identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03D97C79011AFFD6F8EFFFAEFB847EAF.text	03D97C79011AFFD6F8EFFFAEFB847EAF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tibionemini Motyka, Kusy, Arias-Bohart, Bybee et Bocak 2025	<div><p>Tibionemini Motyka, Kusy, Arias-Bohart, Bybee et Bocak,  new tribe</p><p>http://zoobank.org/ urn:lsid:zoobank.org:act: BB8AA288-640C-4A66-A32F-B1BDEC480B14</p><p>Type genus.  Tibionema Solier, 1851 .</p><p>=  Tibionemini Ulrich, 1988 (in litt.), unavailable name.</p><p>Type genus.  Tibionema Solier, 1851 .</p><p>=  Tibionemini Motyka, Kusy, Arias, Bybee et Bocak, 2023 (Motyka et al. 2023b), unavailable name (Bouchard et al. 2024). urn:lsid:zoobank.org:pub: 91027F71-C0E4-4746-A440-DBD9C1894D6A.</p><p>Type genus.  Tibionema Solier, 1851 .</p><p>Diagnosis.  Tibionemini is proposed in Pityobiinae sensu Motyka et al. (2023b) as a monogeneric tribe. Currently, the nominotypical  Tibionema is the only genus placed in it, and its diagnosis presented below is simultaneously the diagnosis of the tribe.</p><p>Tibionema has a medium-to-large body, flat pronotum and elytra (Fig. 1A), distant longitudinal bulges in the pronotum, and acutely prominent posterior angles of the pronotum (Fig. 1A). The prosternum is narrow, and the prosternal process is long and slender (Fig. 1B). The lateral edges of the pronotum are complete (Fig. 1E, F). There is a sharp keel between mesocoxae (Fig. 1B). Antennae are 11-segmented and serrate; the scapus is parallelsided, antennomeres 2 and 3 are short and ball-like, and antennomeres 1–3 are almost bare and shining (Fig. 1D). The abdomen has five visible segments, a short intercoxal process, a straight posterior margin of the penultimate segment, and a triangular, apically rounded last visible abdominal segment. Internal abdominal segments consist of long, narrow penultimate, and small ultimate sternites (Fig. 1H).  Tibionema and  Pityobius differ in the number of antennomeres (twelve in  Pityobius versus eleven in  Tibionema); the posterior pronotal angles bent toward the body in  Tibionema (also in  Hapatesus and  Oxynopterus), but not in  Pityobius .  Tibionema has a prominent intercoxal keel in the mesosternum (Fig. 1B) that has not been observed in other Pityobiinae.  Campyloxenus differs in much longer antennomere 3. The larva and pupa of  T. abdominalis were described by Angulo (Angulo, 1970).</p><p>Remark. Here,  Tibionemini trib. nov. is proposed due to the unavailability of the same name published earlier by Motyka et al. (2023b). As we cannot get a response from the journal and rectify this omission in another way, we have to re-publish the description to make it valid in the sense of the Code (ICZN 2012).</p></div>	https://treatment.plazi.org/id/03D97C79011AFFD6F8EFFFAEFB847EAF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bocak, Ladislav;Motyka, Michal;Kusy, Dominik;Arias-Bohart, Elizabeth T.;Bybee, Seth M.;Kazantsev, Sergey V.;Yamamoto, Shuhei	Bocak, Ladislav, Motyka, Michal, Kusy, Dominik, Arias-Bohart, Elizabeth T., Bybee, Seth M., Kazantsev, Sergey V., Yamamoto, Shuhei (2025): Validation of earlier proposed taxa Tibionemini new tribe and Hiekeolycus winkleri new species. Zootaxa 5609 (2): 293-297, DOI: 10.11646/zootaxa.5609.2.10, URL: https://doi.org/10.11646/zootaxa.5609.2.10
03D97C790119FFD6F8EFFECFFE887995.text	03D97C790119FFD6F8EFFECFFE887995.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hiekeolycus Winkler 1987	<div><p>Hiekeolycus Winkler, 1987, nom. resurr.</p><p>Hiekeolycus Winkler, 1987: 66 .</p><p>Type species:  Hiekeolycus berendti Winkler, 1987 (original designation).</p><p>Diagnosis.  Hiekeolycus resembles related  Helcophorus Fairmaire in general appearance. The genera differ in the structure of the elytral humeri. The humeral bulge is built only by costa 3 in  Hiekeolycus (Fig. 2A,C) and by costae 3 and 4 in  Helcophorus .</p><p>Justification of the name resurrection.  Hiekeolycus Winkler, 1987 has been considered a synonym of  Helcophorus Fairmaire, 1881 (Kazantsev 2004, 2013) as these genera are similar, and the elytral humeri of  H. berendti and  H. ekaterinae were poorly seen (Kazantsev &amp; Perkovsky 2022; Winkler 1987). We compared all Eocene species with extant  Helcophorus and noted differently arranged costae. Therefore, we resurrect  Hiekeolycus as a separate genus.</p></div>	https://treatment.plazi.org/id/03D97C790119FFD6F8EFFECFFE887995	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bocak, Ladislav;Motyka, Michal;Kusy, Dominik;Arias-Bohart, Elizabeth T.;Bybee, Seth M.;Kazantsev, Sergey V.;Yamamoto, Shuhei	Bocak, Ladislav, Motyka, Michal, Kusy, Dominik, Arias-Bohart, Elizabeth T., Bybee, Seth M., Kazantsev, Sergey V., Yamamoto, Shuhei (2025): Validation of earlier proposed taxa Tibionemini new tribe and Hiekeolycus winkleri new species. Zootaxa 5609 (2): 293-297, DOI: 10.11646/zootaxa.5609.2.10, URL: https://doi.org/10.11646/zootaxa.5609.2.10
03D97C790119FFD7F8EFF8FEFD387E3B.text	03D97C790119FFD7F8EFF8FEFD387E3B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hiekeolycus winkleri Kazantsev et Yamamoto 2025	<div><p>Hiekeolycus winkleri Kazantsev et Yamamoto,  sp. nov.</p><p>http:/zoobank.org/ urn:lsid:zoobank.org:act: 812CA068-62DB-4EFC-B5AD-5B1DB5E15EE8</p><p>(Fig. 2A–E).</p><p>=  Hiekeolycus winkleri Kazantsev et Yamamoto in Motyka et al. 2023a, invalid name due to the absence of the secondary repository in journal’s ZooBank registration. Original registration in the ZooBank: http://zoobank.org/ urn:lsid:zoobank. org:act: D92B0B93-4077-46F7-8FB6-A7047C57EF85</p><p>Material examined.   Holotype: ♂, specimen No. AMNH Ba-SY2, deposited in the  American Museum of Natural History, New York, NY, USA. Baltic amber, Eocene.</p><p>Diagnosis.  H. winkleri sp. nov. differs from contemporary  H. berendti in the less conspicuous anterior and more acute posterior pronotal angles and distally widened hind trochanters (Fig. 2D, E). Furthermore,  H. berendti has denser and longer elytral vestiture and fewer elytral cells in each interstice than  H. winkleri (Fig. 2A–E).</p><p>Description. Male. Body elongated; dark brown (putatively red in the live specimen; Fig. 2). Head narrower than pronotum. Eyes prominent, interocular dorsal distance 1.4 times eye diameter (Fig. 2B); terminal palpomeres long, wide distally; antennae reaching over elytral mid-length; antennomere 3 longer than antennomere 2 (Fig. 2C), combined subequal in length to antennomere 4 (Fig. 2C). Pronotum 1.6 times wider than long, anteriorly rounded, with widely rounded anterior and sharply prominent posterior angles; median areole rhomboidal; lateral carinae almost straight; noticeable fronto-lateral carina ventrally (Fig. 2A). Elytra flat, elongated, 3.3 times longer than width at humeri, four primary costae, costa 3 stoutest and basally forming humeral bulge; interstices with transverse cells (Fig. 2A,B). Femurs widened distally, tibiae with pair of small spurs; frontal and middle trochanters elongate, distally rounded, hind trochanters distally widened (Fig. 2D); tarsomeres with plantar pads in tarsomeres 1–4 (Fig. 2A,B,D). Abdomen with eight visible ventrites, penultimate ventrite with broad triangular incision (Fig. 2E).</p><p>Measurements. Body length 11.2 mm; pronotal length 1.4 mm; pronotal width 2.2 mm.</p><p>Etymology.  H. winkleri sp. nov. is named in honor of our late colleague Josef R. Winkler (1930–1993), who described the first erotine taxa from Baltic amber in the late 1980s.</p><p>Locality and horizon. Baltic amber, Eocene.</p></div>	https://treatment.plazi.org/id/03D97C790119FFD7F8EFF8FEFD387E3B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bocak, Ladislav;Motyka, Michal;Kusy, Dominik;Arias-Bohart, Elizabeth T.;Bybee, Seth M.;Kazantsev, Sergey V.;Yamamoto, Shuhei	Bocak, Ladislav, Motyka, Michal, Kusy, Dominik, Arias-Bohart, Elizabeth T., Bybee, Seth M., Kazantsev, Sergey V., Yamamoto, Shuhei (2025): Validation of earlier proposed taxa Tibionemini new tribe and Hiekeolycus winkleri new species. Zootaxa 5609 (2): 293-297, DOI: 10.11646/zootaxa.5609.2.10, URL: https://doi.org/10.11646/zootaxa.5609.2.10
03D97C790118FFD7F8EFF805FEFA7A03.text	03D97C790118FFD7F8EFF805FEFA7A03.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hiekeolycus ekaterinae (Kazantsev et Perkovsky 2022) Bocak & Motyka & Kusy & Arias-Bohart & Bybee & Kazantsev & Yamamoto 2025	<div><p>Hiekeolycus ekaterinae (Kazantsev et Perkovsky, 2022),  comb. nov.</p><p>Helcophorus ekaterinae Kazantsev et Perkovsky, 2022: 86 .</p><p>Remark. We transfer  Helcophorus ekaterinae to  Hiekeolycus based on the updated genus diagnosis. The new combination with  Hiekeolycus proposed in 2023 is invalid due to incomplete data in the work’s registration in ZooBank.</p></div>	https://treatment.plazi.org/id/03D97C790118FFD7F8EFF805FEFA7A03	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bocak, Ladislav;Motyka, Michal;Kusy, Dominik;Arias-Bohart, Elizabeth T.;Bybee, Seth M.;Kazantsev, Sergey V.;Yamamoto, Shuhei	Bocak, Ladislav, Motyka, Michal, Kusy, Dominik, Arias-Bohart, Elizabeth T., Bybee, Seth M., Kazantsev, Sergey V., Yamamoto, Shuhei (2025): Validation of earlier proposed taxa Tibionemini new tribe and Hiekeolycus winkleri new species. Zootaxa 5609 (2): 293-297, DOI: 10.11646/zootaxa.5609.2.10, URL: https://doi.org/10.11646/zootaxa.5609.2.10
