi comb. nov. A, head and antennae; B, forewing; C, hindwing; D, hind leg. Scale bars: 0.1 mm (A, D); 1 mm (B, C).	2025-12-08	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.		Zenodo	biologists	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.			
03D2A373FFDCFFAFFF26FDB8FA142A30.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17870918/files/figure.png	https://doi.org/10.5281/zenodo.17870918	FIGURE 2. Adult Triozidus burckhardti sp. nov. A, head; B, antenna; C, hind leg; D, hindwing; E, forewing. Scale bars: 0.1 mm (A, B, D); 1 mm (C, E).	FIGURE 2. Adult Triozidus burckhardti sp. nov. A, head; B, antenna; C, hind leg; D, hindwing; E, forewing. Scale bars: 0.1 mm (A, B, D); 1 mm (C, E).	2025-12-08	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.		Zenodo	biologists	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.			
03D2A373FFDCFFAFFF26FDB8FA142A30.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17870929/files/figure.png	https://doi.org/10.5281/zenodo.17870929	FIGURE 5. Adult Triozidus stackelbergi comb. nov. A, head; B, antenna; C, forewing, with small dark spot at trifurcation of R+M+Cu (arrowed); D, hindwing; E, hind leg. Scale bars: 0.1 mm (A, B, E); 1 mm (C, D).	FIGURE 5. Adult Triozidus stackelbergi comb. nov. A, head; B, antenna; C, forewing, with small dark spot at trifurcation of R+M+Cu (arrowed); D, hindwing; E, hind leg. Scale bars: 0.1 mm (A, B, E); 1 mm (C, D).	2025-12-08	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.		Zenodo	biologists	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.			
03D2A373FFDCFFAFFF26FDB8FA142A30.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17870961/files/figure.png	https://doi.org/10.5281/zenodo.17870961	FIGURE 13. Adult Triozidus yangorum sp. nov. A, head; B, antenna; C, terminal antennal segments showing uneven length terminal setae (arrowed); D, forewing; E, hindwing; F, hind leg. Scale bars: 0.1 mm (A, B, C, F); 1 mm (D, E).	FIGURE 13. Adult Triozidus yangorum sp. nov. A, head; B, antenna; C, terminal antennal segments showing uneven length terminal setae (arrowed); D, forewing; E, hindwing; F, hind leg. Scale bars: 0.1 mm (A, B, C, F); 1 mm (D, E).	2025-12-08	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.		Zenodo	biologists	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.			
03D2A373FFDCFFAFFF26FDB8FA142A30.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17870967/files/figure.png	https://doi.org/10.5281/zenodo.17870967	FIGURE 16. Phylogenetic placement of Triozidus in family Triozidae is indicated by a black square representing a subgroup within Group B (see Percy et al. 2018), the related genera are expanded on in the text. An asterix indicates the placement of the type species of Heterotrioza, H. chenopodii in Group D, and Trioza, T. urticae in Group M, and Dyspersa sensu Cho et al. (2022) in Group A. Configuration of the three Triozidus taxa to one another based on the 714 bp COI fragment data (unique haplotypes only) is shown, with T. burckhardti placement indicated based on the backbone analysis (see text); taxon placements within Triozidus are uncertain in both analyses due to short internal branches and low to moderate bootstrap support, with a slightly different taxon configuration recovered in the ML backbone constraint analysis (see text). Illustrations of forewing, paramere and aedeagus are shown for all six Triozidus species to indicate comparative sizes; an asterisk by T. ceratophorus and T. eleutherococci indicates the illustrations are redrawn with reference to Li (2011) and Konovalova (1980, 1988) respectively, and in these cases paramere and aedeagus may not be to scale, plus the apex of the aedeagus is partly obscured in the illustration for T. eleutherococci. The annotation of the mitogenome of T. yangorum is graphically represented at top right.	FIGURE 16. Phylogenetic placement of Triozidus in family Triozidae is indicated by a black square representing a subgroup within Group B (see Percy et al. 2018), the related genera are expanded on in the text. An asterix indicates the placement of the type species of Heterotrioza, H. chenopodii in Group D, and Trioza, T. urticae in Group M, and Dyspersa sensu Cho et al. (2022) in Group A. Configuration of the three Triozidus taxa to one another based on the 714 bp COI fragment data (unique haplotypes only) is shown, with T. burckhardti placement indicated based on the backbone analysis (see text); taxon placements within Triozidus are uncertain in both analyses due to short internal branches and low to moderate bootstrap support, with a slightly different taxon configuration recovered in the ML backbone constraint analysis (see text). Illustrations of forewing, paramere and aedeagus are shown for all six Triozidus species to indicate comparative sizes; an asterisk by T. ceratophorus and T. eleutherococci indicates the illustrations are redrawn with reference to Li (2011) and Konovalova (1980, 1988) respectively, and in these cases paramere and aedeagus may not be to scale, plus the apex of the aedeagus is partly obscured in the illustration for T. eleutherococci. The annotation of the mitogenome of T. yangorum is graphically represented at top right.	2025-12-08	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.		Zenodo	biologists	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.			
03D2A373FFDCFFAFFF26FDB8FA142A30.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17870923/files/figure.png	https://doi.org/10.5281/zenodo.17870923	FIGURE 3. Terminalia of Triozidus burckhardti sp. nov. A, male terminalia (lateral view); B, paramere (inner surface); C, distal segment of aedeagus; D, female terminalia (lateral view). Scale bars: 0.1 mm.	FIGURE 3. Terminalia of Triozidus burckhardti sp. nov. A, male terminalia (lateral view); B, paramere (inner surface); C, distal segment of aedeagus; D, female terminalia (lateral view). Scale bars: 0.1 mm.	2025-12-08	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.		Zenodo	biologists	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.			
03D2A373FFDCFFAFFF26FDB8FA142A30.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17870935/files/figure.png	https://doi.org/10.5281/zenodo.17870935	FIGURE 6. Terminalia of Triozidus stackelbergi comb. nov. A, male terminalia (lateral view); B, paramere (inner surface); C, distal portion of aedeagus showing medial semicircular projections (arrowed); D, female terminalia (lateral view). Scale bars: 0.1 mm.	FIGURE 6. Terminalia of Triozidus stackelbergi comb. nov. A, male terminalia (lateral view); B, paramere (inner surface); C, distal portion of aedeagus showing medial semicircular projections (arrowed); D, female terminalia (lateral view). Scale bars: 0.1 mm.	2025-12-08	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.		Zenodo	biologists	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.			
03D2A373FFDCFFAFFF26FDB8FA142A30.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17870963/files/figure.png	https://doi.org/10.5281/zenodo.17870963	FIGURE 14. Terminalia of Triozidus yangorum sp. nov. A, male terminalia (lateral view); B, paramere (inner surface); C, distal portion of aedeagus, showing medial narrow thumb-like projections (arrowed); D, female terminalia (lateral view). Scale bars: 0.1 mm.	FIGURE 14. Terminalia of Triozidus yangorum sp. nov. A, male terminalia (lateral view); B, paramere (inner surface); C, distal portion of aedeagus, showing medial narrow thumb-like projections (arrowed); D, female terminalia (lateral view). Scale bars: 0.1 mm.	2025-12-08	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.		Zenodo	biologists	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.			
03D2A373FFDCFFAFFF26FDB8FA142A30.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17870916/files/figure.png	https://doi.org/10.5281/zenodo.17870916	FIGURE 1. Triozidus burckhardti sp. nov. and Triozidus yangorum sp. nov. adults, immature and galls on the host plant, Eleutherococcus trifoliatus. A, T. burckhardti female; B, closed galls of cf. T. burckhardti with a single gall on each petiolule of the trifoliate leaf; C, T. yangorum male; D, T. yangorum fifth instar immature within gall chamber; E, close up showing three petiolule galls of T. yangorum on a single leaf, the upper already opened; F, the scandent host, E. trifoliatus, showing distribution of petiolule galls.	FIGURE 1. Triozidus burckhardti sp. nov. and Triozidus yangorum sp. nov. adults, immature and galls on the host plant, Eleutherococcus trifoliatus. A, T. burckhardti female; B, closed galls of cf. T. burckhardti with a single gall on each petiolule of the trifoliate leaf; C, T. yangorum male; D, T. yangorum fifth instar immature within gall chamber; E, close up showing three petiolule galls of T. yangorum on a single leaf, the upper already opened; F, the scandent host, E. trifoliatus, showing distribution of petiolule galls.	2025-12-08	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.		Zenodo	biologists	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.			
03D2A373FFDCFFAFFF26FDB8FA142A30.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17870937/files/figure.png	https://doi.org/10.5281/zenodo.17870937	FIGURE 7. Fifth instar immature of Triozidus stackelbergi comb. nov. A, habitus; B, tarsal arolium showing asymmetric development of tarsal claws (vestigial claw arrowed); C, forewing pad marginal sectasetae; D, circumanal ring showing regular single row of pores (inset); E, antenna. Scale bars: 0.1 mm.	FIGURE 7. Fifth instar immature of Triozidus stackelbergi comb. nov. A, habitus; B, tarsal arolium showing asymmetric development of tarsal claws (vestigial claw arrowed); C, forewing pad marginal sectasetae; D, circumanal ring showing regular single row of pores (inset); E, antenna. Scale bars: 0.1 mm.	2025-12-08	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.		Zenodo	biologists	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.			
03D2A373FFDCFFAFFF26FDB8FA142A30.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17870951/files/figure.png	https://doi.org/10.5281/zenodo.17870951	FIGURE 11. Fifth instar immature of Triozidus ukogi comb. nov. A, habitus; B, tarsal arolium showing asymmetric development of tarsal claws (vestigial claw arrowed); C, forewing pad marginal simple setae; D circumanal ring showing single narrow row of elongate pores (inset); E antenna. Scale bars: 0.1 mm.	FIGURE 11. Fifth instar immature of Triozidus ukogi comb. nov. A, habitus; B, tarsal arolium showing asymmetric development of tarsal claws (vestigial claw arrowed); C, forewing pad marginal simple setae; D circumanal ring showing single narrow row of elongate pores (inset); E antenna. Scale bars: 0.1 mm.	2025-12-08	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.		Zenodo	biologists	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.			
03D2A373FFDCFFAFFF26FDB8FA142A30.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17870965/files/figure.png	https://doi.org/10.5281/zenodo.17870965	FIGURE 15. Fifth instar immature of Triozidus yangorum sp. nov. A, habitus; B, tarsal arolium showing asymmetric development of tarsal claws (vestigial claw arrowed); C, forewing pad marginal simple setae; D, circumanal ring showing intermittent irregular pore shape (inset); E, antenna. Scale bars: 0.1 mm.	FIGURE 15. Fifth instar immature of Triozidus yangorum sp. nov. A, habitus; B, tarsal arolium showing asymmetric development of tarsal claws (vestigial claw arrowed); C, forewing pad marginal simple setae; D, circumanal ring showing intermittent irregular pore shape (inset); E, antenna. Scale bars: 0.1 mm.	2025-12-08	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.		Zenodo	biologists	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.			
03D2A373FFDDFFAEFF26FF0CFCB82AB9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17870916/files/figure.png	https://doi.org/10.5281/zenodo.17870916	FIGURE 1. Triozidus burckhardti sp. nov. and Triozidus yangorum sp. nov. adults, immature and galls on the host plant, Eleutherococcus trifoliatus. A, T. burckhardti female; B, closed galls of cf. T. burckhardti with a single gall on each petiolule of the trifoliate leaf; C, T. yangorum male; D, T. yangorum fifth instar immature within gall chamber; E, close up showing three petiolule galls of T. yangorum on a single leaf, the upper already opened; F, the scandent host, E. trifoliatus, showing distribution of petiolule galls.	FIGURE 1. Triozidus burckhardti sp. nov. and Triozidus yangorum sp. nov. adults, immature and galls on the host plant, Eleutherococcus trifoliatus. A, T. burckhardti female; B, closed galls of cf. T. burckhardti with a single gall on each petiolule of the trifoliate leaf; C, T. yangorum male; D, T. yangorum fifth instar immature within gall chamber; E, close up showing three petiolule galls of T. yangorum on a single leaf, the upper already opened; F, the scandent host, E. trifoliatus, showing distribution of petiolule galls.	2025-12-08	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.		Zenodo	biologists	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.			
03D2A373FFDDFFAEFF26FF0CFCB82AB9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17870918/files/figure.png	https://doi.org/10.5281/zenodo.17870918	FIGURE 2. Adult Triozidus burckhardti sp. nov. A, head; B, antenna; C, hind leg; D, hindwing; E, forewing. Scale bars: 0.1 mm (A, B, D); 1 mm (C, E).	FIGURE 2. Adult Triozidus burckhardti sp. nov. A, head; B, antenna; C, hind leg; D, hindwing; E, forewing. Scale bars: 0.1 mm (A, B, D); 1 mm (C, E).	2025-12-08	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.		Zenodo	biologists	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.			
03D2A373FFDDFFAEFF26FF0CFCB82AB9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17870923/files/figure.png	https://doi.org/10.5281/zenodo.17870923	FIGURE 3. Terminalia of Triozidus burckhardti sp. nov. A, male terminalia (lateral view); B, paramere (inner surface); C, distal segment of aedeagus; D, female terminalia (lateral view). Scale bars: 0.1 mm.	FIGURE 3. Terminalia of Triozidus burckhardti sp. nov. A, male terminalia (lateral view); B, paramere (inner surface); C, distal segment of aedeagus; D, female terminalia (lateral view). Scale bars: 0.1 mm.	2025-12-08	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.		Zenodo	biologists	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.			
03D2A373FFDDFFAEFF26FF0CFCB82AB9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17870967/files/figure.png	https://doi.org/10.5281/zenodo.17870967	FIGURE 16. Phylogenetic placement of Triozidus in family Triozidae is indicated by a black square representing a subgroup within Group B (see Percy et al. 2018), the related genera are expanded on in the text. An asterix indicates the placement of the type species of Heterotrioza, H. chenopodii in Group D, and Trioza, T. urticae in Group M, and Dyspersa sensu Cho et al. (2022) in Group A. Configuration of the three Triozidus taxa to one another based on the 714 bp COI fragment data (unique haplotypes only) is shown, with T. burckhardti placement indicated based on the backbone analysis (see text); taxon placements within Triozidus are uncertain in both analyses due to short internal branches and low to moderate bootstrap support, with a slightly different taxon configuration recovered in the ML backbone constraint analysis (see text). Illustrations of forewing, paramere and aedeagus are shown for all six Triozidus species to indicate comparative sizes; an asterisk by T. ceratophorus and T. eleutherococci indicates the illustrations are redrawn with reference to Li (2011) and Konovalova (1980, 1988) respectively, and in these cases paramere and aedeagus may not be to scale, plus the apex of the aedeagus is partly obscured in the illustration for T. eleutherococci. The annotation of the mitogenome of T. yangorum is graphically represented at top right.	FIGURE 16. Phylogenetic placement of Triozidus in family Triozidae is indicated by a black square representing a subgroup within Group B (see Percy et al. 2018), the related genera are expanded on in the text. An asterix indicates the placement of the type species of Heterotrioza, H. chenopodii in Group D, and Trioza, T. urticae in Group M, and Dyspersa sensu Cho et al. (2022) in Group A. Configuration of the three Triozidus taxa to one another based on the 714 bp COI fragment data (unique haplotypes only) is shown, with T. burckhardti placement indicated based on the backbone analysis (see text); taxon placements within Triozidus are uncertain in both analyses due to short internal branches and low to moderate bootstrap support, with a slightly different taxon configuration recovered in the ML backbone constraint analysis (see text). Illustrations of forewing, paramere and aedeagus are shown for all six Triozidus species to indicate comparative sizes; an asterisk by T. ceratophorus and T. eleutherococci indicates the illustrations are redrawn with reference to Li (2011) and Konovalova (1980, 1988) respectively, and in these cases paramere and aedeagus may not be to scale, plus the apex of the aedeagus is partly obscured in the illustration for T. eleutherococci. The annotation of the mitogenome of T. yangorum is graphically represented at top right.	2025-12-08	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.		Zenodo	biologists	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.			
03D2A373FFD0FFA3FF26FA83FCDC2A01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17870967/files/figure.png	https://doi.org/10.5281/zenodo.17870967	FIGURE 16. Phylogenetic placement of Triozidus in family Triozidae is indicated by a black square representing a subgroup within Group B (see Percy et al. 2018), the related genera are expanded on in the text. An asterix indicates the placement of the type species of Heterotrioza, H. chenopodii in Group D, and Trioza, T. urticae in Group M, and Dyspersa sensu Cho et al. (2022) in Group A. Configuration of the three Triozidus taxa to one another based on the 714 bp COI fragment data (unique haplotypes only) is shown, with T. burckhardti placement indicated based on the backbone analysis (see text); taxon placements within Triozidus are uncertain in both analyses due to short internal branches and low to moderate bootstrap support, with a slightly different taxon configuration recovered in the ML backbone constraint analysis (see text). Illustrations of forewing, paramere and aedeagus are shown for all six Triozidus species to indicate comparative sizes; an asterisk by T. ceratophorus and T. eleutherococci indicates the illustrations are redrawn with reference to Li (2011) and Konovalova (1980, 1988) respectively, and in these cases paramere and aedeagus may not be to scale, plus the apex of the aedeagus is partly obscured in the illustration for T. eleutherococci. The annotation of the mitogenome of T. yangorum is graphically represented at top right.	FIGURE 16. Phylogenetic placement of Triozidus in family Triozidae is indicated by a black square representing a subgroup within Group B (see Percy et al. 2018), the related genera are expanded on in the text. An asterix indicates the placement of the type species of Heterotrioza, H. chenopodii in Group D, and Trioza, T. urticae in Group M, and Dyspersa sensu Cho et al. (2022) in Group A. Configuration of the three Triozidus taxa to one another based on the 714 bp COI fragment data (unique haplotypes only) is shown, with T. burckhardti placement indicated based on the backbone analysis (see text); taxon placements within Triozidus are uncertain in both analyses due to short internal branches and low to moderate bootstrap support, with a slightly different taxon configuration recovered in the ML backbone constraint analysis (see text). Illustrations of forewing, paramere and aedeagus are shown for all six Triozidus species to indicate comparative sizes; an asterisk by T. ceratophorus and T. eleutherococci indicates the illustrations are redrawn with reference to Li (2011) and Konovalova (1980, 1988) respectively, and in these cases paramere and aedeagus may not be to scale, plus the apex of the aedeagus is partly obscured in the illustration for T. eleutherococci. The annotation of the mitogenome of T. yangorum is graphically represented at top right.	2025-12-08	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.		Zenodo	biologists	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.			
03D2A373FFD0FFA6FF26F8C8FAD32C42.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17870925/files/figure.png	https://doi.org/10.5281/zenodo.17870925	FIGURE 4. Triozidus stackelbergi adults and galls on Eleutherococcus divaricatus. A, male; B, female; C, twig and male flower galls on E. divaricatus; D, detail of male flower galls.	FIGURE 4. Triozidus stackelbergi adults and galls on Eleutherococcus divaricatus. A, male; B, female; C, twig and male flower galls on E. divaricatus; D, detail of male flower galls.	2025-12-08	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.		Zenodo	biologists	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.			
03D2A373FFD0FFA6FF26F8C8FAD32C42.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17870929/files/figure.png	https://doi.org/10.5281/zenodo.17870929	FIGURE 5. Adult Triozidus stackelbergi comb. nov. A, head; B, antenna; C, forewing, with small dark spot at trifurcation of R+M+Cu (arrowed); D, hindwing; E, hind leg. Scale bars: 0.1 mm (A, B, E); 1 mm (C, D).	FIGURE 5. Adult Triozidus stackelbergi comb. nov. A, head; B, antenna; C, forewing, with small dark spot at trifurcation of R+M+Cu (arrowed); D, hindwing; E, hind leg. Scale bars: 0.1 mm (A, B, E); 1 mm (C, D).	2025-12-08	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.		Zenodo	biologists	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.			
03D2A373FFD0FFA6FF26F8C8FAD32C42.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17870935/files/figure.png	https://doi.org/10.5281/zenodo.17870935	FIGURE 6. Terminalia of Triozidus stackelbergi comb. nov. A, male terminalia (lateral view); B, paramere (inner surface); C, distal portion of aedeagus showing medial semicircular projections (arrowed); D, female terminalia (lateral view). Scale bars: 0.1 mm.	FIGURE 6. Terminalia of Triozidus stackelbergi comb. nov. A, male terminalia (lateral view); B, paramere (inner surface); C, distal portion of aedeagus showing medial semicircular projections (arrowed); D, female terminalia (lateral view). Scale bars: 0.1 mm.	2025-12-08	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.		Zenodo	biologists	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.			
03D2A373FFD0FFA6FF26F8C8FAD32C42.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17870937/files/figure.png	https://doi.org/10.5281/zenodo.17870937	FIGURE 7. Fifth instar immature of Triozidus stackelbergi comb. nov. A, habitus; B, tarsal arolium showing asymmetric development of tarsal claws (vestigial claw arrowed); C, forewing pad marginal sectasetae; D, circumanal ring showing regular single row of pores (inset); E, antenna. Scale bars: 0.1 mm.	FIGURE 7. Fifth instar immature of Triozidus stackelbergi comb. nov. A, habitus; B, tarsal arolium showing asymmetric development of tarsal claws (vestigial claw arrowed); C, forewing pad marginal sectasetae; D, circumanal ring showing regular single row of pores (inset); E, antenna. Scale bars: 0.1 mm.	2025-12-08	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.		Zenodo	biologists	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.			
03D2A373FFD0FFA6FF26F8C8FAD32C42.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17870967/files/figure.png	https://doi.org/10.5281/zenodo.17870967	FIGURE 16. Phylogenetic placement of Triozidus in family Triozidae is indicated by a black square representing a subgroup within Group B (see Percy et al. 2018), the related genera are expanded on in the text. An asterix indicates the placement of the type species of Heterotrioza, H. chenopodii in Group D, and Trioza, T. urticae in Group M, and Dyspersa sensu Cho et al. (2022) in Group A. Configuration of the three Triozidus taxa to one another based on the 714 bp COI fragment data (unique haplotypes only) is shown, with T. burckhardti placement indicated based on the backbone analysis (see text); taxon placements within Triozidus are uncertain in both analyses due to short internal branches and low to moderate bootstrap support, with a slightly different taxon configuration recovered in the ML backbone constraint analysis (see text). Illustrations of forewing, paramere and aedeagus are shown for all six Triozidus species to indicate comparative sizes; an asterisk by T. ceratophorus and T. eleutherococci indicates the illustrations are redrawn with reference to Li (2011) and Konovalova (1980, 1988) respectively, and in these cases paramere and aedeagus may not be to scale, plus the apex of the aedeagus is partly obscured in the illustration for T. eleutherococci. The annotation of the mitogenome of T. yangorum is graphically represented at top right.	FIGURE 16. Phylogenetic placement of Triozidus in family Triozidae is indicated by a black square representing a subgroup within Group B (see Percy et al. 2018), the related genera are expanded on in the text. An asterix indicates the placement of the type species of Heterotrioza, H. chenopodii in Group D, and Trioza, T. urticae in Group M, and Dyspersa sensu Cho et al. (2022) in Group A. Configuration of the three Triozidus taxa to one another based on the 714 bp COI fragment data (unique haplotypes only) is shown, with T. burckhardti placement indicated based on the backbone analysis (see text); taxon placements within Triozidus are uncertain in both analyses due to short internal branches and low to moderate bootstrap support, with a slightly different taxon configuration recovered in the ML backbone constraint analysis (see text). Illustrations of forewing, paramere and aedeagus are shown for all six Triozidus species to indicate comparative sizes; an asterisk by T. ceratophorus and T. eleutherococci indicates the illustrations are redrawn with reference to Li (2011) and Konovalova (1980, 1988) respectively, and in these cases paramere and aedeagus may not be to scale, plus the apex of the aedeagus is partly obscured in the illustration for T. eleutherococci. The annotation of the mitogenome of T. yangorum is graphically represented at top right.	2025-12-08	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.		Zenodo	biologists	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.			
03D2A373FFD5FFA4FF26FE09FE9F2AF9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17870939/files/figure.png	https://doi.org/10.5281/zenodo.17870939	FIGURE 8. Triozidus ukogi adults, immature and galls on Eleutherococcus spinosus. A, adults (male above), female (lateral and dorsal views, below) (images: Yorio Miyatake); B, immature within gall chamber (above) detail of immature (below); C, closed “spindle-shaped” gall on the petiole; D, petiolule gall. Scale bar in A 1 mm.	FIGURE 8. Triozidus ukogi adults, immature and galls on Eleutherococcus spinosus. A, adults (male above), female (lateral and dorsal views, below) (images: Yorio Miyatake); B, immature within gall chamber (above) detail of immature (below); C, closed “spindle-shaped” gall on the petiole; D, petiolule gall. Scale bar in A 1 mm.	2025-12-08	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.		Zenodo	biologists	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.			
03D2A373FFD5FFA4FF26FE09FE9F2AF9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17870945/files/figure.png	https://doi.org/10.5281/zenodo.17870945	FIGURE 9. Adult Triozidus ukogi comb. nov. A, head and antennae; B, forewing; C, hindwing; D, hind leg. Scale bars: 0.1 mm (A, D); 1 mm (B, C).	FIGURE 9. Adult Triozidus ukogi comb. nov. A, head and antennae; B, forewing; C, hindwing; D, hind leg. Scale bars: 0.1 mm (A, D); 1 mm (B, C).	2025-12-08	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.		Zenodo	biologists	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.			
03D2A373FFD5FFA4FF26FE09FE9F2AF9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17870947/files/figure.png	https://doi.org/10.5281/zenodo.17870947	FIGURE 10. Terminalia of Triozidus ukogi comb. nov. A, male terminalia (lateral view); B, paramere (inner surface); C, distal portion of aedeagus showing medial broadly semicircular projections (arrowed); D, female terminalia (lateral view). Scale bars: 0.1 mm.	FIGURE 10. Terminalia of Triozidus ukogi comb. nov. A, male terminalia (lateral view); B, paramere (inner surface); C, distal portion of aedeagus showing medial broadly semicircular projections (arrowed); D, female terminalia (lateral view). Scale bars: 0.1 mm.	2025-12-08	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.		Zenodo	biologists	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.			
03D2A373FFD5FFA4FF26FE09FE9F2AF9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17870951/files/figure.png	https://doi.org/10.5281/zenodo.17870951	FIGURE 11. Fifth instar immature of Triozidus ukogi comb. nov. A, habitus; B, tarsal arolium showing asymmetric development of tarsal claws (vestigial claw arrowed); C, forewing pad marginal simple setae; D circumanal ring showing single narrow row of elongate pores (inset); E antenna. Scale bars: 0.1 mm.	FIGURE 11. Fifth instar immature of Triozidus ukogi comb. nov. A, habitus; B, tarsal arolium showing asymmetric development of tarsal claws (vestigial claw arrowed); C, forewing pad marginal simple setae; D circumanal ring showing single narrow row of elongate pores (inset); E antenna. Scale bars: 0.1 mm.	2025-12-08	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.		Zenodo	biologists	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.			
03D2A373FFD5FFA4FF26FE09FE9F2AF9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17870967/files/figure.png	https://doi.org/10.5281/zenodo.17870967	FIGURE 16. Phylogenetic placement of Triozidus in family Triozidae is indicated by a black square representing a subgroup within Group B (see Percy et al. 2018), the related genera are expanded on in the text. An asterix indicates the placement of the type species of Heterotrioza, H. chenopodii in Group D, and Trioza, T. urticae in Group M, and Dyspersa sensu Cho et al. (2022) in Group A. Configuration of the three Triozidus taxa to one another based on the 714 bp COI fragment data (unique haplotypes only) is shown, with T. burckhardti placement indicated based on the backbone analysis (see text); taxon placements within Triozidus are uncertain in both analyses due to short internal branches and low to moderate bootstrap support, with a slightly different taxon configuration recovered in the ML backbone constraint analysis (see text). Illustrations of forewing, paramere and aedeagus are shown for all six Triozidus species to indicate comparative sizes; an asterisk by T. ceratophorus and T. eleutherococci indicates the illustrations are redrawn with reference to Li (2011) and Konovalova (1980, 1988) respectively, and in these cases paramere and aedeagus may not be to scale, plus the apex of the aedeagus is partly obscured in the illustration for T. eleutherococci. The annotation of the mitogenome of T. yangorum is graphically represented at top right.	FIGURE 16. Phylogenetic placement of Triozidus in family Triozidae is indicated by a black square representing a subgroup within Group B (see Percy et al. 2018), the related genera are expanded on in the text. An asterix indicates the placement of the type species of Heterotrioza, H. chenopodii in Group D, and Trioza, T. urticae in Group M, and Dyspersa sensu Cho et al. (2022) in Group A. Configuration of the three Triozidus taxa to one another based on the 714 bp COI fragment data (unique haplotypes only) is shown, with T. burckhardti placement indicated based on the backbone analysis (see text); taxon placements within Triozidus are uncertain in both analyses due to short internal branches and low to moderate bootstrap support, with a slightly different taxon configuration recovered in the ML backbone constraint analysis (see text). Illustrations of forewing, paramere and aedeagus are shown for all six Triozidus species to indicate comparative sizes; an asterisk by T. ceratophorus and T. eleutherococci indicates the illustrations are redrawn with reference to Li (2011) and Konovalova (1980, 1988) respectively, and in these cases paramere and aedeagus may not be to scale, plus the apex of the aedeagus is partly obscured in the illustration for T. eleutherococci. The annotation of the mitogenome of T. yangorum is graphically represented at top right.	2025-12-08	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.		Zenodo	biologists	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.			
03D2A373FFC8FFBFFF26F886FE9F2A85.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17870916/files/figure.png	https://doi.org/10.5281/zenodo.17870916	FIGURE 1. Triozidus burckhardti sp. nov. and Triozidus yangorum sp. nov. adults, immature and galls on the host plant, Eleutherococcus trifoliatus. A, T. burckhardti female; B, closed galls of cf. T. burckhardti with a single gall on each petiolule of the trifoliate leaf; C, T. yangorum male; D, T. yangorum fifth instar immature within gall chamber; E, close up showing three petiolule galls of T. yangorum on a single leaf, the upper already opened; F, the scandent host, E. trifoliatus, showing distribution of petiolule galls.	FIGURE 1. Triozidus burckhardti sp. nov. and Triozidus yangorum sp. nov. adults, immature and galls on the host plant, Eleutherococcus trifoliatus. A, T. burckhardti female; B, closed galls of cf. T. burckhardti with a single gall on each petiolule of the trifoliate leaf; C, T. yangorum male; D, T. yangorum fifth instar immature within gall chamber; E, close up showing three petiolule galls of T. yangorum on a single leaf, the upper already opened; F, the scandent host, E. trifoliatus, showing distribution of petiolule galls.	2025-12-08	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.		Zenodo	biologists	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.			
03D2A373FFC8FFBFFF26F886FE9F2A85.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17870959/files/figure.png	https://doi.org/10.5281/zenodo.17870959	FIGURE 12.Adults of Triozidus yangorum sp. nov. A, Holotype, male, dorsal view; B, Holotype, male, lateral view; C, female, dorsal view; D, female, lateral view. Scale bars: 1 mm.	FIGURE 12.Adults of Triozidus yangorum sp. nov. A, Holotype, male, dorsal view; B, Holotype, male, lateral view; C, female, dorsal view; D, female, lateral view. Scale bars: 1 mm.	2025-12-08	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.		Zenodo	biologists	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.			
03D2A373FFC8FFBFFF26F886FE9F2A85.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17870961/files/figure.png	https://doi.org/10.5281/zenodo.17870961	FIGURE 13. Adult Triozidus yangorum sp. nov. A, head; B, antenna; C, terminal antennal segments showing uneven length terminal setae (arrowed); D, forewing; E, hindwing; F, hind leg. Scale bars: 0.1 mm (A, B, C, F); 1 mm (D, E).	FIGURE 13. Adult Triozidus yangorum sp. nov. A, head; B, antenna; C, terminal antennal segments showing uneven length terminal setae (arrowed); D, forewing; E, hindwing; F, hind leg. Scale bars: 0.1 mm (A, B, C, F); 1 mm (D, E).	2025-12-08	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.		Zenodo	biologists	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.			
03D2A373FFC8FFBFFF26F886FE9F2A85.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17870963/files/figure.png	https://doi.org/10.5281/zenodo.17870963	FIGURE 14. Terminalia of Triozidus yangorum sp. nov. A, male terminalia (lateral view); B, paramere (inner surface); C, distal portion of aedeagus, showing medial narrow thumb-like projections (arrowed); D, female terminalia (lateral view). Scale bars: 0.1 mm.	FIGURE 14. Terminalia of Triozidus yangorum sp. nov. A, male terminalia (lateral view); B, paramere (inner surface); C, distal portion of aedeagus, showing medial narrow thumb-like projections (arrowed); D, female terminalia (lateral view). Scale bars: 0.1 mm.	2025-12-08	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.		Zenodo	biologists	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.			
03D2A373FFC8FFBFFF26F886FE9F2A85.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17870965/files/figure.png	https://doi.org/10.5281/zenodo.17870965	FIGURE 15. Fifth instar immature of Triozidus yangorum sp. nov. A, habitus; B, tarsal arolium showing asymmetric development of tarsal claws (vestigial claw arrowed); C, forewing pad marginal simple setae; D, circumanal ring showing intermittent irregular pore shape (inset); E, antenna. Scale bars: 0.1 mm.	FIGURE 15. Fifth instar immature of Triozidus yangorum sp. nov. A, habitus; B, tarsal arolium showing asymmetric development of tarsal claws (vestigial claw arrowed); C, forewing pad marginal simple setae; D, circumanal ring showing intermittent irregular pore shape (inset); E, antenna. Scale bars: 0.1 mm.	2025-12-08	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.		Zenodo	biologists	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.			
03D2A373FFC8FFBFFF26F886FE9F2A85.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17870967/files/figure.png	https://doi.org/10.5281/zenodo.17870967	FIGURE 16. Phylogenetic placement of Triozidus in family Triozidae is indicated by a black square representing a subgroup within Group B (see Percy et al. 2018), the related genera are expanded on in the text. An asterix indicates the placement of the type species of Heterotrioza, H. chenopodii in Group D, and Trioza, T. urticae in Group M, and Dyspersa sensu Cho et al. (2022) in Group A. Configuration of the three Triozidus taxa to one another based on the 714 bp COI fragment data (unique haplotypes only) is shown, with T. burckhardti placement indicated based on the backbone analysis (see text); taxon placements within Triozidus are uncertain in both analyses due to short internal branches and low to moderate bootstrap support, with a slightly different taxon configuration recovered in the ML backbone constraint analysis (see text). Illustrations of forewing, paramere and aedeagus are shown for all six Triozidus species to indicate comparative sizes; an asterisk by T. ceratophorus and T. eleutherococci indicates the illustrations are redrawn with reference to Li (2011) and Konovalova (1980, 1988) respectively, and in these cases paramere and aedeagus may not be to scale, plus the apex of the aedeagus is partly obscured in the illustration for T. eleutherococci. The annotation of the mitogenome of T. yangorum is graphically represented at top right.	FIGURE 16. Phylogenetic placement of Triozidus in family Triozidae is indicated by a black square representing a subgroup within Group B (see Percy et al. 2018), the related genera are expanded on in the text. An asterix indicates the placement of the type species of Heterotrioza, H. chenopodii in Group D, and Trioza, T. urticae in Group M, and Dyspersa sensu Cho et al. (2022) in Group A. Configuration of the three Triozidus taxa to one another based on the 714 bp COI fragment data (unique haplotypes only) is shown, with T. burckhardti placement indicated based on the backbone analysis (see text); taxon placements within Triozidus are uncertain in both analyses due to short internal branches and low to moderate bootstrap support, with a slightly different taxon configuration recovered in the ML backbone constraint analysis (see text). Illustrations of forewing, paramere and aedeagus are shown for all six Triozidus species to indicate comparative sizes; an asterisk by T. ceratophorus and T. eleutherococci indicates the illustrations are redrawn with reference to Li (2011) and Konovalova (1980, 1988) respectively, and in these cases paramere and aedeagus may not be to scale, plus the apex of the aedeagus is partly obscured in the illustration for T. eleutherococci. The annotation of the mitogenome of T. yangorum is graphically represented at top right.	2025-12-08	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.		Zenodo	biologists	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.			
