taxonID	type	description	language	source
03E287812423505464C5FD3EAFE7F9A2.taxon	diagnosis	Diagnosis. Usually macropterous. Head longer than wide, with a pair of postocular setae. Antennae 8 - segmented, segment VIII rather long and weakly constricted at base, sense cone formula of segments III and IV variable, but usually (1 + 2) and (2 + 2 + 1) respectively, but rarely fewer. Maxillary stylets variable, rather close together and retracted to postocular setae or eyes, often V-shaped and not reaching postocular setae. Prothoracic notopleural suture usually complete, rarely incomplete; anteromarginal setae reduced; mesopresternum variable, almost complete or reduced medially. Metanotum with or without a pair of small setae before median pair of setae; metathoracic sternopleural suture absent; both metafurcal arms usually fused medially (cf. Fig. 253). Fore wings weakly constricted medially, with duplicated cilia. Fore femur with an inner sub-basal tubercle and with several small tubercles before sub-basal tubercle; fore tibia with a sub-apical scale or tubercle; fore tarsal tooth arising at apex of inner margin and directed forwardly.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781242C505764C5F935A89FFDC4.taxon	description	(Figs 6, 11 – 19) Female (macroptera). Distended body length: 1.5 – 2.0 mm. Body brown to dark brown (Fig. 6), tube the darkest. Fore femora largely yellow, shaded with brown postero-externally, but sometimes more brown or largely yellow, mid and hind femora brown; tibiae and tarsi yellow. Antennal segments I – II and VII – VIII pale brown; segments III – VI largely yellow, segment VI scarcely shaded distally. Fore wings and major body setae pale. Head longer than wide (Fig. 11), 1.35 – 1.40 times as long as wide, 1.37 times in holotype; dorsal surface almost smooth, sculptured laterally, but with a few transverse fine wrinkles or striae at middle. Cheeks weakly rounded, serrated. Postocular setae shorter than eyes, weakly expanded, 110 µm apart from each other, 17 – 18 µm apart from eyes in holotype. Eyes largely developed, 0.36 – 0.38 times as long as head, ommatidia scarcely spaced. Ocelli normal, posterior pair in contact with eyes, 30 µm apart from each other in holotype. Antennae (Fig. 19) about 1.7 times as long as head; segment VIII very weakly constricted at base, about 0.9 times as long as segment VII; segments III a little longer than IV; segments III and IV each with (1 + 2) and (2 + 2 + 1) sense cones respectively, these sense cones are pale and slender, often very difficult to observe. Maxillary stylets rather short and wider apart, not reaching postocular setae, almost V-shaped; maxillary bridge wide, slightly wider than half of head width, placed just before basal collar of head, 7 – 10 µm apart from basal collar of head at middle in holotype. Pronotum (Fig. 11) 0.69 times as long as head, 1.46 times as wide as long in holotype, almost smooth; only three pairs of major prothoracic setae weakly expanded; ml reduced to small setae, but often somewhat longer; pa and epim well developed, pa almost as long as postocular setae, or longer, epim longer than pa. Prosternal basantra and ferna well-developed (Fig. 13). Prospinasternum well-developed, widely fan-shaped, 35 µm wide in holotype. Mesopresternum transverse, nearly complete. Mesonotal lateral setae not reduced. Metanotum sculptured with polygonal reticulation, with no small setae before median pair of setae; median pair of setae short and acute, 25 µm apart from each other, 52 – 55 µm apart from anterior margin of metanotum in holotype. Fore femora typical of the genus (Figs 15 & 16), with an inner sub-basal tubercle and an irregular row of several minute tubercles; fore tibial inner apical scale small (Fig. 17). Fore wing with 2 – 7 duplicated cilia, with 6 cilia in holotype; sub-basal setae S 1 and S 2 expanded or weakly expanded, S 2 longer than S 1, S 3 much longer than S 2, blunt or very weakly expanded. Pelta (Fig. 14) trapezoid or hat-shaped, usually with indistinct lateral wings, 1.05 times as wide as long in holotype. Tergites I – VIII S 1 setae weakly expanded. Tergites III – VIII S 2 setae: III – VI weakly expanded or blunt, VII pointed, VIII expanded. Tergite IX S 1 setae blunt, shorter than tube, S 2 finely pointed, longer than tube; intermediate setae about half of S 1. Tube (Fig. 18) about 0.6 times as long as head, 0.62 times in holotype, 2.0 – 2.2 times as long as wide; terminal setae about 1.5 times as long as tube, or a little longer. Measurements (holotype female in µm). Body length 1950 (distended). Head length 210, from anterior margin of eyes 193, width across eyes 152, maximum width across cheeks 153, minimum width across base 127; eyes length 79; diameter of posterior ocelli 16 – 17; postocular setae 50 – 55. Maxillary bridge width 81. Antenna total length 355, segments III – VIII length (width) as follows: 52 (28), 49 (31), 45 (26), 42 (22), 35 (18), 32 (11). Pronotum length 145, width 212. Setae on prothorax: aa 32, ml minute, pa 58 – 60, epim 70 – 73. Fore wing length 800. Sub-basal wing setae: S 1 40 – 41, S 2 48 – 50, S 3 78 – 86. Pelta length 78, width 82. Tergite IX setae: S 1 101 – 107, S 2 140 – 144. Tube length 130, maximum width 60; terminal setae 190 – 200. Male (macroptera). Distended body length: 1.4 – 1.8 mm. Very similar to macropterous female in color and structure. Head (Fig. 12) 1.2 – 1.3 times as long as wide; maxillary bridge wider, 0.55 – 0.60 times as wide as head; fore wing with 4 – 6 duplicated cilia; tergite IX S 1 setae longer than 0.4 times as long as S 2; tube 0.64 – 0.66 times as long as head; terminal tube setae 1.3 – 1.4 times as long as tube. Measurements (paratype male in µm). Body length 1520 (distended). Head length 167, from anterior margin of eyes 155, width across eyes 136, maximum width across cheeks 135, minimum width across base 110; eyes length 72; diameter of posterior ocelli 12 – 15; postocular setae about 40. Maxillary bridge width 77. Antenna total length 290, segments III – VIII length as follows: 40, 39, 37, 37, 32, 29. Pronotum length 114, width 172. Setae on prothorax: aa 18, ml less than 10, pa 52 – 55, epim 52 – 52. Fore wing length 620. Sub-basal wing setae: S 1 36 – 38, S 2 40 – 43, S 3 65 – 67. Pelta length 70, width 74. Tergite IX setae: S 1 75 – 80, S 2 35 – 38. Tube length 110, maximum width 25; terminal setae 145 – 150.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781242C505764C5F935A89FFDC4.taxon	materials_examined	Type material. Holotype: macropterous female, Thailand, nr. Chiang Mai, Doi Suthep, on bamboo, 1. ix. 1991, TN & SO. Paratypes: Thailand, 15 females and 1 male, collected together with holotype; 1 male, nr. Chiang Mai, Mae-sa, on bamboo, 3. ix. 1991, TN & SO; 1 female, nr. Chiang Mai, Hong Kai, 4. ix. 1991, TN & SO; 4 females, nr. Chiang Mai, Phrow, on bamboo, 22. viii. 1992, TN & SO; 3 females, nr. Chiang Mai, Ka Jan, on bamboo, 3. ix. 1992, TN & SO; 3 females, Chiang Mai Prov., Mae Sa Nam, on bamboo, 20. iii. 1999, P. Jangvitaya; 14 females and 11 males, Phuket Is., Phuket Hill, on bamboo, 22. viii. 1991, TN & SO; 2 females and 2 males, data very similar to above, but 9. ix. 1992; 2 females and 1 male, Phuket Is., nr. Rawai Beach, on bamboo, 15. ix. 1992, TN & SO. Non-paratypic specimens. Thailand, 1 female and 1 male, Bangkok, Bangkhen, Campus of Kasetsart University, NBCRC, on bamboo, 22. viii. 1990, TN & SO; 3 females, nr. Kanchanabri, on bamboo, 30. viii. 1991, TN & SO. Vietnam, 8 females and 1 male, Son La Province, Moc Chau, on bamboo, 5. iii. 2000, SO; 1 female and 1 male, Hoa Binh Province, Hoa Binh, on bamboo, 6. iii. 2000, SO; 1 female, Hatay Province, Mt. Bavi, on bamboo, 29. vii. 2000, SO; 4 females and 1 male, Bac Can Province, Ba Be National Park, on bamboo, 2. viii. 2000; SO. Indonesia, Java, 1 female, Malang, Batu, Pujon, on bamboo, 22. viii. 2005, SO; 1 female, Malang, Sundeng, Sawo Sajar, 24. viii. 2005, SO; 1 female, Banyuwangi, Malan Sari, on bamboo, 27. viii. 2005, SO.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781242C505764C5F935A89FFDC4.taxon	discussion	Comments. This species is described from Thailand based on a good number of specimens collected from bamboo, and included in the coimbatorensis - group. It is related to A. coimbatorensis that is also taken from bamboo in India, and these two species share the maxillary stylets short and V-shaped even in female and the prothoracic midlateral setae reduced. However, from coimbatorensis, bambusicola can be distinguished by the following features: mid and hind tibiae yellow; antennal segments IV – VI largely yellow; prothoracic anteroangular setae developed, expanded; fore tibial inner sub-apical scale small; tergite IX S 2 setae longer than tube. A. huanglianensis from Yunnan, China, is also similar to this species, but it has the pronotal midlateral setae well-developed and the fore femora largely brown. Several females and males listed under the non-paratypic specimens may represent this species, but have some differences in the length of setae, such as prothoracic anteroangular setae, sub-basal wing setae and S 1 and S 2 setae on the abdominal tergite IX. In particular, three females from ‘ nr. Kanchanabri’, Thailand, have the prothoracic notopleural sutures incomplete.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781242F505064C5FDADA8BCF837.taxon	description	(Figs 7, 20 – 27) Female (macroptera). Distended body length: 1.95 – 2.20 mm. Body brown (Fig. 7). Femora brown, extreme apices of fore femora yellow; fore tibiae yellow, mid and hind tibiae brown (cf. Fig. 25), with extreme bases and apices slightly paler; tarsi yellow. Antennal segments I – II and VII – VIII brown, somewhat paler than thorax, segments III – VI yellow, often segment VI scarcely shaded distally. Fore wings and major body setae pale. Head much longer than wide (Fig. 20), 1.43 times as long as wide in holotype; dorsal surface almost smooth, sculptured laterally. Cheeks sub-parallel, very weakly rounded, weakly serrated. Postocular setae shorter than eyes, weakly expanded, 111 µm apart from each other, 17 µm apart from eyes in holotype. Eyes a little longer than one-third of head length, 0.35 times as long as head in holotype. Ocelli well-developed; posterior pair slightly separated from eyes, 29 µm apart from each other in holotype. Antennae (Fig. 26) about 1.7 times as long as head; segment VIII rather slender, very weakly constricted at base, a little shorter than 0.8 times as long as segment VII; segments III and IV subequal in length; segments III and IV each with (1 + 2) and (2 + 2 + 1) sense cones respectively, these sense cones are short and slender, very difficult to observe. Maxillary stylets reaching eyes, rather close together at middle, about one-third of head width apart; maxillary bridge rather weak, situated at 32 µm from basal collar of head in holotype. Pronotum (Fig. 20) 0.58 times as long as head, 1.74 times as wide as long in holotype, almost smooth; four pairs of major prothoracic setae well-developed, expanded; aa and ml subequal in length; pa almost as long as postocular setae; epim the longest. Prosternal basantra and ferna well-developed (Fig. 23). Prospinasternum about 30 µm wide in holotype. Mesopresternum narrowly boat-shaped. Mesonotal lateral setae well-developed, weakly expanded, about 30 µm long in holotype. Metanotum sculptured with polygonal reticulation, with a pair of small setae before median pair of setae; median pair of setae 28 – 31 µm in length, 53 µm apart from each other, about 52 – 58 µm apart from anterior margin of metanotum in holotype. Fore femora (Fig. 22) typical of the genus, but not so enlarged, with a few small tubercles at inner margin before sub-basal tubercle; fore tibial inner apical scale slender. Fore wing with 6 – 7 duplicated cilia, with 6 cilia in holotype; sub-basal setae S 1 and S 2 expanded, S 2 longer than S 1, S 3 much longer than S 2, weakly expanded. Pelta trapezoid (Fig. 24), 1.23 times as wide as long in holotype. Tergites I – VIII S 1 setae very weakly expanded, but often blunt. Tergites III – VIII S 2 setae: III – V short and blunt or very weakly expanded, VI – VIII pointed or nearly pointed. Tergite IX S 1 and S 2 setae pointed, S 1 almost as long as tube, or a little shorter; S 2 a little longer than tube; intermediate setae shorter than half of S 1. Tube (Fig. 27) shorter than 0.6 times as long as head, 0.55 times in holotype, about 1.8 times as long as wide; terminal setae about 1.4 times as long as tube. Measurements (holotype female in µm). Body length about 2200 (distended). Head length 223, from anterior margin of eyes 205, width across eyes 154, maximum width across cheeks 156, minimum width across base 133; eyes length 79; diameter of posterior ocelli 17 – 20; postocular setae 63 – 64. Maxillary bridge width 53. Antenna total length 385, segments III – VIII length (width) as follows: 54 (31), 58 (32), 51 (26), 46 (22), 42 (20), 32 (12). Pronotum length 130, width 226. Setae on prothorax: aa 46 – 47, ml about 50, pa 63 – 64, epim 75. Fore wing length 850. Sub-basal wing setae: S 1 45 – 46, S 2 56 – 58, S 3 95. Pelta length 91, width 112. Tergite IX setae: S 1 119 – 120, S 2 124 – 137. Tube length 122, maximum width 68; terminal setae 170. Male (macroptera). Distended body length: about 1.5 mm. Very similar to macropterous female in color and structure. Head shorter (Fig. 21), shorter than 1.2 times as long as wide; maxillary stylets relatively wider apart, almost V-shaped; maxillary bridge a little narrower than half of head width; fore wing with 5 – 6 duplicated cilia; tergite IX S 2 setae 0.28 – 0.33 times as long as S 1; tube about 0.6 times as long as head. Measurements (paratype male in µm). Body length 1540 (distended). Head length 155, from anterior margin of eyes 143, width across eyes 135, maximum width across cheeks 138, minimum width across base 113; eyes length 57; diameter of posterior ocelli 12 – 14; postocular setae about 40. Maxillary bridge width 63. Antenna total length 310, segments III – VIII length as follows: 44, 43, 40, 37, 33, 30. Pronotum length 101, width 178. Setae on prothorax: aa 30, ml 30, pa 45 – 48, epim 52 – 54. Fore wing length 660. Sub-basal wing setae: S 1 35, S 2 42 – 44, S 3 66 – 68. Pelta length 66, width 82. Tergite IX setae: S 1 90 – 93, S 2 26 – 30. Tube length 95, maximum width 53; terminal setae 143 – 148.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781242F505064C5FDADA8BCF837.taxon	materials_examined	Type material. Holotype: macropterous female, Vietnam, Lam Dong province, Bao Loc, Dam Bri, on bamboo, 26. xii. 2001, SO. Paratypes: Vietnam, 2 females and 2 males, collected together with holotype.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781242F505064C5FDADA8BCF837.taxon	discussion	Comments. This species is described from Vietnam based on three females and two males collected from bamboo, and was obtained together with Vietothrips bicolor gen. et sp. nov. described below. It is somewhat similar to A. fuscipes sp. nov. from Sulawesi, Indonesia, in having brownish mid and hind tibiae, but can be distinguished by the following features: ocelli well-developed; maxillary stylets reaching eyes; antennal segment IV with a small outer sense cone in addition to four major sense cones, (2 + 2 + 1); tergite IX S 1 and S 2 setae shorter. A. crus from Taiwan and A. ramachandrai from somewhat wider range of Asia are also somewhat similar to this species, but have antennal segments IV to VI with apices brownish.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812429505264C5FF65AAA6F967.taxon	description	(Figs 8, 28 – 36) Female (macroptera). Distended body length: 2.0 – 2.3 mm. Body brown to dark brown (Fig. 8). Femora brown, fore femora with apices slightly paler; fore tibiae yellow, mid and hind tibiae pale brown (cf. Fig. 35), at least not clear yellow, with paler apices; tarsi yellow. Antennal segments I – II brown, but somewhat paler than head, segment II slightly paler antero-externally; segments III – V yellow; segment VI shaded with brown at anterior half, yellowish at basal half; segments VII – VIII brown. Fore wings very slightly shaded. Major body setae pale. Head much longer than wide (Fig. 28), 1.37 times as long as wide in holotype; dorsal surface almost smooth, sculptured laterally, but with a few transverse fine wrinkles at middle. Cheeks sub-parallel, very weakly rounded, weakly serrated. Postocular setae shorter than eyes, weakly expanded, 120 µm apart from each other, about 15 µm apart from eyes in holotype. Eyes about one-third as long as head, or a little longer, 0.35 times as long as head in holotype. Ocelli relatively small; posterior pair slightly separated from eyes, about 30 µm apart from each other in holotype. Antennae (Fig. 36) about 1.7 times as long as head; segment VIII very weakly constricted at base, a little longer than 0.8 times as long as segment VII; segment III a little shorter than segment IV; segments III and IV each with (1 + 2) and (2 + 2) sense cones respectively, these sense cones are short and slender, very difficult to observe. Maxillary stylets retracted to postocular setae, rather close together at middle, a little wider than one-third of head width; maxillary bridge rather distinct, situated at about 40 µm from basal collar of head. Pronotum 0.62 times as long as head, 1.57 times as wide as long in holotype, almost smooth; four pairs of major prothoracic setae well-developed, expanded; pa almost as long as postocular setae, or a little longer; epim the longest. Prosternal basantra and ferna well-developed (Fig. 29). Prospinasternum usually longer than wide, 25 µm wide in holotype. Mesopresternum narrowly boat-shaped (Fig. 30) or divided (Fig. 29). Mesonotal lateral setae well-developed, weakly expanded, 30 µm long in holotype. Metanotum sculptured with polygonal reticulation (Fig. 31), with a pair of small setae before median pair of setae; median pair of setae relatively stout, pointed, 36 – 37 µm in length, 39 µm apart from each other, about 70 µm apart from anterior margin of metanotum in holotype. Fore femora typical of the genus (Fig. 34), but not so enlarged, with a few small tubercles at inner margin before sub-basal tubercle; fore tibial inner apical scale slender (Fig. 33). Fore wing with 6 – 8 duplicated cilia, with 6 – 7 in holotype; sub-basal setae S 1 and S 2 expanded, S 2 longer than S 1, S 3 much longer than S 2, pointed or bluntly pointed. Pelta trapezoid or nearly triangular (Fig. 32), 1.16 times as wide as long in holotype. Tergites II – VIII S 1 setae very weakly expanded. Tergites III – VIII S 2 setae: III – V short and blunt or very weakly expanded, VI – VIII pointed or nearly pointed. Tergite IX S 1 and S 2 setae pointed, longer than tube, S 2 longer than S 1; intermediate setae short, shorter than one-third of S 1. Tube shorter than 0.6 times as long as head, 0.56 times in holotype, about 1.9 times as long as wide; terminal setae about 1.5 times as long as tube, or a little longer. Measurements (holotype female in µm). Body length about 2200 (distended). Head length 228, from anterior margin of eyes 210, width across eyes 164, maximum width across cheeks 167, minimum width across base 143; eyes length 79; diameter of posterior ocelli 13 – 15; postocular setae 60 – 65. Maxillary bridge width 65. Antenna total length about 380, segments III – VIII length (width) as follows: 57 (30), 59 (32), 49 (28), 45 (26), 40 (22), 33 (13). Pronotum length 142, width 223. Setae on prothorax: aa 55, ml 50, pa 67 – 68, epim 70 – 80. Fore wing length about 900. Sub-basal wing setae: S 1 50 – 52, S 2 67 – 70, S 3 110 – 112. Pelta length 93, width 108. Tergite IX setae: S 1 136, S 2 158 – 160. Tube length 128, maximum width 69; terminal setae?. Male (macroptera). Distended body length: 1.65 – 1.90 mm. Very similar to macropterous female in color and structure. Head 1.30 – 1.35 times as long as wide; maxillary bridge somewhat wider, 0.43 – 0.46 times as wide as head; fore wing with 5 – 7 duplicated cilia; tergite IX S 2 setae shorter than one-fourth of S 1; tube longer than 0.6 times as long as head. Measurements (paratype male in µm). Body length about 1800 (distended). Head length 197, from anterior margin of eyes 182, width across eyes 146, maximum width across cheeks 148, minimum width across base 129; eyes length 73; diameter of posterior ocelli 12 – 14; postocular setae 57 – 60. Maxillary bridge width 66. Antenna total length 370, segments III – VIII length as follows: 54, 57, 45, 44, 40, 32. Pronotum length 118, width 210. Setae on prothorax: aa 45 – 47, ml about 50, pa 53 – 58, epim 66 – 68. Fore wing length 760. Sub-basal wing setae: S 1 45 – 46, S 2 57 – 58, S 3 87. Pelta length 78, width 84. Tergite IX setae: S 1 113 – 115, S 2 23 – 25. Tube length 127, maximum width 53; terminal setae 163.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812429505264C5FF65AAA6F967.taxon	materials_examined	Type material. Holotype: macropterous female, Indonesia, Sulawesi, 11 km E from Malino, Kanreapia, alt. about 1500 m, on grass, 2. viii. 1984, SO. Paratypes: Indonesia, Sulawesi, 4 females and 6 males, collected together with holotype.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812429505264C5FF65AAA6F967.taxon	discussion	Comments. This species is described from Sulawesi, Indonesia, based on five females and six males taken from grasses, and has the mid and hind tibiae brown. Amongst previously described 12 species of the genus, only two species, A. crus and A. ramachandrai, have the mid and hind tibiae brown, at least not yellow, and the other ten species have the tibiae clear yellow. Although crus and ramachandrai have antennal segment IV with one outer small sense cone in addition to the four major sense cones, (2 + 2 + 1), fuscipes has only four major sense cones, (2 + 2).	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781242A504D64C5F8F9AB9EFEC0.taxon	description	(Figs 9, 37 – 44, 253) Female (macroptera). Distended body length: 1.8 – 2.4 mm. Body bicolored (Fig. 9). Head brownish yellow, paler than abdomen, paler at anterior one-third; pterothorax and abdomen including tube brown, extreme base of tube yellowish; prothorax and legs yellow. Antennal segments I and II pale brown, segments VII and VIII brown; segments III – V largely yellow, but segments IV and V slightly shaded at anterior half, segment VI brown at distal half, yellowish at basal half. Fore wings and major body setae pale. Head much longer than wide (Fig. 37), 1.40 – 1.55 times as long as wide, 1.49 times in holotype; dorsal surface almost smooth, sculptured laterally, but with a few transverse fine wrinkles at middle. Cheeks sub-parallel, very weakly rounded, serrated. Postocular setae shorter than eyes, weakly expanded, 108 µm apart from each other, about 15 µm apart from eyes in holotype. Eyes about one-third as long as head. Ocelli normal, posterior pair 25 µm apart from each other in holotype. Antennae (Fig. 44) about 1.8 times as long as head; segment VIII rather slender, very weakly constricted at base, about 0.8 times as long as segment VII; segments III and IV subequal in length, each with (1 + 1) and (1 + 2) sense cones respectively, these sense cones are pale and slender, very difficult to observe. Maxillary stylets retracted to postocular setae, rather close together at middle; maxillary bridge rather distinct, situated far from basal collar of head, much narrower than half of head width. Pronotum (Fig. 37) 0.59 times as long as head, 1.57 times as wide as long in holotype, almost smooth; four pairs of major prothoracic setae weakly expanded; aa and ml subequal in length, much shorter than pa; pa almost as long as postocular setae; epim the longest. Prosternal basantra and ferna well-developed (Fig. 38). Prospinasternum circular or irregularly fan-shaped, 26 µm wide in holotype. Mesopresternum narrowly boat-shaped or scarcely divided. Mesonotal lateral setae well-developed, expanded. Metanotum sculptured with polygonal reticulation (Fig. 39), with a pair of small setae before median pair of setae; median pair of setae short and acute, 36 µm apart from each other, 57 – 65 µm apart from anterior margin of metanotum in holotype. Fore femora not so enlarged (Fig. 42), sub-basal inner tubercle rather small (Fig. 43), often reduced to small hump in small individuals; fore tibial inner apical scale small (Fig. 41), but it is not visible depending on the angle of the specimen. Fore wing with 3 – 8 duplicated cilia, with 5 – 6 in holotype; sub-basal setae S 1 and S 2 expanded or weakly expanded, S 2 longer than S 1, S 3 much longer than S 2, blunt or very weakly expanded, rarely bluntly pointed. Pelta trapezoid (Fig. 40), 1.13 times as wide as long in holotype. Tergites II – VIII S 1 setae very weakly expanded, but those on VII often blunt or bluntly pointed. Tergites III – VIII S 2 setae: III – V short and blunt, VI – VIII pointed or nearly pointed, but those on VII often blunt or very weakly expanded. Tergite IX S 1 and S 2 setae subequal in length, finely pointed, longer than tube; intermediate setae short, about one-fourth of S 1. Tube about 0.6 times as long as head, about 2.0 times as long as wide; terminal setae about 1.2 times as long as tube, or a little longer. Measurements (holotype female in µm). Body length 1940 (distended). Head length 217, from anterior margin of eyes 196, width across eyes 143, maximum width across cheeks 146, minimum width across base 125; eyes length 73; diameter of posterior ocelli 13 – 15; postocular setae 54. Maxillary bridge width 62. Antenna total length 387, segments III – VIII length (width) as follows: 54 (29), 57 (30), 52 (27), 45 (23), 43 (20), 36 (13). Pronotum length 127, width 200. Setae on prothorax: aa about 30, ml 35, pa about 50, epim 70 – 75. Fore wing length 700. Sub-basal wing setae: S 1 33 – 35, S 2 42, S 3 78 – 80. Pelta length 85, width 98. Tergite IX setae: S 1 168, S 2 165 – 167. Tube length 126, maximum width 117; terminal setae 160. Measurements (paratype large female from Thailand in µm). Body length 2430 (distended). Head length 230, from anterior margin of eyes 208, width across eyes 145, maximum width across cheeks 150, minimum width across base 132; eyes length 75; diameter of posterior ocelli 15 – 17; postocular setae 58 – 62. Maxillary bridge width 52. Antenna total length 415, segments III – VIII length as follows: 60, 60, 55, 50, 45, 37. Pronotum length 148, width 226. Setae on prothorax: aa about 40, ml 35 – 38, pa 55 – 60, epim 75. Fore wing length 850. Sub-basal wing setae: S 1 40 – 43, S 2 55 – 57, S 3 95 – 100. Pelta length 97, width 110. Tergite IX setae: S 1 165 – 172, S 2 165 – 172. Tube length 138, maximum width 66; terminal setae 165 – 170. Male (macroptera). Distended body length: 1.5 – 1.8 mm. Very similar to macropterous female in color and structure. Head 1.35 – 1.45 times as long as wide; maxillary stylets somewhat wider apart; tergite IX S 2 setae shorter than one-fourth of S 1; tube a little shorter than 0.6 times as long as head. Measurements (paratype male in µm). Body length 1770 (distended). Head length 192, from anterior margin of eyes 172, width across eyes 133, maximum width across cheeks 135, minimum width across base 115; eyes length 68; diameter of posterior ocelli 12 – 13; postocular setae 40 – 45. Maxillary bridge width 65. Antenna total length 345, segments III – VIII length as follows: 50, 50, 47, 41, 40, 32. Pronotum length 115, width 183. Setae on prothorax: aa 25, ml 25, pa 35, epim 55. Fore wing length 680. Sub-basal wing setae: S 1 32 – 33, S 2 35, S 3 70 – 72. Pelta length 75, width 80. Tergite IX setae: S 1 130 – 140, S 2 27 – 28. Tube length 108, maximum width 56; terminal setae 135 – 138.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781242A504D64C5F8F9AB9EFEC0.taxon	materials_examined	Type material. Holotype: macropterous female. Japan, the Ryukyus, Okinawa-hontou Is., Tomigusuku-shi, Tomigusuku-joshi park, on Poaceae, 2. v. 2015, K. Minoura. Paratypes: Japan, the Ryukyus, same locality and habitat as in holotype, 6 females, 12. vi. 2002, M. Masumoto, 2 females, 21. ii. 2015, 2 females and 1 male, 28. ii. 2015, K. Minoura; 1 male, Ishigaki-jima Is., Mt. Banna-dake, on grass, 1. ix. 1988, SO; 1 male, Ishigaki-jima Is., Mt. Omotodake, on grass, 2. ix. 1988, SO. Taiwan, 1 female and 1 male, Pingtung Hsien, Kenting Nat. Park, on grass, 22. v. 1972, SO; same locality and habitat as above, 1 female and 1 male, 18. iii. 1984, 2 females and 2 males, 19. iii. 1984, SO; Chiai Hsien, Kuantzulin, 1 female, on grass, 1 female, on dead leaves, 1. iv. 1993, and branches, TN & SO; 1 female, Lan-Yu, 4 – 6. vi. 1980, H. Makihara. Thailand, Chiang Mai, paddy field, 3 females, 4. v. 1978, 1 female and 4 males, 24. vi. 1979, K. Yasumatsu; Kamphaeng Saen Campus of Kastsart University, on grass, 2 females, 29. xii. 1987, 1 female, 12. i. 1988, SO; 1 female and 2 males, Sarabri, Farm of Kasetsart University, on grass, 20. viii. 1991, TN & SO; 1 female and 2 males, nr. Chiang Mai, Doi Pui, on grass, 1. ix. 1992, SO. Vietnam, 1 male, Ha Noi, Farm of Ha Noi Agricultural University, 29. vii. 2000, SO; 1 male, Lam Dong Province, Bao Loc, Dam Bri, on bamboo, 27. xii. 2001, SO; 1 male, same locality as above, on grass, 22. viii. 2007, SO. Borneo, 1 female, Sabah, Jl. Kimanis, 15 km from Kimanis, on dead leaves and branches, 9. ix. 1990, TN & SO.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781242A504D64C5F8F9AB9EFEC0.taxon	discussion	Comments. This species is described from a somewhat wider range of eastern Asia based on a good number of females and males collected from grasses, and may be included in the melastomae - group. It is peculiar within the genus in having (1 + 1) and (1 + 2) sense cones on antennal segments III and IV respectively, because most congeners have (1 + 2) and (2 + 2 + 1) or (2 + 2) sense cones. However, it has the fore femur armed with an inner sub-basal tubercle (Fig. 42) and the fore tibia with an inner sub-apical scale (Fig. 41) in both sexes; these are important characteristics of the genus Androthrips. The interspecific variation in the number of sense cones within the genus is not unique because it is also observed in other genera included in the Haplothripini, such as Karnyothrips, Neandrothrips and Mirothrips (Cavalleri et al. 2016; Okajima & Masumoto 2025 a, 2025 b). A. graminicola is apparently very similar to Karnyothrips species especially in small individuals, because the fore femoral sub-basal tubercle and the fore tibial sub-apical scale are scarcely developed. However, unlike Karnyothrips, the metanotum has a pair of anterior small setae in addition to median pair of setae (Fig. 39) and the prosternal basantra are more closely located to the ferna (Fig. 38) in this species.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812435504F64C5FD23A94AFE54.taxon	description	(Figs 10, 45 – 52) Female (macroptera). Distended body length: 2.05 – 2.45 mm. Body bicolored (Fig 10). Head, thorax, abdominal segment I and legs yellow, often head very weakly shaded with brown; abdominal segments II – VIII bicolored, shaded with brown medially, yellow marginally, brown area darkened and enlarged in posterior segments, segment IX largely yellow, scarcely shaded with brown; tube brown, with basal one-fifth yellow. Antennae largely yellow, segments VII – VIII very slightly shaded. Fore wings and major body setae pale. Head much longer than wide (Fig. 45), 1.43 times as long as wide in holotype; dorsal surface almost smooth, sculptured laterally, but with a few transverse fine wrinkles at middle. Cheeks weakly rounded, weakly serrated. Postocular setae shorter than eyes, blunt or very weakly expanded, 137 µm apart from each other, about 20 µm apart from eyes in holotype. Eyes well-developed, a little longer than one-third of head length. Ocelli normal, posterior pair in contact with eyes, 38 µm apart from each other in holotype. Antennae (Fig. 52) about 1.7 times as long as head; segment VIII rather slender, very weakly constricted at base, more than 0.9 times as long as segment VII; segment III asymmetrical, almost as long as segment IV; segments III and IV each with (1 + 2) and (2 + 2 + 1) sense cones respectively. Maxillary stylets not reaching postocular setae, rather wider apart, V-shaped; maxillary bridge rather distinct, about 25 µm apart from basal collar of head at middle in holotype, about half of head width. Pronotum (Fig. 45) well-developed, 0.83 times as long as head, 1.26 times as wide as long in holotype, almost smooth; four pairs of major prothoracic setae developed; aa and ml blunt or very weakly expanded, ml a little longer than aa; pa and epim weakly expanded, longer than postocular setae. Prosternal basantra and ferna well-developed (Fig. 46). Prospinasternum rather small, longer than wide, 25 µm wide in holotype. Mesopresternum almost complete. Mesonotal lateral setae well-developed, weakly expanded. Metanotum distinctly sculptured with polygonal reticulation (Fig. 47), with no small setae before median pair of setae; median pair of setae short and acute, 55 µm apart from each other, 75 – 78 µm apart from anterior margin of metanotum in holotype. Fore femora enlarged (Fig. 50), sub-basal inner tubercle distinct, with two rows of small but distinct tubercles at inner margin before sub-basal tubercle; fore tibial inner apical scale distinct (Fig. 49). Fore wing with 5 – 7 duplicated cilia, with 7 cilia in holotype; sub-basal setae S 1 and S 2 weakly expanded, S 2 longer than S 1, S 3 much longer than S 2, blunt or very weakly expanded. Pelta trapezoid (Fig. 48), 1.24 times as wide as long in holotype. Tergites II – VIII S 1 and S 2 setae weakly expanded, but S 2 on tergite II short and pointed. Tergite IX S 1 and S 2 setae blunt or very weakly expanded, shorter than tube, S 2 longer than S 1; intermediate setae short, much shorter than half of S 1. Tube (Fig. 51) a little longer than 0.6 times as long as head, 0.63 times in holotype, a little longer than 2.0 times as long as wide, 2.25 times in holotype; terminal setae about 1.5 times as long as tube. Measurements (holotype female in µm). Body length 2440 (distended). Head length 258, from anterior margin of eyes 235, width across eyes 179, maximum width across cheeks 180, minimum width across base 151; eyes length 93; diameter of posterior ocelli 18 – 21; postocular setae 65. Maxillary bridge width 89. Antenna total length 440, segments III – VIII length (width) as follows: 67 (32), 68 (33), 55 (27), 52 (23), 44 (20), 42 (13). Pronotum length 215, width 270. Setae on prothorax: aa? 35, ml about 50, pa 81, epim 80 – 84. Fore wing length 980. Sub-basal wing setae: S 1 50, S 2 65, S 3 104. Pelta length 95, width 118. Tergite IX setae: S 1 110 – 120, S 2 125. Tube length 162, maximum width 72; terminal setae 245. Male (macroptera). Distended body length: 2.0 – 2.1 mm. Very similar to macropterous female in structure. Abdomen largely brownish, segments VI – IX almost entirely brown; tube entirely dark brown. Head a little longer than 1.45 times as long as wide; antenna shorter than 1.7 times as long as head; pronotum about 1.7 times as long as head, about 1.3 times as long as wide; tergite IX S 2 setae about half of S 1, or a little longer; tube slightly shorter than 0.6 times as long as head; terminal tube setae about 1.6 times as long as tube. Measurements (paratype male in µm). Body length about 2000 (distended). Head length 240, from anterior margin of eyes 220, width across eyes 158, maximum width across cheeks 163, minimum width across base 134; eyes length 70; diameter of posterior ocelli 16 – 21; postocular setae 65 – 70. Maxillary bridge width 78. Antenna total length 390, segments III – VIII length as follows: 59, 57, 50, 47, 42, 39. Pronotum length 167, width 218. Setae on prothorax: aa? 30, ml? 35, pa 75 – 78, epim 70. Fore wing length 810. Sub-basal wing setae: S 1 47 – 48, S 2 53 – 55, S 3 100 – 105. Pelta length 77, width 100. Tergite IX setae: S 1 102 – 105, S 2 55. Tube length 137, maximum width 65; terminal setae 220.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812435504F64C5FD23A94AFE54.taxon	materials_examined	Type material. Holotype: macropterous female, Indonesia, Java, Malang, Sundeng, Sawo Sajar, 450 m alt., on bamboo, 24. viii. 2005, SO. Paratypes: Indonesia, Java, 2 females and 2 males, collected together with holotype.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812435504F64C5FD23A94AFE54.taxon	discussion	Comments. This species is described from Java, Indonesia, based on three females and two males taken from bamboo, and probably belongs in the coimbatorensis group. It is somewhat similar to A. bambusicola sp. nov. described above from Thailand, A. coimbatorensis from India and A. huanglianensis from China also collected from bamboo in having short and V-shaped maxillary stylets and a well-developed mesopresternum that is nearly complete. However, although bambusicola, coimbatorensis and huanglianensis have uniformly brown bodies, javae has the body bicolored and largely yellowish. A. collaris which was collected from tobacco leaf in Sumatra, Indonesia, also has a bicolored body and its color pattern is somewhat similar to this species according to the original description (Karny 1925). However, javae can be distinguished from collaris by the followings: head entirely yellow (brownish yellow with dark sides in collaris); antennal segments VII and VIII yellow, weakly shaded with brown (dark brown in collaris); pterothorax yellow (grayish brown in collaris).	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812437504864C5FE28A893F9A8.taxon	description	Okajima (1993 b) and Dang et al. (2014) suggested all eight species classified into both of these two genera inhabit bamboo. However, Veerabahuthrips species live under the leaf sheath of bamboo and feed on scale insects, at least in bambusae species-group (the two species-groups in Veerabahuthrips, the bambusae - group and the exilis - group, are recognized below as two different genera). In contrast, Mychiothrips species live on the leaf or stem of bamboo and probably prey on small arthropods other than scale insects; there is even a slight possibility that these species are phytophagous. At least Mychiothrips fruticola, the type-species of the genus known from Japan, has never been found under the sheath of bamboo, and it has not been confirmed to be predaceous, despite our extensive observations. In general, the mid and hind legs of Veerabahuthrips are somewhat short and thick relative to their body size (Figs 57 – 61), while those of Mychiothrips are rather long and slender (Figs 53 – 56). These morphological differences between Mychiothrips and Veerabahuthrips seem important because they are associated with different behaviors. Presumably, the short mid and hind legs of Veerabahuthrips are adapted to living in closed, narrow spaces between the leaf sheath and stem of bamboo, and since they prey on immobile scale insects they do not need to run fast. Podothrips and Okajimathrips also have similar short mid and hind legs (cf. Figs 94 & 108) and prey on scale insects in the narrow space under the sheath of bamboo or grass. In contrast, the long legs among Mychiothrips are possibly adapted to running fast to prey on small mobile arthropods and / or to escape from natural enemies that may be common in open spaces. These long mid and hind legs of Mychiothrips do not show size related variation, and the legs are also elongate even in small individuals. In contrast to our observation, Ng observed in Veerabahuthrips simplex, of the exilis - group (see below), that the sharp fore femoral teeth are used to grip a leaf vein in windy conditions (Ng & Mound 2015), but it is uncertain whether this observation was in its natural state or experimental. In our observations, Veerabahuthrips species rarely roamed freely on the leaves of bamboo. However, it is also true that Ng’s observation has an insight that cannot be ignored. Observing the fore legs of Veerabahuthrips species, it seems that when they are folded, the largest femoral tooth and the large tarsal tooth always fit well (Figs 64 – 67), as if they are adapted to gripping something. The possibility that Veerabahuthrips species walks on bamboo leaves in rare occasions cannot be ruled out, but it is not clear because there are few observations. Otherwise, there is a slight possibility that, unlike the bambusae - group, the exilis - group may also live on bamboo leaves rather than under the leaf sheaths. However, unlike Mychiothrips which live on the leaves, the mid and hind legs of the exilis - group are short as already mentioned. In any case, further observations will be necessary to understand the behavior and life history of these thrips and to make a more accurate classification.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812430504964C5F934A93DFEC0.taxon	description	(Figs 53, 75)	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812430504964C5F934A93DFEC0.taxon	description	Specimens examined. Thailand, holotype female and most of paratypes (data described in Okajima, 1993); additionally, some females and males (detailed data omitted).	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812431504B64C5FE9CAEBAFE54.taxon	description	(Figs 3, 54 – 56, 62 – 63, 68, 76)	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781243C504464C5FF65AE21FBE8.taxon	description	Type-species. Namothrips bambusae sp. nov.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781243C504464C5FF65AE21FBE8.taxon	discussion	Comments. The genus Namothrips presumably belongs to the Phlaeothrips - lineage, and it may be nearly allied to the Oriental genus Ablemothrips Ananthakrishnan. These two genera share the head long and elevated dorsally, the maxillary stylets elongated and close together medially, the fore wings weakly constricted medially and without duplicated cilia, and the fore tarsal tooth absent in both sexes. Ablemothrips has the prothoracic notopleural suture complete, the prosternum without basantra, the antennal segments VII and VIII closely and widely fused, and sternite VIII of male with no pore plate (Okajima 1999). However, Namothrips has the notopleural sutures incomplete, the prosternal basantra weakly developed, antennal segments VII and VIII well separated, and has a pair of pore plates on sternite VIII of males. Moreover, Ablemothrips has the postocular cephalic setae close together in female but far apart from each other in male, whereas they are close together in both sexes in Namothrips. The genus name is derived from the type locality ‘ Viêt Nam’. Gender: masculine.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781243C504664C5FB75AF02FD58.taxon	description	(Figs 83 – 93) Female (macroptera). Distended body length: about 1.5 mm. Body sharply bicolored brown and yellow (Fig. 83). Head and pterothorax brown, but head is yellow on ventral surface (Fig. 85); abdominal segment IX and tube pale brown, somewhat paler than head; abdominal segments I – VIII yellow, but segments I and VI – VIII very weakly shaded with brown. Legs yellow, but mid coxae are brown. Antennal segments I – II and VII – VIII brown, segments III – VI yellow. Fore wings weakly shaded with brown. Prominent body setae pale. Head 1.9 times as long as wide (Fig. 84), distinctly elevated dorsally (Fig. 85), widest across cheeks just behind eyes; dorsal surface covered with fine wrinkles and sculptured with weak reticulation (Fig. 86). Cheeks almost straight, weakly narrowed towards base, each with a short postocular cheek seta. Postocular setae much shorter than eyes, weakly expanded, about 40 µm apart from each other in holotype. Eyes about one-fourth as long as head. Posterior ocelli 10 – 12 µm in diameter, about 10 µm apart from each other in holotype. Antennae (Fig. 92) 1.35 times as long as head in holotype; segment VI a little longer than segment III. Pronotum (Fig. 84) 0.42 times as long as head and 1.55 times as wide as long in holotype; sculptured with reticulation marginally, semicircular lines sub-medially. Four pairs of pronotal setae and epimeral setae well developed, expanded at apices. Mesonotum distinctly sculptured; mesopresternum somewhat wider, not narrowed or eroded medially. Metanotum sculptured medially with longitudinal reticulation or lines (Fig. 87), with fine dots among lines, smooth laterally; median pair of setae short, 25 – 27 µm apart from anterior margin, 22 µm apart from each other in holotype. Three sub-basal setae expanded, S 2 a little shorter than S 1. Pelta weak (Fig. 89), narrowly hat-shaped, with no campaniform sensilla. Wing-retaining setae well developed and sigmoid (Fig. 90), but those on tergite II are short and rather straight. Tergite IX S 1 and S 2 setae pointed. Tube (Fig. 93) 0.43 times as long as head, about 2.0 times as long as wide in holotype. Measurements (holotype female in μm). Body length about 1500 (distended). Head length 208, from anterior margin of eyes 195, width across cheeks 110, across eyes 108; eyes length 50, width 38 – 40; postocular setae 35 – 38. Antenna total length 280, segments I – VIII length (width) as follows: 27 (25), 38 (24), 37 (21), 35 (22), 38 (20), 40 (20), 32 (18), 24 (10). Pronotum length 87, width 135. Setae on prothorax: am 40 – 42, aa 45 – 48, ml 37 – 42, pa 50, epim 53 – 55, cox 20. Fore wing length 580. Sub-basal wing setae: S 1 45, S 2 38 – 43, S 3 50 – 52. Tergite IX setae: S 1 40 – 42, S 2 60 – 70. Tube length 90, maximum width 46; terminal setae about 90. Male (macroptera). Distended body length: 1.1 – 1.2 mm. Color and structure very similar to female. Pore plate on abdominal sternite VIII, weak, separated both sides (Fig. 91); tergite IX S 2 setae shorter, almost as long as S 1 or shorter. Tube shorter than 0.4 times as long as head. Measurements (paratype male in μm). Body length about 1120 (distended). Head length 183, from anterior margin of eyes 165, width 95; eyes length 40, width 35; postocular setae about 20. Antenna total length 260, segments I – VIII length (width) as follows: 27 (22), 33 (21), 32 (17), 30 (19), 33 (20), 37 (18), 30 (17), 20 (10). Pronotum length 65, width 100. Setae on prothorax: am 20, aa 25, ml 25, pa 30, epim 40, cox 15. Fore wing length 490. Sub-basal wing setae: S 1 26, S 2 23, S 3 30. Tergite IX setae: S 1 about 30, S 2 about 30. Tube length 70, maximum width 35; terminal setae 70.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781243C504664C5FB75AF02FD58.taxon	materials_examined	Type material. Holotype: macropterous female, Vietnam, Lam Dong Province, Bao Loc, Dam Bri, on bamboo, 27. xii. 2001, S. Okajima. Paratypes: Vietnam, 5 females and 1 male, collected with holotype; 6 females and 2 males, data very similar to holotype, but 28. xii. 2001.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781243C504664C5FB75AF02FD58.taxon	discussion	Comments. This species is described from Vietnam based on 12 females and 3 males taken from bamboo. It was collected together with Vietothrips bicolor gen. et sp. nov. described below, but their feeding habit is unknown.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781243E504664C5FD24AFE7F8A0.taxon	diagnosis	Diagnosis. Body slightly flattened, macropterous. Head longer than wide, with a pair of postocular setae. Postocular setae arose near cheeks. Eyes and ocelli well-developed. Antennae eight-segmented; segments III and IV each with (1 + 1) sense cones. Mouth cone short and rounded, relatively pointed; maxillary stylets retracted to postocular setae, close together medially; maxillary bridge present. Pronotum with three pairs of setae developed, am and ml reduced; notopleural sutures incomplete. Basantra well developed, longer than wide, often fused to ferna. Mesopresternum divided into two lateral triangles; metathoracic sternopleural sutures well-developed, furcal arms separated from one another, without spinula. Fore tarsal tooth present in both sexes (Fig. 99), falciform; fore tibia with an inner subapical tubercle. Fore wings parallel-sided, without duplicated cilia. Pelta wide, nearly semicircular. Tergites II – VII with two pairs of wing-retaining setae. Tube shorter than head, distinctly constricted at apex; terminal setae a little longer than tube. Male tergite IX extruded posteriorly. Male sternite VIII with a pair of pore plates.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781243F504064C5FF65AB22FF24.taxon	description	(Figs 94 – 104, 254) Female (macroptera). Distended body length: 2.5 – 2.7 mm. Body largely yellow (Fig. 94); anterior half of head and apex of tube brown, but rarely abdominal segment IX and tube brown; the rest of body and legs yellow. Antennal segment I brown, segments II – VI yellow, segments VII – VIII scarcely shaded with brown. Fore wings scarcely shaded, major body setae pale. Head about 1.3 times as long as wide (Fig. 96), 1.33 times in holotype, scarcely protruded in front of eyes, widest across cheeks at a level of postocular setae, dorsal surface almost smooth. Cheeks rounded, almost smooth. Postocular setae shorter than eyes, expanded, situated near cheeks, 175 µm apart from each other, 32 – 33 µm apart from eyes in holotype. Eyes relatively small, about one-fourth of head length, ommatidia separated from each other. Ocelli small, posterior ocelli almost as large as ommatidium, about 80 µm apart from each other. Antennae (Fig. 104) about 1.6 times as long as head; segment VIII pedicellate, distinctly constricted at base, a little longer than segment VII; segment III 1.85 times as long as wide in holotype. Maxillary stylets 14 µm apart from each other in holotype; maxillary bridge somewhat weak, a little narrower than one-fifth of head width. Pronotum 0.94 times as long as head and 1.03 times as wide as long in holotype, almost smooth. Prosternum and mesopresternum very similar to those of the type-species (Fig. 101). Mesonotum almost smooth, lateral pair of setae short and acute. Metanotum almost smooth at middle; median pair of setae short, 72 – 73 µm apart from anterior margin, 65 µm apart from each other in holotype. Fore wing sub-basal setae short and pointed. Pelta weak. Tergites II – VIII S 2 setae: II short, but weakly expanded, III, IV and VIII expanded, V – VII finely pointed. Tergite IX S 1 and S 2 setae shorter than tube, pointed, S 1 much shorter than S 2. Tube (Fig. 102) 0.77 times as long as head, 2.47 times as long as wide in holotype. Measurements (holotype female in µm). Body length 2670 (distended). Head length 248, from anterior margin of eyes 220, width across eyes 170, maximum width across cheeks 187; eyes length 63; diameter of posterior ocelli 8 – 10; postocular setae 35 – 40. Maxillary bridge width 35. Antenna total length 395, segments III – VIII length (width) as follows: 50 (27), 55 (30), 50 (27), 50 (25), 43 (20), 45 (15). Pronotum length 233, width 240. Setae on prothorax: aa about 20, pa 25, epim 52 – 53. Fore wing length 850. Sub-basal wing setae: S 1 17, S 2 15, S 3 17. Pelta length? 80, width 132. Tergite IX setae: S 1 45 – 50, S 2 150 – 155. Tube length 190, maximum width 77; terminal setae about 200. Male (macroptera). Distended body length: 2.0 – 2.2 mm. Very similar to female in color and structure. Head a little shorter than 1.3 times as long as head; tube about 0.7 times as long as head. Pore plates on sternite VIII are very similar to those of the type-species (Fig. 103). Measurements (paratype male in µm). Body length 2140 (distended). Head length 208, from anterior margin of eyes 184, width across eyes 150, maximum width across cheeks 162; diameter of posterior ocelli 7 – 9; postocular setae about 35. Maxillary bridge width 30. Antenna total length 360, segments III – VIII length as follows: 47, 48, 43, 43, 37, 45. Pronotum length 200, width 202. Setae on prothorax: aa 20 – 22, pa 20 – 25, epim 45 – 47. Fore wing length 740. Sub-basal wing setae: S 1 11 – 15, S 2 15 – 20, S 3 15 – 16. Pelta length 78, width 103. Tergite IX setae: S 1 43 – 45, S 2 40 – 45. Tube length 150, maximum width 61; terminal setae 190.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781243F504064C5FF65AB22FF24.taxon	materials_examined	Type material. Holotype: macropterous female, Thailand, nr. Chiang Mai, Doi Suthep, on bamboo, 1. ix. 1991, TN & SO. Paratypes: Thailand, 31 females and 11 males, collected together with holotype; same locality and habitat as in holotype, 4 females, 3. ix. 1991, 27 females and 10 males, 24. viii. 1992, TN & SO, 1 female, 2. ix. 1992, SO; 4 females, nr. Chiang Mai, Ka Jan, on bamboo, 3. ix. 1992, SO; 2 females, nr. Chiang Mai, Hong kai, on bamboo, 4. ix. 1991, TN & SO; 2 females, Chiang Mai, on bamboo, 2. ix. 1991, TN & SO; 2 females, nr. Chiang Mai, Doi Inthanon, on bamboo, 29. viii. 1992, TN & SO. Non-paratypic specimens. Thailand, 4 females and 1 male, nr. Chiang Mai, Ka Jan, on bamboo, 3. ix. 1992, SO.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781243F504064C5FF65AB22FF24.taxon	discussion	Comments. This species has the body largely yellow (Fig. 94), although O. kentingensis has the body sharply bicolored brown and yellow (Fig. 105). Previously, this species has been considered as a color form of kentingensis, but it appears to have several differences in structure, and it can be distinguished from kentingensis by the key above. Therefore, caution should be exercised because this species is mixed in the specimens collected from Thailand that Okajima has identified as ‘ O. kentingensis ’ in the past. Incidentally, Fig. 133 in Dang et al. (2014) is likely to be this species, not kentingensis, based on the aspect ratio of the head and the condition of the maxillary stylets. Four females and one male listed in the non-paratypic specimens may represent this species, but these have the head largely pale brown (Fig. 97), and abdominal segment IX and tube largely brown (Fig. 95). These specimens were collected together with four females that show the typical color pattern of borealis, and the two color forms are indistinguishable in structure. This species and O. kentingensis are ectopically distributed, but the boundary is not clear at this time.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812438504164C5F8F7AE2BFE0C.taxon	description	(Figs 105 – 107)	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812439504264C5FDD0AFE7F840.taxon	description	Sixteen species are recognized here from East Asia, of which six species are newly described. Among them, only two species, P. ferrugineus and P. lucasseni, are uniformly brown, but the remaining 14 species are sharply bicolored brown and yellow. All species are predators and inhabit bamboos and / or grasses, especially bamboos infested by scale insects. Curiously, despite repeated investigations conducted by Okajima, no species other than a widely distributed P. lucasseni have been observed from Java and Bali, Indonesia. Diagnosis. Body slightly flattened, usually macropterous. Head a little longer than wide, almost smooth dorsally, often with a pair of lateral teeth at cheeks just behind eyes; cheeks without stout setae; a pair of postocular setae developed; postocellar setae minute. Ocelli developed but small, anterior ocellus situated at inter-antennal projection. Antennae 8 - segmented, segment III with (1 + 1) sense cones, IV with (1 + 1) or (1 + 1 + 1) sense cones. Mouth cone short and rounded, maxillary stylets at least usually reaching at postocular setae and rather close together, maxillary bridge present. Prothorax with three or four pairs of setae developed, am setae reduced, ml setae often reduced; notopleural suture complete. Basantra well developed (cf. Fig. 112), usually longer than wide, ferna and prospinasternum developed, mesopresternum boat shaped. Metathoracic sternopleural suture present, but absent in some Australian species; furcal arms usually separated from one another (cf. Fig. 251), without spinula. Fore tibia frequently with an inner subapical tooth, fore tarsus with well-developed apical tooth in both sexes. Fore wing constricted at middle (cf. Fig. 255), with or without duplicated cilia. Pelta usually bell-shaped, often semicircular, usually with a pair of campaniform sensilla. Abdominal tergites II – VII each with two pairs of wing-retaining setae, second pair on VII usually reduced to straight setae in female; tube shorter than head, anal setae longer than tube. Male abdominal tergite IX with S 2 setae short, sternite VIII without pore plate.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781243B507C64C5FF65AB01FE54.taxon	description	(Figs 108 – 115) Female (macroptera). Distended body length 2.2 – 2.6 mm. Body bicolored (Fig. 108). Head and abdominal segments VII – X brown, thorax yellow with pleural areas of metathorax shaded, abdominal segments I – VI yellow with an anteromedian brown marking on each tergites III – VI. Antennal segment I brown, II brown with apex yellowish, III – VI yellow with distal half of V – VI shaded, VII – VIII brown; all legs yellow. Fore wings pale, slightly shaded at middle; major body setae pale. Head about 1.3 times as long as wide (Fig. 109), widest just behind compound eyes, without lateral tooth. Cheeks weakly rounded and weakly narrowed towards base, often with an indistinct small tooth just behind eyes, slightly constricted near base. Postocular setae short and brunt at apex, 156 µm apart from each other in holotype. Posterior ocelli 65 µm apart from each other in holotype. Antennal segment III with (1 + 1) sense cones, segment IV and V each with (1 + 1 + 1) sense cones; antennal segment VIII slightly constricted at base. Ratio length / width of antennal segments III – VIII (Fig. 115) in holotype as followings: 2.1, 2.1, 2.4, 2.4, 2.8, 2.9. Maxillary stylet reaching near postocular setae, their distance about 0.3 times as wide as width across cheeks. Pronotum (Fig. 109) a little shorter than wide, about 0.95 times as long as wide in holotype, almost smooth and weakly sculptured along posterior margin and with a longitudinal line at middle. Prothoracic setae short, aa and epim expanded, ml blunt, pa pointed, epim the longest (right one short and pointed in holotype). Prosternum and mesopresternum typical of the genus (Fig. 112). Metanotum almost smooth medially; median pair of setae 43 µm apart from each other, 84 – 87 µm apart from anterior margin. Fore femora unarmed; fore tibial inner apical tooth distinct (Fig. 111). Fore wings with duplicated cilia, rarely only one cilium; three sub-basal setae short, S 1 setae expanded and the longest, S 2 setae pointed, S 3 setae pointed and longer than S 2. Pelta hut-shaped (Fig. 113). Abdominal tergite VIII with S 1 setae bluntly expanded; tergite IX S 1 and S 2 setae pointed, S 1 about 0.7 times (missing at distal parts of both S 1 in holotype) and S 2 about 1.2 times as long as tube. Tube (Fig. 114) distinctly narrowed at basal fifth, almost straight-sided, gently constricted at apex, 0.56 times as long as head in holotype, about 2.2 times as long as basal width; terminal setae 1.7 – 1.8 times as long as tube. Measurements (holotype female in µm). Body length about 2370 (distended). Head length 250, from anterior margin of eyes 225, width across eyes 188, maximum width across cheeks 193; eye length 75; diameter of posterior ocelli 15; postocular setae 19 – 23. Maxillary bridge width 60. Antenna total length 408, segments III – VIII length (width) as followings: 58 (28), 58 (28), 60 (25), 55 (23), 55 (20), 38 (13). Pronotum length 213, width 230. Setae on prothorax: aa 14 – 15, ml 8 – 12, pa 19 – 23, epim 20 – 43. Fore wing length 850. Sub-basal wing setae: S 1 15 – 20, S 2 10 – 11, S 3 15. Pelta length 90, width 113. Tergite IX setae: S 1?, S 2 163 – 168. Tube length 140, basal width 65; terminal setae 243 – 250. Male (macroptera). Distended body length 1.6 – 2.1 mm. Body color and structures similar to female but smaller, abdominal tergite VII slightly pale at middle. Maxillary stylet not reaching postocular setae (Fig. 110), their distance wider than that of female, about 0.3 times as wide as head. Measurements (paratype male in µm). Body length about 2050 (distended). Head length 215, from anterior margin of eyes 198, width across eyes 173, maximum width across cheeks 178; eye length 75; diameter of posterior ocelli 13; postocular setae 22 – 26. Maxillary bridge width 40. Antenna total length 389, segments III – VIII length (width) as followings: 55 (25), 58 (28), 58 (25), 53 (23), 51 (19), 38 (11). Pronotum length 190, width 213. Setae on prothorax: aa 15 – 16, ml 8 – 10, pa 18, epim 38 – 43. Fore wing length 780. Sub-basal wing setae: S 1 18 – 19, S 2 15, S 3 15 – 18. Pelta length 83, width 105. Tergite IX setae: S 1 66 – 74, S 2 43. Tube length 128, basal width 53; terminal setae 218 – 235.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781243B507C64C5FF65AB01FE54.taxon	materials_examined	Type material. Holotype: female, Borneo, East Malaysia, Sabah, nr. Keningaw, Taman Bunsit, on bamboo, 2. ix. 1991, TN & SO. Paratypes: 1 female and 4 males collected together with holotype. Vietnam, 1 female, Tuyen Quang Province, Ham Yan Distr., Tan Lap, Lam Truong, on bamboo, 19. viii. 2000, SO; 3 females, Nghe An Prov., Aanh Son District, Cao Son, on bamboo, 9. viii. 2007, SO; 1 female, Thai Nguyen Province, on bamboo, 22. viii. 2000; 1 female, Hoa Binh Province, Hoa Binh, on bamboo, 6. iii. 2000, SO; 3 females, Bac Can Province, Ba Be National Park, on bamboo, 2. viii. 2000, SO; Lam Dong Province, Bao Loc, Dam Bri, on bamboo, 1 female, 26. xii. 2001. 2 females, 27. xii. 2001, 10 females and 2 males, 28. xii. 2001, SO.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781243B507C64C5FF65AB01FE54.taxon	discussion	Comments. This species is very similar to P. flavipes sp. nov. described below from Thailand in body color. It has abdominal segment VII brown and the fore wings with duplicated cilia, whereas flavipes has abdominal segment VII yellow with an anteromedian brown marking and the fore wings without duplicated cilia. The specimens from Vietnam have S 1 and S 2 setae on abdominal tergite IX somewhat shorter. P. semiflavus is also somewhat similar to this species but can easily be distinguished by having abdominal segment VII brown, antennal segment IV with (1 + 1 + 1) sense cones, and the fore wing with duplicated cilia. The body color of this species is very similar to that of P. canizoi from India (Bhatti 1978). However, canizoi has a pair of distinct teeth on the cheeks behind the compound eyes, and is not so closely related to this species. It is probably more closely related to P. erami from Iran than to all bicolored Podothrips species distributed in East Asia (Minaei 2015).	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812404507E64C5F9DEAF6FFD2C.taxon	description	(Figs 116 – 124) Female (macroptera). Distended body length 1.9 – 2.2 mm. Body bicolored (Fig. 116). Head, thorax and tube brown, pleural areas of metathorax and tube darker but tube slightly yellow at base and apex; abdominal segments I – VII yellow; tergites III – VII each with an anteromedian brown marking, but paler on VII; tergite VI and VII often shaded at posterior angles; segment VIII yellow with posterior third brown, segment IX brown with anterior half yellow. Antennal segments I – VI brownish yellow but I and II slightly darker, V – VI with apices brown, VII – VII brown. All legs yellow. Fore wings pale slightly shaded at middle; major body setae pale. Head 1.1 – 1.2 times as long as wide (Fig. 118), 1.15 times in holotype, widest just behind compound eyes, without lateral tooth. Cheeks rounded and weakly narrowed towards base, slightly constricted near base. Postocular setae short and pointed, 145 µm apart from each other in holotype. Posterior ocelli relatively small, 66 µm apart from each other in holotype. Antennal segment III with (0 + 1) sense cone, IV and V each with (1 + 1 + 1) sense cones; segment VIII not narrowed at base. Ratio length / width of antennal segments III – VIII (Fig. 124) in holotype as followings: 2.0, 1.7, 2.1, 2.1, 2.5, 2.0. Maxillary stylet reaching postocular setae, their distance about 0.2 times as wide as width across cheeks. Pronotum (Fig. 118) almost as long as wide, almost smooth and weakly sculptured along posterior margin and with a longitudinal line at middle; prothoracic setae relatively short, aa, ml and epim expanded at apex, pa pointed, epim the longest. Prosternum and mesopresternum typical of the genus (Fig. 120). Metanotum almost smooth anteromedially; median pair of setae 58 µm apart from each other, 78 – 80 µm apart from anterior margin in holotype. Fore femora unarmed; fore tibial inner apical tooth distinct (Fig. 121). Fore wings without duplicated cilia; three sub-basal setae short, S 1 and S 3 subequal in length, pointed, S 2 expanded and the longest, but S 1 rarely expanded at almost as long as S 2 setae. Pelta hut-shaped (Fig. 122). Abdominal tergite VIII S 1 setae expanded at apex; tergite IX S 1 and S 2 setae pointed, S 1 about 0.6 times as long as tube, S 2 almost as long as tube. Tube (Fig. 123) distinctly narrowed at basal fifth, gently tapered towards apex, constricted at apex, about 0.6 times as long as head, about 2.2 times as long as basal width; terminal setae 1.4 – 1.5 times as long as tube. Measurements (holotype female in µm). Body length about 2180 (distended). Head length 213, from anterior margin of eyes 195, width across eyes 175, maximum width across cheeks 185; eyes length 63; diameter of posterior ocelli 10; postocular setae 22 – 29. Maxillary bridge width 38. Antenna total length 350, segments III – VIII length (width) as followings: 45 (23), 48 (28), 53 (25), 48 (23), 50 (20), 30 (15). Pronotum length 215, width 218. Setae on prothorax: aa 18 – 20, ml 14 – 19, pa 14 – 25, epim 34 – 36. Fore wing length 740. Sub-basal wing setae: S 1 9 – 10, S 2 15 – 18, S 3 15 – 18. Pelta length 73, width 125. Tergite IX setae: S 1 84, S 2 128 – 133. Tube length 132, basal width 60; terminal setae 189 – 193. Male (macroptera). Distended body length 1.4 – 1.7 mm. Very similar to female in structure but smaller and paler. Body almost largely yellow (Fig. 117), but slightly shaded along inner margin of compound eyes and pleural areas of metathorax, posterior third of abdominal segment IX brown, tube brown with base yellowish. Antennal segments I – VI yellow with apices of V – VI shaded, VII – VIII brown. Head (Fig. 119) angular at cheeks just behind compound eyes, maxillary stylets not reaching postocular setae, their distance slightly wider than in female, about 0.2 times as wide as head. Measurements (paratype male in µm). Body length about 1410 (distended). Head length 178, from anterior margin of eyes 165, width across eyes 145, maximum width across cheeks 148; eyes length 60; diameter of posterior ocelli 10; postocular setae 18 – 20. Maxillary bridge width 45. Antenna total length 296, segments III – VIII length (width) as followings: 38 (23), 43 (24), 43 (21), 40 (20), 43 (18), 25 (13). Pronotum length 140, width 163. Setae on prothorax: aa 10 – 16, ml 10 – 11, pa 10 – 13, epim 30. Fore wing length 570. Sub-basal wing setae: S 1 5, S 2 9 – 12, S 3 10. Pelta length 58, width 85. Tergite IX setae: S 1 43 – 48, S 2 40. Tube length 115, basal width 43; terminal setae 153 – 160.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812404507E64C5F9DEAF6FFD2C.taxon	materials_examined	Type material. Holotype: female, Thailand, nr. Kanchanaburi, on bamboo, 30. viii. 1991, TN & SO. Paratypes: 23 females & 5 males collected together with holotype.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812404507E64C5F9DEAF6FFD2C.taxon	discussion	Comments. This species is very similar to P. bellatulus in female body color, but it can easily be distinguished from the latter species by the following features: antennal segments III and IV with (0 + 1) and (1 + 1 + 1) sense cones respectively (those segments each with (1 + 1) sense cones in bellatulus); head, thorax and abdominal segment VIII largely yellow in male (head, thorax and abdominal segment VIII largely brown in male in bellatulus).	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812406507864C5FCC5ABC0FC30.taxon	description	(Figs 125 – 133) Female (macroptera). Distended body length 2.0 – 2.4 mm. Body bicolored (Fig. 125). Head, prothorax, pleural areas of pterothorax and abdominal segments VIII – X brown; both meso- and metanota and meso- and metasterna yellow or slightly shaded; abdominal segments I – VII yellow, tergites III – VII each with an anteromedian brown marking but paler on VII. Fore femora brown, fore tibiae yellow with outer margin slightly shaded, fore tarsi yellow; mid- and hind legs yellow. Antennal segment I brown, segment II brown with apex slightly yellow, segments III – VI yellow, but VI slightly shaded at distal third, segment VII yellow with distal third shaded, segment VIII brown. Fore wings pale, slightly shaded at middle; major body setae pale. Head 1.2 times as long as wide (Fig. 127), 1.23 times in holotype, widest just behind compound eyes, without lateral tooth. Cheeks distinctly serrate, weakly rounded and weakly but somewhat distinctly narrowed towards base, slightly constricted near base. Postocular setae short and pointed, 153 µm apart from each other in holotype. Posterior ocelli relatively small, about 70 µm apart from each other. Antennal segment III with (1 + 1) sense cones, segments IV and V each with (1 + 1 + 1) sense cones; antennal segment VIII not constricted at base. Ratio length / width of antennal segments III – VIII (Fig. 133) in holotype as followings: 2.0, 1.9, 2.1, 2.2, 2.7, 2.7. Maxillary stylet reaching postocular setae, their distance about 0.1 times as wide as width across cheeks. Pronotum (Fig. 127) almost as long as wide, almost smooth and weakly sculptured along posterior margin, with a longitudinal line at middle; aa, ml and epim expanded at apex, aa a little longer than ml, pa pointed, epim the longest. Prosternum (Fig. 129) and mesopresternum typical of the genus. Metanotum almost smooth medially, median pair of setae small, about 60 µm apart from each other, 95 µm apart from anterior margin in holotype. Fore femora often slightly rugose at ventro-external portion, without inner tubercles or hump; fore tibial inner apical tooth distinct (Fig. 130). Fore wings without duplicated cilia; three sub-basal setae short, S 1 and S 2 subequal in length and expanded at apex, S 3 setae pointed and almost as long as or slightly shorter than S 2. Pelta hut-shaped (Fig. 131). Abdominal tergite VIII S 1 setae expanded at apex; tergite IX S 1 and S 2 setae pointed, S 1 about 0.6 – 0.7 times as long as tube, S 2 almost as long as tube or a little longer. Tube (Fig. 132) distinctly narrowed at basal sixth, tapered to apex, slightly constricted at apex, about 0.6 times as long as head, about 2.0 times as long as basal width; terminal setae about 1.4 times as long as tube. Measurements (holotype female in µm). Body length about 2420 (distended). Head length 238, from anterior margin of eyes 218, width across eyes 185, maximum width across cheeks 193; eyes length 78; diameter of posterior ocelli 10; postocular setae 27. Maxillary bridge width 48. Antenna total length 379, segments III – VIII length (width) as followings: 50 (25), 55 (29), 53 (25), 51 (23), 53 (20), 35 (13). Pronotum length 220, width 220. Setae on prothorax: aa 21 – 25, ml 15 – 16, pa 27, epim 40 – 43. Fore wing length 850. Sub-basal wing setae: S 1 23, S 2 20 – 23, S 3 13 – 15. Pelta length 58, width 130. Tergite IX setae: S 1 96 – 97, S 2 150 – 151. Tube length 148, basal width 63; terminal setae 213 – 215. Male (macroptera). Distended body length 1.7 – 1.8 mm. Very similar to female in structure, but smaller and paler. Thorax yellow but slightly shaded with brown at pleural areas of pterothorax (Fig. 126); all legs yellow; antennal segment I pale brown, II pale brown with apex pale. Head (Fig. 128) usually with distinct serrations at cheeks, maxillary stylets not reaching postocular setae, and their distance wider than in female, about 0.3 times as wide as width across cheeks. Measurements (paratype male in µm). Body length about 1690 (distended). Head length 193, from anterior margin of eyes 175, width across eyes 158, maximum width across cheeks 163; eyes length 68; diameter of posterior ocelli 13; postocular setae 15 – 18. Maxillary bridge width 55. Antenna total length 330, segments III – VIII length (width) as followings: 50 (24), 50 (25), 48 (24), 45 (20), 48 (18), 28 (11). Pronotum length 165, width 180. Setae on prothorax: aa 21, ml 11 – 14, pa 17, epim 35. Fore wing length 650. Sub-basal wing setae: S 1 10 – 13, S 2 13 – 15, S 3 13 – 15. Pelta length 60, width 83. Tergite IX setae: S 1 54, S 2 40 – 45. Tube length 103, basal width 50; terminal setae 170 – 173.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812406507864C5FCC5ABC0FC30.taxon	materials_examined	Type material. Holotype: female, Thailand, nr. Chiang Mai, Hong kai, on bamboo, 4. ix. 1991, TN & SO. Paratypes: Thailand, 24 females and 9 males collected together with holotype; 1 female, foot of Doi Inthanon, on grass, 4. ix. 1991, TN & SO; 1 female, Fang, on bamboo, 23. viii. 1992, SO; nr. Chiang Mai: Doi Suthep, on bamboo, 9 females and 3 males, 1. ix. 1991, TN & SO, 8 females and 6 males, 24. viii. 1992, TN & SO, 1 male, 2. ix. 1992, SO; 2 females, Pha Hean, on bamboo, 3. ix. 1992, TN; 1 female, Chiang Rai, on bamboo, 4. ix. 1992, SO; 1 female and 1 male, Doi Inthanon, on bamboo, 29. viii. 1992, TN & SO; 12 females and 4 males, Hang Dong, on bamboo, 25. viii. 1992, SO; 12 females and 2 males, Chiang Mai, on bamboo, 2. ix. 1991, TN & SO; 3 females and 1 male, Farm of Chiang Mai University, on bamboo, 26. viii. 1992, TN; 28 females and 11 males, Mae-sa, on bamboo, 3. ix. 1991, TN & SO; 26 females and 21 males, Chiang Dao, on bamboo, 23. viii. 1992, TN & SO; 25 females and 21 males, nr. Chiang Mai, Phrow, on bamboo, 22. viii. 1992, TN & SO. Non-paratypic specimens. Thailand, 1 male, Chiang Mai, Farm of Chiang Mai University, on bamboo, 26. viii. 1992, TN. Vietnam, 18 females and 8 males, Lam Dong Province, Finom Ward, Don Duong, on bamboo, 25. xii. 2001, SO; 1 female and 1 male, Thai Nguten Province, on bamboo, 22. viii. 2000, SO; Son La Province: 4 females, Moc Chau, on bamboo, 5. iii. 2000, SO; 1 female, Deo Pha Din, on bamboo, 13. viii. 2000, SO. Doubtful specimens. Thailand, 1 male, nr. Chiang Mai, Mae-sa, on bamboo, 3. ix. 1991, TN & SO; 1 female, foot of Doi Inthanon, on grass, 4. ix. 1991, TN & SO	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812406507864C5FCC5ABC0FC30.taxon	discussion	Comments. This species is described from Thailand based on a good number of females and males, and is very similar to P. taiwanus sp. nov. described below. It can be distinguished from taiwanus by the key above. Especially, males can clearly be distinguished from each other by the color of prothorax and the serration of the cheeks. The pronotum is yellow and the cheeks are usually with distinct serrations in asiaticus, whereas the pronotum is brown and the head with weak serration in taiwanus. In the male, this species cannot be distinguished satisfactorily from P. flavipes sp. nov. described below, but it can be distinguished by the difference in the shape of S 1 setae on tergite IX as in the key couplet 16 above. P. bellatulus is also similar to this species in the female body color and the shape of tube, but it can be distinguished by the following: antennal segment IV with (1 + 1 + 1) sense cones (with (1 + 1) sense cones in bellatulus); pterothorax largely yellow except for pleural areas brown in female (thorax largely brown in both sexes in bellatulus); cheeks distinctly serrated in male (weakly serrated in bellatulus). The male is somewhat similar in color to P. semiflavus but has antennal segment IV with (1 + 1 + 1) sense cones. The specimens from Vietnam listed under the non-paratypic specimens are probably conspecific, but have the prothoracic anteroangular and midlateral setae slightly longer and the male prothorax often slightly shaded. A female and a male from Thailand listed under the doubtful specimens are very similar to this species, but these have the tube much shorter (120 µm in female, 88 µm in male).	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812400507864C5FBCCA8ADF96A.taxon	description	(Figs 134 – 135)	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812400507864C5FBCCA8ADF96A.taxon	description	Specimens examined. Thailand, holotype female and paratype (allotype) male, Phuket, on bamboo, 19. viii. 1976, SO; Phuket, Phuket Hill, on bamboo, 6 females and 5 males, 27. viii. 1991, 6 females and 2 males, 9. ix. 1992, TN & SO; 4 females and 1 male, Phuket, Gu gu suri Road, on bamboo, 24. viii. 1991, TN & SO; Phuket, nr. Rawai Beach, on bamboo, 38 females and 22 males, 26. viii. 1991, 7 females & 6 males, 15. ix. 1992, TN & SO; 1 male, nr. Kanchanaburi, on bamboo, 30. viii. 1991, TN & SO.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812400507A64C5F8FBA974F98C.taxon	description	(Figs 136 – 137)	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812400507A64C5F8FBA974F98C.taxon	description	The specimens examined listed below from India, Southeast Asia and Japan exhibit structural variation in the shape and form of certain setae, such as postocular setae, prothoracic setae and sub-basal wing setae. It is difficult to know whether these specimens contain additional distinct species or not. The postocular setae are long and pointed in the specimens from India but those are rather shorter and expanded in the specimens from Singapore (Ritchie, 1974). In our observation, the prothoracic posteroangular setae are usually expanded in the specimens from Southeast Asia and Japan, and are rarely pointed. The sub-basal wing setae S 1 are rarely pointed in the specimens from Singapore. The postocular setae are rarely blunt or pointed in the specimens from Peninsular Malaysia. Moreover, the male posteromarginal setae S 1 on tergite IX of the specimens from India are pointed, whereas those setae are expanded, or at least blunt, in the specimens from Singapore, Peninsular Malaysia, Thailand and Japan. In a long series of specimens from Thailand examined here, a few specimens have (1 + 1) sense cones on the antennal segment IV, but it is difficult to determine whether the small outer sense cone is absent or missing. The description of this species is given below based on the specimens from Thailand, because the original description is insufficient for the present study. Female (macroptera). Distended body length 1.7 – 2.3 mm. Body bicolored (Fig. 136). Head, prothorax and abdominal segments VIII – X brown but segment VIII slightly yellowish in anterior to antecostal suture; pterothorax yellow with pleural areas shaded, abdominal segments I – VII yellow, tergites III – VI each with a brown marking anteromedially. All legs yellow. Antennal segments I – II brown with apex of II yellowish, III – V yellow, VI yellow with distal third shaded, VII brown with base yellowish, VIII brown. Fore wings pale, slightly shaded with brown at middle. Head widest just behind compound eyes (Fig. 137), without lateral tooth. Postocular setae developed, pointed, blunt or expanded. Cheeks rounded and somewhat distinctly narrowed towards base, slightly constricted near base. Antennal segment III with (1 + 1) sense cones, IV usually with (1 + 1 + 1) sense cones, but often with (1 + 1), V with (1 + 1 + 1) sense cones; segment VIII not constricted at base. Maxillary stylet reaching postocular setae. Prothoracic aa, ml and epim expanded, pa pointed, blunt or expanded. Fore femora unarmed; fore tibial inner apical tooth weak. Fore wings without duplicated cilia, sub-basal setae short, S 1 and S 2 expanded, S 3 setae pointed. Pelta hut-shaped. Abdominal tergite VIII S 1 setae expanded; tergite IX S 1 and S 2 setae pointed, S 1 about 0.7 times as long as tube, S 2 about 1.7 times as long as tube. Tube almost straight-sided, tapered to apex, about 0.5 times as long as head, 2.2 times as long as basal width. Male (macroptera). Distended body length 1.6 – 2.0 mm. Very similar in color and structure to female, but smaller; abdominal tergite IX with S 1 setae pointed or expanded. Specimens examined. India, 2 males, Bangalore, Lal Bagh, on bamboo, 25. iii. 1964, T. N. Ananthakrishnan; 1 female, Chalakudi, on bamboo, 5. xii. 1964, T. N. Ananthakrishnan. Thailand, 1 female and 1 male, Kamphaeng Saen, Campus of Kasetsart University, on bamboo, 12. i. 1988, SO; 4 females, Nakornprathom, Puttamonton, on bamboo, 18. ix. 1991, J. Pornthip; 1 male, Hang Dong, on bamboo, 25. viii. 1992, SO; Bangkok, Bangkhen, Campus of Kasetsart University, NBCRC, on bamboo, 4 females, 28. xii. 1987, SO, 4 females and 2 males, 11. i. 1988, SO, 5 females and 2 males, 22. viii. 1990, TN & SO; 2 females and 1 male, Chiang Mai, on bamboo, 2. ix. 1991, TN & SO; 56 females and 17 males, Phuket, Gu gu suri Road, on bamboo, 24. viii. 1991, TN & SO; 7 females and 4 males, Phuket Is., nr. Rawai Beach, on bamboo, 15. ix. 1992, TN & SO; 14 females and 8 males, nr. Rawai Beach, on bamboo, 26. Viii. 1991, TN & SO. Singapore, 11 females and 7 males, Macritchie Res., on bamboo, 17. viii. 1990, TN & SO. Peninsular Malaysia, 21 females and 2 males, Tapha, on bamboo, 17. ix. 1990, TN & SO. Japan, Okinawaken, Minamidaitô-jima Is., on bamboo, 1 females, 21. iii. 2008, K. Minoura. 12 females and 6 males 22. iii. 2016, K. Minoura. 9 females and 6 males, 22. iii. 2016, Kurahashi.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812402507B64C5F950A8F7F899.taxon	description	(Figs 144 – 153)	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812402507B64C5F950A8F7F899.taxon	description	The specimens from Vietnam listed below under the doubtful specimens (Figs 146 – 153) are very similar to this species. However, they are slightly different from etsukoae in the following features: head entirely brown (yellowish posteriorly in etsukoae); abdominal segment VIII entirely brown in both sexes (largely brown, but anterior portion yellowish in female, largely yellow in male in etsukoae); three sub-basal wing setae pointed (sub-basal wing setae S 2 expanded in etsukoae); fore wing with 4 – 7 duplicated cilia (with 2 – 4 duplicated cilia in etsukoae); metanotal median pair of setae short, about 15 µm long in female (more than 20 µm in etsukoae). These differences are minor, and it is not possible to know whether they are local populations of the same species until more specimens are examined. Specimens examined. Thailand, holotype female and all paratypes (4 females and 5 males), and 1 non-paratypic male (see Minoura & Masumoto, 2020). Doubtful specimens. Vietnam, 6 females and 4 males, Thua Thien Hue Prov., Phu Loc District, Bach Ma National Park, on bamboo, 14. viii. 2007, SO.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781240C507464C5FD7BA875FAB2.taxon	description	(Fig. 251)	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781240C507464C5FD7BA875FAB2.taxon	description	Specimens examined. Peninsular Malaysia, holotype female (see Okajima 1978). Thailand: 2 paratype males (including allotype) (see Okajima 1978); 1 female, Chiang Rai, on bamboo, 4. ix. 1992, SO; 16 females and 10 males, Phuket, Gu gu suri Road, on bamboo, 24. viii. 1991, TN & SO; Phuket, nr. Rawai Beach, on bamboo, 1 female and 1 male, 26. viii. 1991, 8 females and 4 males on bamboo, 15. ix. 1992, TN & SO. Vietnam, 1 male, Dong Nai Province, Tan Phu, Phu An, on dead leaves and branches, 29. xii. 2001, SO; 1 female, Nghe An Prov., Anh Son District, Cao Son, ca 35 m alt., on bamboo, 9. viii. 2007, SO. Borneo: 1 male, Sabah, nr. Keningaw, Taman Bunsit, on bamboo, 2. ix. 1990, TN & SO.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781240C507664C5FA32A96AFC80.taxon	description	(Figs 154 – 161) Female (macroptera). Distended body length 2.1 – 2.3 mm. Body bicolored (Fig. 154). Head and abdominal segments VIII – X brown, but segment VIII usually slightly yellowish before antecostal suture; thorax yellow but metathoracic pleural areas shaded with brown; abdominal segments I – VII yellow, tergites III – VII each with an anteromedian brown marking, but paler on VII. All legs yellow, but fore tibiae shaded at outer margin. Antennal segments I – II brown with apex of II yellowish; III – VII yellow with apices of V – VII shaded; VIII brown. Fore wings pale, slightly shaded at middle; major body setae pale. Head about 1.2 times as long as wide (Fig. 155), 1.23 times in holotype, widest just behind compound eyes, without lateral tooth. Cheeks rounded but somewhat distinctly narrowed towards base, slightly constricted near base. Postocular setae rather developed, pointed, 153 µm apart from each other in holotype. Posterior ocelli 65 µm apart from each other in holotype. Antennal segment III with (1 + 1) sense cones, segments IV and V each with (1 + 1 + 1) sense cones; segment VIII not constricted at base. Ratio length / width of antennal segments III – VIII (Fig. 161) in holotype as followings: 2.1, 2.0, 2.2, 2.2, 2.9, 2.5. Maxillary stylet reaching postocular setae, their distance about 0.1 times as wide as width across cheeks. Pronotum (Fig. 155) about 0.9 times as long as wide, almost smooth and weakly sculptured along posterior margin, with a longitudinal line at middle; aa, ml and epim expanded, pa pointed, epim the longest. Prosternum and mesopresternum typical of the genus (Fig. 158). Metanotum almost smooth anteromedially; median pair of setae 58 µm apart from each other, 80 – 84 µm apart from anterior margin in holotype. Fore femora unarmed; fore tibial inner apical tooth distinct (Fig. 157). Fore wings without duplicated cilia; sub-basal wing setae short, S 1 and S 2 subequal in length, expanded, S 3 shorter than S 2, pointed. Pelta hut-shaped (Fig. 159). Abdominal tergite VIII S 1 setae expanded; tergite IX S 1 pointed or blunt, S 2 pointed, S 1 about 0.7 times as long as tube; S 2 almost as long as tube. Tube (Fig. 160) distinctly narrowed at basal one-eighth, weakly tapered towards apex, constricted at apex, 0.6 times as long as head, about 2.3 times as long as basal width; terminal setae 1.4 – 1.5 times as long as tube. Measurements (holotype female in µm). Body length about 2250 (distended). Head length 238, from anterior margin of eyes 218, width across eyes 185, maximum width across cheeks 193; eyes length 78; diameter of posterior ocelli 15; postocular setae 37 – 40. Maxillary bridge width 45. Antenna total length 384, segments III – VIII length (width) as followings: 53 (25), 55 (28), 55 (25), 50 (23), 56 (19), 33 (13). Pronotum length 203, width 225. Setae on prothorax: aa 24 – 33, ml 18 – 19, pa 26 – 33, epim 45 – 48. Fore wing length 810. Sub-basal wing setae: S 1 23, S 2 23 – 28, S 3 15. Pelta length 78, width 120. Tergite IX setae: S 1 105 – 106, S 2 143 – 145. Tube length 143, basal width 63; terminal setae 193 – 209. Male (macroptera). Distended body length 1.4 – 1.6 mm. Very similar to female in color and structure, but smaller. Maxillary stylets not reaching postocular setae, their distance wider than that of female, about 0.3 times as wide as width across cheeks (Fig. 156). Abdominal tergite IX with S 1 setae expanded. Measurements (paratype male in µm). Body length about 1450 (distended). Head length 183, from anterior margin of eyes 170, width across eyes 150, maximum width across cheeks 153; eyes length 63; diameter of posterior ocelli 13; postocular setae 20 – 22. Maxillary bridge width 58. Antenna total length 314, segments III – VIII length (width) as followings: 43 (23), 48 (24), 45 (23), 41 (20), 48 (18), 28 (11). Pronotum length 135, width 153. Setae on prothorax: aa 14 – 18, ml 13 – 15, pa 14 – 16, epim 30 – 35. Fore wing length 550. Sub-basal wing setae: S 1 15, S 2 20, S 3 13 – 15. Pelta length 55, width 80. Tergite IX setae: S 1 46 – 47, S 2 38. Tube length 99, basal width 43; terminal setae 150 – 153.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781240C507664C5FA32A96AFC80.taxon	materials_examined	Type material. Holotype: female, Thailand, Kamphaeng Saen, Campus of Kasetsart University, on bamboo, 12. i. 1988, SO. Paratypes: Thailand, 9 females and 1 male collected together with holotype; Bangkok, Bangkhen, Campus of Kasetsart University, NBCRC, on bamboo, 1 female and 1 male, 28. xii. 1987, SO, 2 females, 22. viii. 1990, TN & SO; 1 male, Nakornprathom, Puttamonton, on bamboo, 18. ix. 1991, J. Pornthip. Non-paratypic specimens. Thailand, 1 male, Bangkok, Bangkhen, Campus of Kasetsart University, on dead branches, 19. viii. 1991, TN & SO.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781240C507664C5FA32A96AFC80.taxon	discussion	Comments. This species is very similar to P. bicolor in having similar body color, antennal segment IV with (1 + 1 + 1) sense cones, the fore wings without duplicated cilia, and the sub-basal wing setae S 1 and S 2 expanded. However, flavipes has the prothorax largely yellow in both sexes, though bicolor has the prothorax largely brown in both sexes. Moreover, flavipes is somewhat similar to a widely distributed species, P. semiflavus, in having the body color similar. It can be distinguished by the following features: antennal segment IV with (1 + 1 + 1) sense cones (with (1 + 1) sense cones in semiflavus); fore wing sub-basal setae S 1 and S 2 expanded (all sub-basal wing setae pointed in semiflavus); male tube much shorter, about 100 µm (about 140 µm in semiflavus); male tergite IX with S 1 setae expanded at apex (pointed in semiflavus). A male listed under the non-paratypic specimen has S 1 setae on the abdominal tergite IX blunt at apex, at least not expanded	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781240E507764C5FC51A983FDC4.taxon	description	(Figs 162 – 169) Female (macroptera). Distended body length 2.1 – 2.5 mm. Body bicolored (Fig. 162). Head, prothorax and abdominal segments VIII – X brown; pterothorax yellow with pleural areas shaded with brown; abdominal segments I – VII yellow, tergites III – VII each with an anteromedian brown marking, but paler on VII. All legs yellow. Antennal segment I brown, II brown with apex paler; III – VI yellow, apices of V – VI shaded; VII – VIII brown. Fore wings pale, slightly shaded with brown at middle; major body setae pale. Head 1.3 times as long as wide (Fig. 163), 1.31 times in holotype, widest just behind compound eyes, without lateral tooth. Cheeks slightly rounded and weakly narrowed towards base, slightly constricted near base. Postocular setae short and pointed, 156 µm apart from each other in holotype. Posterior ocelli relatively large, almost as large as anterior one, 67 µm apart from each other in holotype. Antennal segment III with (1 + 1) sense cones; segments IV and V each with (1 + 1 + 1) sense cones; segment VIII elongate, not constricted at base, tapered at distal half (Fig. 168). Ratio length / width of antennal segments III – VIII (Fig. 169) in holotype as followings: 2.3, 2.1, 2.7, 2.5, 2.9, 3.3. Maxillary stylet reaching postocular setae, their distance about 0.2 times as wide as width across cheeks. Pronotum (Fig. 163) about 0.9 times as long as wide, 0.93 times in holotype, almost smooth and weakly sculptured along posterior margin, with a longitudinal line at middle; aa, ml and pa pointed, epim expanded and the longest. Prosternum and mesopresternum typical of the genus (Fig. 165). Metanotum almost smooth medially; median pair of setae 57 µm apart from each other, 77 – 82 µm apart from anterior margin in holotype. Fore femora unarmed; fore tibial inner apical tooth weak (Fig. 164). Fore wings usually with 1 – 3 duplicated cilia (only one at left in holotype); sub-basal wing setae short and acute, at most blunt. Pelta hut-shaped (Fig. 166). Abdominal tergite VIII S 1 setae blunt or pointed; tergite IX S 1 and S 2 setae pointed, S 1 0.7 – 0.8 times as long as tube, S 2 about 1.5 times as long as tube. Tube (Fig. 167) slightly narrowed at basal fifth, straight-sided, weakly tapered to apex, 0.5 times as long as head, about 2.2 times as long as basal width; terminal setae 1.6 – 1.8 times as long as tube. Measurements (holotype female in µm). Body length about 2340 (distended). Head length 260, from anterior margin of eyes 238, width across eyes 193, maximum width across cheeks 198; eyes length 90; diameter of posterior ocelli 15; postocular setae 28 – 34. Maxillary bridge width 48. Antenna total length 415, segments III – VIII length (width) as followings: 63 (28), 58 (28), 63 (23), 53 (21), 53 (18), 43 (13). Pronotum length 213, width 230. Setae on prothorax: aa 13 – 16, ml 14 – 18, pa 21 – 23, epim 43. Fore wing length 830. Sub-basal wing setae: S 1 15 – 18, S 2 12 – 14, S 3 15 – 18. Pelta length 75, width 110. Tergite IX setae: S 1 92 – 98, S 2 194 – 197. Tube length 130, basal width 60; terminal setae 213 – 231. Male. Unknown.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781240E507764C5FC51A983FDC4.taxon	materials_examined	Type material. Holotype: female, Thailand, nr. Chiang Mai, on bamboo, 24. viii. 1992, TN & SO. Paratypes: 2 females collected together with holotype; 1 female, nr. Chiang Mai, Doi Suthep, on bamboo, 2. ix. 1992, SO.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781240E507764C5FC51A983FDC4.taxon	discussion	Comments. This species is described from Thailand based on four females. It is very similar to P. flavipes sp. nov. described above also from Thailand, but it can be distinguished from flavipes by the following features: prothorax brown; fore wing usually with duplicated cilia (without duplicated cilia in flavipes); sub-basal wing setae S 1 and S 2 pointed, at most blunt, not expanded; fore tibia with weak inner sub-apical teeth (with distinct sub-apical teeth in flavipes). P. odonaspicola from Japan is also very similar to this species in having the body color pattern similar and the fore wing duplicated cilia present, but it has antennal segment IV with (1 + 1) sense cones, the fore tibiae without distinct inner apical tooth and the fore femora largely brown with apical one-third yellowish. Moreover, antennal segment VIII of gracilicornis is longer and more slender.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781240F507064C5F981AE11FE28.taxon	description	Specimens examined. Taiwan, Pintung Hsien, Kenting National Park, on grass, 2 females, 22. v. 1972, 10 females and 3 males, 18. iii. 1984, 8 females and 7 males, 19. iii. 1984, SO; 2 females, Kaohsiung Hsien, Liukuei, 22. iii. 1984, SO. Indonesia, Bali, 2 female, Tabanan, Bedugul, ca 1250 m alt., on grass, 7. iii. 2005, SO. Peninsular Malaysia, 1 female, Tapha, on grass, 26. vii. 1976, SO; 3 females and 4 males, N. E. 13 Km from Tapha, sweeping, 27. vii. 1976, SO. Thailand, Chiang Mai, rice field, 3 females, 4. v. 1978, 1 female, 7. v. 1978, 1 male, 24. vi. 1979, K. Yasumatsu; 1 female, nr. Chiang Mai, foot of Doi Pui, on grass, 24. viii. 1992, TN & SO; 1 female, foot of Doi Inthanon, on grass, 29. viii. 1992, TN & SO; 8 females and 2 males, Doi Suthep, 500 m alt, on grass, 8. viii. 1976, SO; Kamphaeng Saen, Campus of Kasetsart University, on grass, 1 male, 22. xii. 1978, 5 females and 12 males, 29. xii. 1987, 20 females and 19 males, 12. i. 1988, SO; 1 females and 2 males, Mangasan Ubot, Phibun, 10. xi. 1977, K. Yasumatsu; 1 female and 1 male, Bannanpriew, Chacheongsao, wild rice, 12. vii. 1978, K. Yasumatsu. Vietnam, 1 male, Ha Noi, Farm of Ha Noi Agricultural University, habitat unknown, 29. vii. 2000, SO. Japan: 1 female and 2 males (see Okajima, 2006). Many females and males from India, Peninsular Malaysia (detailed data are omitted).	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812408507064C5FDB4AF41FC3A.taxon	description	(Figs 138 – 139)	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812408507064C5FDB4AF41FC3A.taxon	description	Specimens examined. Taiwan, holotype female, Pingtung Hsien, Kenting National Park, on grass, 26. v. 1972, SO; same locality and habitat, 10 female and 2 males, 18. iii. 1984, 9 females and 9 males, 19. iii. 1984, SO.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812408507064C5FBCBAF09FA07.taxon	description	(Figs 5, 140 – 141, 255)	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812408507064C5FBCBAF09FA07.taxon	description	Specimens examined. Japan, Honshu, many females and males (detailed data omitted, see Okajima 2006).	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812408507164C5F9D9A837FE54.taxon	description	(Figs 142 – 143)	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812408507164C5F9D9A837FE54.taxon	description	Specimens examined. Japan, many females and males from Hachijo Is., Okinawa Is. and the Ogasawara Islands (detailed data are omitted, see Okajima 2006). Additional record: Japan, 3 females, Honshu, Chiba-ken, Tateyamashi, Mera, on bamboo, 12. vi. 1999, TN.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812409507164C5FE28ABC5FBF2.taxon	description	Specimens examined. Uganda, Lugazi, ex Aulacaspis regalensis on sugar cane, 1 female, 16. i. 1969, 1 male, 31. i. 1969, D. J. Greathead.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812409507364C5FB73AF4AFDBD.taxon	description	(Figs 170 – 177) Female (macroptera). Distended body length 2.2 – 2.4 mm. Body bicolored (Fig 170). Head, prothorax, metathoracic pleural areas and abdominal segments IX – X brown; mesonotum usually yellowish brown, rarely yellow; abdominal segment VIII brown with anterior fourth and median half yellowish; metathorax yellow with episternum shaded; abdominal segments I – VII yellow, tergites III – VI (or VII) each with an anteromedian brown marking, those on VI and VII paler. Fore femora brown, fore tibiae yellow with outer margin shaded, fore tarsi yellow; mid- and hind legs yellow. Antennal segment I brown, II brown with apex yellowish; III – VI yellow, but VI rarely shaded at distal third; VII – VIII usually brown, VII often yellowish at basal two-thirds. Fore wings pale, slightly shaded brown at middle; major body setae pale. Head 1.3 times as long as wide (Fig. 171), 1.31 times in holotype, widest just behind compound eyes, without lateral tooth. Cheeks weakly serrate, rounded and weakly narrowed towards base, slightly constricted near base. Postocular setae short and pointed, about 150 µm apart from each other in holotype. Posterior ocelli smaller than anterior ocellus, about 70 µm apart from each other in holotype. Antennal segment III with (1 + 1) sense cones, segments IV and V each with (1 + 1 + 1) sense cones; segment VIII not constricted at base. Ratio length / width of antennal segments III – VIII (Fig. 177) in holotype as followings: 1.9, 1.8, 2.1, 2.2, 2.4, 2.8. Maxillary stylet reaching postocular setae, their distance about 0.2 times as wide as width across cheeks. Pronotum (Fig. 171) almost as long as wide, almost smooth and weakly sculptured along posterior margin, with a longitudinal line at middle; aa, ml and epim expanded, pa pointed, epim the longest. Prosternum and mesopresternum typical of the genus (Fig. 174); basantra elongate, about twice as long as wide in holotype. Metanotum almost smooth medially, median pair of setae 58 µm apart from each other, 83 – 85 µm apart from anterior margin. Fore femora unarmed; fore tibial apical tooth distinct (Fig. 173). Fore wings without duplicated cilia; sub-basal wing setae short, S 1 and S 2 subequal in length, expanded, S 3 pointed, shorter than S 2. Pelta hut-shaped (Fig. 175). Abdominal tergite VIII S 1 setae expanded; tergite IX S 1 and S 2 setae pointed, S 1 0.7 times as long as tube, S 2 almost as long as tube. Tube (Fig. 176) somewhat narrowed at basal sixth, straight-sided, weakly tapered towards apex, 0.6 times as long as head, 2.2 times as long as basal width; terminal setae 1.4 – 1.5 times as long as tube. Measurements (holotype female in µm). Body length about 2390 (distended). Head length 243, from anterior margin of eyes 220, width across eyes 180, maximum width across cheeks 185; eyes length 75; diameter of posterior ocelli 13; postocular setae 37 – 39. Maxillary bridge width 38. Antenna total length 386, segments III – VIII length (width) as followings: 53 (28), 55 (30), 58 (28), 50 (23), 51 (21), 37 (13). Pronotum length 223, width 228. Setae on prothorax: aa 23 – 26, ml 14, pa 23 – 29, epim 38 – 43. Fore wing length 800. Sub-basal wing setae: S 1 18, S 2 18, S 3 15. Pelta length 80, width 120. Tergite IX setae: S 1 104 – 107, S 2 147 – 155. Tube length 150, maximum width 68; terminal setae 205 – 218. Male (macroptera). Distended body length 1.7 – 1.9 mm. Very similar to female in color and structures, but smaller and paler. All legs yellow, pterothorax paler; abdominal segment VIII largely yellow with both sides shaded. Cheeks with weak serrations; maxillary stylets not reaching postocular setae, their distance a little wider than that of female, about 0.2 times as wide as head (Fig. 172). Measurements (paratype male in µm). Body length about 1910 (distended). Head length 215, from anterior margin of eyes 198, width across eyes 153, maximum width across cheeks 165; eyes length 73; diameter of posterior ocelli 13; postocular setae 17 – 29. Maxillary bridge width 50. Antenna total length 343, segments III – VIII length (width) as followings: 48 (25), 53 (28), 53 (25), 45 (23), 48 (18), 33 (11). Pronotum length 185, width 195. Setae on prothorax: aa 17, ml 9 – 11, pa 22, epim 35. Fore wing length 710. Sub-basal wing setae: S 1 15 – 18, S 2 15 – 18, S 3 13 – 15. Pelta length 63, width 108. Tergite IX setae: S 1 65 – 67, S 2 45. Tube length 128, basal width 53; terminal setae 178 – 183.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812409507364C5FB73AF4AFDBD.taxon	materials_examined	Type material. Holotype: female, Taiwan, Kaohsiung-hsien, Liukuel, on bamboo, 5. ix. 1993, TN & SO. Paratypes: Taiwan, 19 females and 11 males collected together with holotype; 13 females and 9 males, Taitunghsien, Chihpen, 10. ix. 1993, TN & SO.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812409507364C5FB73AF4AFDBD.taxon	discussion	Comments. This species is described from Southern Taiwan. It is similar to P. femoralis Dang et al. described from Yunnan, China in the body color and the antennal segment IV with (1 + 1 + 1) sense cones. However, it may can be distinguished from femoralis by the following features (in parentheses are the features of femoralis): metathorax largely yellow except for pleural areas (pterothorax largely brown); pronotal posteroangular setae pointed (expanded); pronotal midlateral setae very small, much shorter than anteroangular setae (developed, almost as long as anteroangular setae); terminal tube setae 1.4 – 1.5 times as long as tube (about 1.7 times as long as tube). Moreover, the tube of taiwanus is somewhat longer, 150 µm long and about 0.6 times as long as head, while that of femoralis is 140 µm long and about 0.5 times as long as head. P. bellatulus described from Thailand is also similar to this new species in the body color, but it can be distinguished from taiwanus by the followings: antennal segment IV with (1 + 1) sense-cones; basantra shorter, much shorter than twice as long as wide; metathorax brown.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781240B506C64C5FD40AFE7FD85.taxon	description	Among eight species including a new species, two species, P. causiapeltus and P. priesneri, have three (1 + 2) and four (2 + 2 + 1) sense cones on antennal segments III and IV respectively. In contrast, the remaining species each have two (1 + 1) sense cones on those segments, but P. nigrocephalus from India has only one (0 + 1) sense cone on segment III. Most other character states are very similar, and we judge here that these eight species belong to a single genus. Interspecific variation within a genus in the number of sense cones present is not particularly uncommon, and is ubiquitous within the Haplothripini (Okajima & Masumoto 2025 a, 2025 b). Diagnosis. Body slightly flattened, usually macropterous. Head longer than wide, with a pair of postocular setae. Postocular setae arose near cheeks. Eyes and ocelli well-developed. Antennae eight-segmented; segment III with (0 + 1), (1 + 1) or (1 + 2) sense cones, segment IV with (1 + 1) or (2 + 2 + 1) sense cones. Mouth cone short and rounded; maxillary stylets retracted rather short and wide apart, at least not reaching postocular setae; maxillary bridge present. Prothorax with three or four pairs of expanded setae, am reduced, ml often reduced; notopleural sutures complete. Basantra present, wider than long. Mesopresternum transverse; metathoracic sternopleural sutures well-developed, furcal arms separated from one another (cf. Fig. 252), without spinula. Fore tarsal tooth present in both sexes, fore tibiae unarmed. Fore wings weakly constricted medially (cf. Fig. 256), with duplicated cilia. Pelta hat-shaped, often trapezoid. Tergites II – VII with two pairs of wing-retaining setae. Tube shorter than head, straight-sided; terminal setae much longer than tube. Male sternite VIII without pore plate.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812414506C64C5FD52AF14F919.taxon	description	(Figs 178 – 184)	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812414506C64C5FD52AF14F919.taxon	description	Specimens examined. Thailand, 6 females and 4 males, Bangkok, Bangkhen, Campus of Kasetsart University, NBCRC, on bamboo, 22. viii. 1990, SO; Phuket, Phuket Hill, on bamboo, 9 females and 7 males, 27. viii. 1991, 7 females and 6 males, 9. ix. 1992, TN & SO; Phuket, nr. Rawai Beach, on bamboo, 1 female, 26. viii. 1991, 15 females and 9 males, 15. ix. 1992, TN & SO; 5 females and 6 males, nr. Kanchanaburi, on bamboo, 30. viii. 1991, TN & SO; 1 male, Chiang Mai, on bamboo, 2. ix. 1991, TN & SO; 2 males, Chiang Dao, on bamboo, 23. viii. 1992, TN & SO; 1 female, nr. Cgiang Mai, Doi Inthanon, on bamboo, 29. viii. 1992, TN & SO; 1 male, Chiang Mai, Farm of Chiang Mai University, on bamboo, 26. viii. 1992, TN; 1 female, nr. Chiang Mai, Doi Suthep, on bamboo, 2. ix. 1992, SO; 3 females and 1 male, nr. Chiang Mai, Ka Jan, on bamboo, 3. ix. 1992, SO. Vietnam, 39 females and 17 males, Hoa Binh Province, Hoa Binh, on bamboo, 6. iii. 2000, SO; Ha Tay Province, Mt. Ba vi, on bamboo, 12 females and 6 males, 8. iii. 2000, 2 females, 29. vii. 2000, SO; 1 male, Bac Can Province, Ba Be National Park, on bamboo, 2. viii. 2000, SO; 17 females and 13 males, Thai Nguyen Province, on bamboo, 22. viii. 2000, SO; 1 male, Lam Dong Province, Bao Loc, Dam Bri, on bamboo, 26. xii. 2001, SO; 2 females, Nghe An Province, Thanh Chuong District, Thanh An, Cau Cau, 9. viii. 2007, SO. Peninsular Malaysia, 1 male, Tapah, on bamboo, 17. ix. 1990, TN & SO.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812414506D64C5F8F9ABC8FEC0.taxon	description	(Fig. 199)	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812414506D64C5F8F9ABC8FEC0.taxon	description	Specimen-examined. China, 1 paratype female, Hainan Is., Jianfeng Mountain, 400 m, on Miscanthus sinensis, 4. iv. 1980, Zhang Weiqiu.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812416506E64C5FF65AEC8FD15.taxon	description	(Figs 185 – 190)	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812416506E64C5FF65AEC8FD15.taxon	description	Specimens examined. Thailand, 1 female, Doi Suthep, 500 m alt., on grass, 8. viii. 1976, SO; 1 female, Sarabri, Farm of Kasetsart University, on grass, 20. viii. 1991, TN & SO; nr. Chiang Mai, Doi Suthep, 1 female, on bamboo, 23 females, on grass, 1. ix. 1991, TN & SO, 2 females, on grass, 2. ix. 1992, SO; 17 females, foot of Doi Suthep, on grass, 24. viii. 1992, SO; 1 female, nr. Chiang Mai, Pha Hean, habitat uncertain, 3. ix. 1992, SO.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812416506F64C5FCFDA878F84D.taxon	description	(Figs 191 – 198) Female (macroptera). Distended body length: 2.2 – 2.4 mm. Body brown (Fig. 191). Femora brown, fore femora with extreme apices scarcely paler; fore tibiae yellow, mid and hind tibiae brown; tarsi yellow. Antennal segments I – II and VIII pale brown, segment II yellowish antero-externally; segments III – VI yellow; segment VII yellow to pale brown, gradually darkened distally. Fore wings and major body setae pale. Head about 1.2 times as long as wide (Fig. 192), widest across cheeks at a level of postocular setae, dorsal surface with a few transverse wrinkles at middle, with transverse reticulation at base. Cheeks rounded, weakly serrated. Postocular setae shorter than eyes, blunt or very weakly expanded, 168 µm apart from each other, 20 – 24 µm apart from eyes in holotype. Eyes about one-third of head length, about 90 µm apart from each other. Posterior ocelli slightly separated from eyes, 47 µm apart from each other. Antennae (Fig. 197) about 1.5 times as long as head; segment VIII scarcely constricted at base, 0.78 times as long as segment VII; segment III longer than segment IV; segments III and IV each with (1 + 1) sense cones. Maxillary stylets not reaching postocular setae, rather close together; maxillary bridge distinct (Fig. 193), about one-fourth of head width or a little wider, situated at basal one-fourth of head. Pronotum (Fig. 192) well-developed, 0.74 times as long as head, 1.28 times as wide as long in holotype, almost smooth; four pairs of major prothoracic setae developed, blunt or weakly expanded, am reduced; ml longer than aa, epim the longest, almost as long as postocular setae. Prospinasternum longer than wide (Fig. 194). Mesopresternum transverse. Mesonotum weakly reticulate, lateral setae short and pointed, about 15 µm long. Metanotum weakly sculptured with polygonal reticulation; median pair of setae short and acute, 52 µm apart from each other, 75 – 77 µm apart from anterior margin of metanotum in holotype. Fore tarsal tooth small (Fig. 198). Fore wing with 5 – 6 duplicated cilia; sub-basal setae S 1 and S 2 weakly expanded, S 3 blunt or very weakly expanded. Pelta hat-shaped (Fig. 195), sculptured weakly, about 1.8 times as wide as long. Tergite IX S 1 and S 2 setae pointed or bluntly pointed, shorter than tube, S 2 longer than S 1; intermediate setae shorter than half of S 1. Tube (Fig. 196) about 0.5 times as long as head, about 1.8 times as long as wide; terminal setae long, about 1.8 times as long as tube. Measurements (holotype female in µm). Body length 2230 (distended). Head length 243, from anterior margin of eyes 225, width across eyes 194, maximum width across cheeks 203, minimum width across base 150; eyes length 80; diameter of posterior ocelli 15 – 17; postocular setae 55. Maxillary bridge width 53. Antenna total length 370, segments III – VIII length (width) as follows: 57 (28), 52 (26), 50 (26), 47 (22), 45 (20), 35 (12). Pronotum length 180, width 230. Setae on prothorax: aa 35, ml 38, pa 43 – 44, epim 56 – 58. Fore wing length 800. Sub-basal wing setae: S 1 29 – 30, S 2 25 – 30, S 3 25 – 30. Pelta length 72, width 130. Tergite IX setae: S 1 105 – 110, S 2 115 – 118. Tube length 124, maximum width 69; terminal setae 220 – 225. Male (macroptera). Distended body length: about 2.0 mm. Very similar to macropterous female in color and structure. Pronotum 0.73 times as long as head, about 1.3 times as wide as long; fore wing with 5 duplicated cilia; tergite IX S 2 setae a little longer than half of S 1. Measurements (paratype male in µm). Body length about 2000 (distended). Head length 223, from anterior margin of eyes 207, width across eyes 181, maximum width across cheeks 185, minimum width across base 140; eyes length 78; diameter of posterior ocelli 13 – 17; postocular setae about 50. Maxillary bridge width 58. Antenna total length 370, segments III – VIII length as follows: 55, 55, 52, 45, 45, 34. Pronotum length 163, width 212. Setae on prothorax: aa about 20, ml about 30, pa 50, epim 55. Fore wing length 760. Sub-basal wing setae: S 1 33, S 2?, S 3?. Pelta length 71, width 117. Tergite IX setae: S 1 105 – 110, S 2 55 – 60. Tube length 117, maximum width 63; terminal setae 225.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812416506F64C5FCFDA878F84D.taxon	materials_examined	Type material. Holotype: macropterous female, Vietnam, Son La Province, Deo Pha Din, on bamboo, 13. viii. 2000, SO. Paratypes: Vietnam, 2 females and 1 male, collected together with holotype.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812416506F64C5FCFDA878F84D.taxon	discussion	Comments. This species is described from Vietnam based on three females and one male taken from bamboo. Similar to two previously known species, P. cymbapogoni and P. flavicornis, it has the body uniformly brown and antennal segments III and IV each with (1 + 1) sense cones. From cymbapogoni, sonlae can easily be distinguished by having brown mid and hind tibiae. From flavicornis, the most similar species, sonlae may can be distinguished by the maxillary bridge much narrower than one-third of the head width, and the prothoracic anteroangular and midlateral setae well-developed.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812410506864C5FDA9ABB7F97E.taxon	diagnosis	Diagnosis. Known only from macropterae, major body setae stout and expanded. Head almost as long as wide, or a little longer, dorsal surface distinctly reticulate, with a pair of postocular setae. Postocular setae arose behind mid-point of eyes. Eyes and ocelli well-developed. Antennae eight-segmented, segment VIII slightly constricted at base; segment III and IV with (1 + 1) and (1 + 2 + 1) sense cones respectively. Mouth cone short and rounded; maxillary stylets short, scarcely retracted into head capsule, V-shaped; maxillary bridge absent, at least invisible. Pronotum with 5 pairs of expanded setae; notopleural sutures complete. Basantra present. Mesopresternum transverse; sternopleural sutures absent, metaspinula present. Fore tarsi unarmed in both sexes. Fore wings weakly constricted medially, with duplicated cilia. Pelta hat-shaped. Tergites II – VIII with 2 pairs of wing-retaining setae. Tube straight-sided, shorter than head; terminal setae a little shorter than tube. Male: sternite VIII without pore plate; tergite IX S 2 setae much shorter than S 1.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812410506A64C5F91BAB2BF9D4.taxon	description	(Figs 201 – 206)	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812410506A64C5F91BAB2BF9D4.taxon	description	Nine females (Figs 208 – 211) and seven males collected from Java, Indonesia, Peninsular Malaysia and Vietnam listed below under the doubtful specimens are apparently very similar to this species, and most of them were collected from bamboo. Moreover, these specimens are not clearly distinguishable from the original description of Bamboosiella rugata, and there is a possibility that they are same species. These specimens are slightly different from mindorensis in several structures as follows: postocellar setae longer than a diameter of posterior ocellus; prothoracic anteroangular setae shorter; prospinasternum larger; metanotal sculptures more or less coarser (Fig. 210); pelta sculptured with polygonal reticulation (Fig. 211), instead of longitudinal reticulation (Fig 205); lateral wings of pelta with lateral corners rounded (Fig. 211), instead of pointed (Fig. 205); tergite VIII with only posterior pair of wing-retaining setae in female from Peninsular Malaysia and Vietnum; tergite IX S 2 setae of female stouter, clearly expanded at apex, instead of blunt. Thus, two taxonomic problems are involved. One is whether these should be distinguished from Bamboosiella and kept in the genus Propesolomonthrips. The other is whether mindorensis and rugata are different species, or these two are local populations of the same species. In any case, further study will be required in comparison with the original specimens of B. rugata. One important character state, the wing-retaining setae on tergite VIII is not mentioned in the original description of rugata. Female (macroptera) from Mindanao Is. (This specimen is not in good condition. In particular, the head and antennae are tilted upwards in front). Distended body length: about 1.4 mm. Color (Fig. 201) very similar to male. Head a little longer than wide (Fig. 202), widest across eyes; postocellar setae shorter than diameter of posterior ocellus. Eyes well-developed, wider apart ventrally, 0.45 times as long as head. Posterior ocelli about 17 µm apart from each other. Postocular setae 96 µm apart from each other. Pronotum (Fig. 202) 0.54 times as long as head, about 2.1 times as wide as long, with distinct transverse wrinkles on basal one-third; five pairs of major setae stout, aa and ml shorter than ml, epim a little shorter than postocular setae. Basantra relatively well-developed (Fig. 203). Prospinasternum relatively large. Mesonotal lateral setae long and stout, expanded, about 30 µm long. Metanotal median pair of setae 55 µm apart from each other, about 30 µm apart from anterior margin of metanotum. Fore wings with 4 duplicated cilia; sub-basal setae S 2 shorter than S 1, S 1 and S 3 subequal. Tergite IX S 2 setae very weakly expanded, longer than S 1; intermediate setae about half of S 1. Tube (Fig. 207) 0.64 times as long as head, about 2.0 times as long as wide. Other structures including the wing-retaining setae on the tergite VIII are very similar to male description. Measurements (macropterous female from Mindanao Is. in µm). Body length 1420 (distended). Head length 160, from anterior margin of eyes 140, width across eyes 140, maximum width across cheeks 138, minimum width across base 118; eyes length 75; diameter of posterior ocelli 15 – 20; postocular setae 45. Antenna total length 270, segments III – VIII length as follows: 36, 38, 37, 38, 35, 23. Pronotum length 86, width 178. Setae on prothorax: am 35, aa 20, ml 25, pa 41 – 43, epim about 40. Fore wing length 610. Sub-basal wing setae: S 1 40, S 2 30, S 3 40. Pelta length 50, width 83. Tergite IX setae: S 1 65, S 2 82 – 83. Tube length 103, maximum width 50; terminal setae about 110. Measurements (macropterous female listed under doubtful specimens from Vietnam in µm). Body length 1380 (distended). Head length 150, from anterior margin of eyes 131, width across eyes 138, maximum width across cheeks 133, minimum width across base 111; eyes length 67; diameter of posterior ocelli 12 – 13; postocular setae 45 – 50. Antenna total length 290, segments III – VIII length as follows: 40, 39, 38, 38, 32, 30. Pronotum length 85, width 161. Setae on prothorax: am 38, aa? 30, ml? 40, pa 42 – 43, epim about 50. Fore wing length 540. Sub-basal wing setae: S 1 38, S 2 45, S 3 40. Pelta length 53, width 67. Tergite IX setae: S 1 65 – 67, S 2 72 – 75. Tube length 97, maximum width 48; terminal setae 91 - 95. Specimens examined. The Philippines, 1 female, Mindanao Is., Mt. Apo, Agko, alt about 1300 m, on dead branches, 30. vii. 1979, SO. Doubtful specimens (these specimens may well be identified to Bamboosiella rugata). Indonesia, Java, 3 females and 5 males, Banyuwangi, Malan Sari, on bamboo, 27. viii. 2005, SO. Peninsular Malaysia, 1 female, about 20 km N from Kuala Lumpur, Templer Park, on grass, 15. viii. 1990, TN & SO. Vietnam, Lam Dong Province, Bao Loc, Dam Bri, on bamboo, 2 females and 1 male, 26. xii. 2001, 2 females and 1 male, 27. xii. 2001, 1 female, 28. xii. 2001, SO.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812412506B64C5F9BDA8DAFC30.taxon	description	Type-species. Squamothrips crassus sp. nov.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812412506B64C5F9BDA8DAFC30.taxon	discussion	Comments. This genus is somewhat similar in appearance to some genera included in the Liothrips - lineage, e. g. Eurhynchothrips and Psephenothrips, in having the body slightly thickened, maxillary stylets long and close together and the prosternal basantra absent. But Squamothrips probably belongs in the Phlaeothrips - lineage, because it has two sense cones on both antennal segments III and IV. Moreover, it is characterized by having the anterior pair of wing-retaining setae on abdominal tergite VIII. Usually, the wing-retaining setae are present on tergites II to VII and only rarely on tergite VIII in Phlaeothripinae. These unusual wing-retaining setae on tergite VIII are found only in a few genera, such as Andrethrips, Lizalothrips, Phylladothrips, Propesolomonthrips and Solomonthrips, and usually two pairs are present, but two Lizalothrips species and Phylladothrips bispinosus have only the anterior pair of these setae. In addition, one Oriental Terthrothrips species, T. strasseni, also has the anterior pair on tergite VIII. However, Squamothrips does not appear to have a close relationship with any of these genera, because there are no important features in common with these genera other than wing-retaining setae. The distinctly reticulate body and distinctly dilated stout body setae of this genus are somewhat similar to those of Azaleothrips and its relatives (Okajima & Masumoto 2014). Moreover, Azaleothrips similarly has both right and left wing-retaining setae situated rather close together (cf. Fig. 221) and often has short and simply curved anterior pair of wing-retaining setae on tergite VIII. These two genera, Squamothrips and Azaleothrips, may have a common ancestor. From Azaleothrips, it can easily be distinguished by the postocular setae arising near the cheeks. The genus name is derived from the Latin ‘ squama ’ meaning scales. The type-species has scale-like sculptures on the surface of the body. Gender: masculine.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812413506464C5FBC1A853FE54.taxon	description	(Figs 212 – 221) Female (macroptera). Distended body length: 1.38 – 1.52 mm. Body brown (Fig. 212). Legs largely yellow, fore and mid femora scarcely shaded with brown. Antennal segments I – II and VII – VIII brown, but segment II paler antero-externally; segments III – VI yellow, segment VI weakly shaded with brown distally. Fore wings and major body setae pale. Head almost as long as wide (Fig. 213); mid-dorsal setae well-developed, rather stout, 16 – 20 µm length, blunt or weakly pointed (left one blunt, but right one weakly expanded in holotype). Cheeks very weakly rounded, with several stout setae, blunt or weakly expanded. Postocular setae shorter than eyes, 112 µm apart from each other, 18 – 20 µm apart from eyes in holotype. Eyes a little longer than one-third of head length, ommatidia slightly separated from each other. Ocelli relatively small; posterior pair slightly separated from eyes, 29 µm apart from each other in holotype. Antennal segment III weakly swollen sub-basally; segment VII the longest (Fig. 220). Mouth cone reaching a level of ferna. Pronotum (Fig. 213) about 0.9 times as long as head, 1.62 times as wide as long in holotype; almost entirely reticulate, but reticle weaker than that on head, with a few discal setae that are somewhat stout, blunt, often weakly expanded, median longitudinal apodeme absent. Ferna rather small (Fig. 214), wider apart from one another; prospinasternum about 75 µm wide. Metanotum strongly sculptured with polygonal reticulation (Fig. 215), median pair of setae about 25 µm long, 56 µm apart from each other, about 70 µm apart from anterior margin of metanotum in holotype. Dorsal surface of fore femur with more than 10 stout setae, blunt or very weakly expanded. Fore wing with 6 – 7 duplicated cilia in holotype. Tergite IX S 1 setae a little longer than S 2. Tube (Fig. 218) about 0.7 times as long as head. Measurements (holotype female in µm). Body length 1520 (distended). Head length 145, from anterior margin of eyes 140, width across eyes 138, maximum width across cheeks 146, minimum width across base 135; eyes length 52; diameter of posterior ocelli 9 – 12; postocular setae 36 – 38. Antenna total length about 300, segments III – VIII length (width) as follows: 40 (25), 39 (27), 37 (25), 38 (23), 43 (18), 27 (10). Pronotum length 130, width 210. Setae on prothorax: am 25, aa? 25, ml?, pa 35, epim about 50. Fore wing length 520. Sub-basal wing setae: S 1 25, S 2 25, S 3 35. Pelta length 55, width 55. Tergite IX setae: S 1 66 – 70, S 2 56 – 60. Tube length 103, maximum width 48; terminal setae 83 – 90. Male. Unknown.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812413506464C5FBC1A853FE54.taxon	materials_examined	Type material. Holotype: macropterous female, Thailand, nr. Chiang Mai, Pha Sang, on grass, 31. viii. 1992. TN & SO. Paratypes: Thailand, nr. Chiang Mai, 1 female, foot of Doi Saket, 25. viii. 1992, TN & SO, 1 female, Ka Jan, on bamboo, 3. ix. 1992, SO, 1 female, Pha Hean, 3. ix. 1992, SO.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812413506464C5FBC1A853FE54.taxon	discussion	Comments. This species is described from near Chiang Mai, northern Thailand, based on four females collected on different occasions. Two of the females were collected from bamboo and grass respectively, but the habitat of the remaining two females is uncertain.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781241D506564C5FF65AB8CFD10.taxon	description	(Figs 61, 71 – 74, 81 – 82) From Veerabahuthrips, this genus can be distinguished by the following characteristics: head not protruded in front of compound eyes (Figs 71 & 72) (distinctly protruded in Veerabahuthrips); cheeks without constriction behind eyes (Figs 71 & 72) (with distinct constriction in Veerabahuthrips); mesonotum with median cleft incomplete, scarcely reaching middle (Figs 73 & 74) (complete or nearly complete in Veerabahuthrips, cf. Figs 69 & 70). Type-species. Takeazamiuma exilis (Okajima) comb. nov.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781241D506564C5FF65AB8CFD10.taxon	discussion	Comments. This genus is closely related to Veerabahuthrips, but probably also related to Mychiothrips. The relationship between these genera is discussed in detail under Mychiothrips above. Two species are transferred from Veerabahuthrips to this new genus: Takeazamiuma exilis (Okajima) comb. nov. and Takeazamiuma simplex (Okajima) comb. nov. These two species can easily be distinguished from each other by the key couplet 2 in Okajima (1993 b, P. 728). Unfortunately, little is known about these thrips other than that they live on bamboo. There are no new findings to be revised for these two species. The genus name is derived from two Japanese words, ‘ take ’ and ‘ azamiuma ’ meaning ‘ bamboo’ and ‘ thrips’ respectively. Gender: feminine.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781241D506564C5FCEDAEF0F9A8.taxon	description	Type-species. Thaithrips unicolor sp. nov.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781241D506564C5FCEDAEF0F9A8.taxon	discussion	Comments. Although this new genus has the fore wings not distinctly constricted medially, it presumably belongs to the tribe Haplothripini. However, the generic relationships of Thaithrips among the Haplothripini is not clear, as the structure of the compound eyes which have several large colored ommatidia on the ventral and ventro-lateral surface, may be unusual in the tribe. Similar ommatidia are found commonly in the family Thripidae and subfamily Idolothripinae of the family Phlaeothripidae, but their function is uncertain. This genus may be related to Vietothrips gen. nov. described below, and the distinction between them is discussed under Vietothrips. The genus name is derived from the type locality ‘ Thai’ or ‘ Thailand’. Gender: masculine.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781241D506664C5F935A951FAD0.taxon	description	(Figs 222 – 230) Female (macroptera). Distended body length: 1.8 – 1.9 mm. Body uniformly brown (Fig. 222), tube somewhat darker. Femora brown, tibiae brown with anterior half yellowish, tarsi yellow. Antennal segments I – II and VI – VIII brown, almost concolorous with head, segment III pale brown with outer portion slightly yellowish, with yellow basal neck, segment IV and V pale brown with basal one-third very slightly yellowish. Fore wings very slightly brownish medially. Prominent setae pale. Head elongate (Fig. 223), about 1.7 times as long as width across cheeks; dorsal surface almost smooth. Cheeks weakly rounded, with no remarkable setae. Postocular setae much shorter than eyes, weakly expanded. Eyes about 0.4 times as long as head, colored large ommatidia on ventro-lateral surface almost as large as posterior ocelli. Posterior ocelli 13 – 15 µm in diameter, 27 µm apart from each other, 20 – 22 µm apart from anterior one in holotype. Antennae (Fig. 229) about 1.3 times as long as head; intermediate segments subequal in length; segment VIII scarcely constricted at base. Maxillary stylets scarcely reaching middle of head, about half of head width apart, maxillary bridge 47 µm width in holotype. Pronotum (Fig. 223) 0.63 times as long as head and 1.12 times as wide as long in holotype; surface almost smooth, only with weak transverse reticulation near posterior margin; aa and pa short and weakly expanded; epim longer than pa. Mesonotum weakly sculptured, with two pairs of campaniform sensilla. Metanotum with weak reticulation (Fig. 225); median pair of setae short and pointed, situated at anterior one-third. Fore tarsal tooth directed latero-posteriorly. Fore wings very weakly narrowed distally (Fig. 228); three sub-basal setae of fore wings short, S 1 and S 2 expanded, S 3 sharply pointed. Pelta bell-shaped (Fig. 227), with weak reticulation, with a pair of campaniform sensilla. Abdominal sternites III – VII with transverse row of about 10 setae. Tergites II – VIII S 2 setae: II reduced, III – VI and VIII expanded, VII long and sharply pointed. Tergite IX S 1 and S 2 setae shorter than tube, S 1 short and expanded, S 2 long and pointed; intermediate setae between S 1 and S 2 long, almost as long as S 1, or a little shorter. Tube about 0.6 times as long as head, 2.4 times as long as width. Terminal setae about 1.3 times as long as tube. Measurements (holotype female in µm). Body length about 1800 (distended). Head length 220, from anterior margin of eyes 205, width 120; eyes length 80, width 42 – 43; postocular setae less than 25. Antenna total length 285, segments I – VIII length (width) as follows: 33 (28), 40 (27), 37 (22), 37 (24), 37 (25), 38 (21), 37 (18), 30 (12). Pronotum length 138, width 155. Setae on prothorax: aa less than 15, pa about 20, epim 35 – 40. Fore wing length 670. Sub-basal wing setae: S 3 25 – 30. Tergite IX setae: S 1 36 – 38, S 2 115. Tube length 120, maximum width 50; terminal setae 150 – 155. Male. Unknown.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781241D506664C5F935A951FAD0.taxon	materials_examined	Type material. Holotype: macropterous female, Thailand, Bangkok, Bangkhen, Campus of Kasetsert University, NBCRC, on bamboo, 11. i. 1988, SO. Paratypes: Thailand, 1 female collected with holotype; 1 female, Phuket, Gu gu suri Road, on bamboo, 24. viii. 1991, TN & SO. Non-paratypic specimen. Indonesia, Java, Malang, Batu, Pujon, on bamboo, 22. VIII. 2005, SO.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781241D506664C5F935A951FAD0.taxon	discussion	Comments. This species is described from Thailand and Java, Indonesia, and is known from four females all from bamboo. The holotype female was taken from bamboo together with numerous individuals of Podothrips bicolor and Okajimathrips kentingensis. The non-paratypic female from Java is very similar to the type series, but has several small differences as follows: intermediate antennal segment slightly darker; postocular setae longer; prosternal basantra rather small; tergite IX S 1 setae longer.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781241E506764C5FAACAE21F863.taxon	description	Type-species. Vietothrips bicolor sp. nov.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781241E506764C5FAACAE21F863.taxon	discussion	Comments. The distinct prosternal basantra and maxillary bridge of this genus are shared with genera belonging to the Haplothripini. Moreover, the slender pseudovirga of the male genitalia is similar to those of Haplothrips species. Vietothrips is possibly related to Thaithrips, and they share compound eyes with several colored large ommatidia on ventral and ventro-lateral surface. From Thaithrips, however, it can be distinguished by the following features: body setae longer; antennal segment IV with three sense cones; eyes and ocelli rather close together; prothoracic midlateral setae well-developed; metanotum sculptured with closely spaced longitudinal lines or reticles; fore tarsus unarmed. The genus name is derived from the type locality ‘ Viêt Nam’. Gender: masculine.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812418506064C5FF65AB4BF93C.taxon	description	(Figs 231 – 240) Female (macroptera). Distended body length: 1.5 – 1.8 mm. Body sharply bicolored brown and yellow (Fig. 231); head, thorax, abdominal segments VIII – IX and tube brown; abdominal segments I – VII yellow. Legs yellow, but fore and mid coxae brown. Antennal segments I and II brown, the remaining segments yellow, segment VIII very weakly shaded with brown. Fore wings weakly shaded with brown, somewhat paler sub-basally. Postocular and prothoracic prominent setae a little shaded, the remaining setae almost hyaline. Head about 2.0 times as long as wide (Fig. 232), widest across eyes; dorsal surface weakly sculptured with transverse reticles. Cheeks straight, with no remarkable setae. Postocular setae shorter than eyes, weakly expanded. Eyes shorter than 0.4 times as long as head, large colored ommatidia on ventro-lateral surface larger than posterior ocelli. Posterior ocelli 15 µm in diameter, about 10 µm apart from each other, 12 – 13 µm apart from anterior one in holotype. Antennae (Fig. 239) about 1.1 times as long as head; segment VIII weakly constricted at base. Maxillary stylets about half of head width apart, maxillary bridge 43 µm width in holotype. Pronotum short (Fig. 232), 0.41 times as long as head and 1.47 times as wide as long in holotype, sculptured laterally and posteriorly; aa, ml, pa and epim expanded; aa and ml subequal in length, shorter than postocular setae; pa and epim subequal in length, almost as long as postocular setae; cox long and sharply pointed. Mesonotum distinctly sculptured, with two pairs of campaniform sensilla. Metanotal median pair of setae minute. Three sub-basal setae of fore wing well developed and expanded. Pelta triangular (Fig. 236), very weakly sculptured, with a pair of campaniform sensilla. Abdominal sternites III – VII with transverse row of 7 – 10 setae. Tergites II – VIII S 2 setae: II reduced, III expanded, IV – VIII long and sharply pointed. Tergite IX S 1 setae much shorter than tube, expanded, S 2 a little shorter than tube, sharply pointed. Tube shorter than 0.6 times as long as head, 2.4 – 2.5 times as long as width. Terminal setae almost as long as tube or a little longer. Measurements (holotype female in μm). Body length about 1700 (distended). Head length 225, from anterior margin of eyes 202, width across cheeks 105, across eyes 110; eyes length 74 – 75, width 40 – 41; postocular setae 50 – 52. Antenna total length 250, segments I – VIII length (width) as follows: 27 (27), 40 (25), 35 (21), 33 (25), 35 (22), 31 (21), 32 (18), 25 (12). Pronotum length 92, width 135. Setae on prothorax: aa 35 – 40, ml 35 – 40, pa 50 – 55, epim 50, cox 70 – 80. Fore wing length 670. Sub-basal wing setae: S 1 30 – 35, S 2 25 – 30, S 3 47 – 50. Tergite IX setae: S 1 55 – 60, S 2 115. Tube length 123, maximum width 50; terminal setae 130. Male (macroptera). Distended body length: 1.2 – 1.4 mm. Color and structure very similar to female. Antennae about 1.2 times as long as head. Fore tarsal inner hamus very small or absent. S 2 setae on abdominal tergite III expanded; tergite IX S 2 setae much shorter than S 1. Tube 0.50 – 0.55 as long as head, about 2.7 times as long as wide. Measurements (paratype male in μm). Body length about 1400 (distended). Head length 203, from anterior margin of eyes 185, width across cheeks 99, across eyes 103; eyes length 70, width 38 – 40; postocular setae 35 – 37. Antenna total length 250, segments I – VIII length (width) as follows: 25 (25), 35 (25), 30 (20), 28 (22), 35 (20), 35 (20), 32 (18), 25 (12). Pronotum length 90, width 123. Setae on prothorax: aa less than 20, ml less than 20, pa about 40, epim about 40, cox 45 – 55. Fore wing length 600. Sub-basal wing setae: S 1 20 – 25, S 2 25 – 28, S 3 40 – 43. Tergite IX setae: S 1 40, S 2 about 20. Tube length 108, maximum width 40; terminal setae 115.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812418506064C5FF65AB4BF93C.taxon	materials_examined	Type material. Holotype: macropterous female, Vietnam, Lam Dong Province, Bao Loc, Dam Bri, on bamboo, 26. xii. 2001, SO. Paratypes: Vietnam, 5 females, collected with holotype; data very similar to holotype, 9 females and 2 males, 27. xii. 2001, 23 females and 13 males, 28. xii. 2001.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812418506064C5FF65AB4BF93C.taxon	discussion	Comments. This species is described from Vietnam based on a good number of females and males collected from bamboo together with Namothrips bambusae described above. It is interesting that both of them have bicolored bodies, but their color patterns are somewhat different. Unfortunately, no biological information about them is known so far.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812418506264C5F8C1AFE7FB47.taxon	description	Type-species. Yurikoazamiuma bidens sp. nov.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E287812418506264C5F8C1AFE7FB47.taxon	discussion	Comments. This genus is included in the Haplothripini, and closely related to Okajimathrips, Podothrips and Praepodothrips, and it is probably most closely related to Podothrips in having similar formed maxillary stylets and bridge. However, unlike Podothrips, Yurikoazamiuma has the basantra smaller and located far from ferna on the prosternum (Fig. 243). These four genera share the somewhat flattened body, metathoracic sternopleural sutures well-developed (cf. Fig. 247) and metafurcal arms separated from each other (cf. Figs 248, 251 & 252), and can be discriminated from each other by the following key. The flattened body with relatively short legs of these genera are probably an adaptation to living under the leaf sheaths of bamboo and / or grass, where they may be predaceous on certain coccids that also live there. The genus name is dedicated to the late Yuriko Okajima. She was the wife of the first author and was well understanding of her husband’s research on thrips during her lifetime. She always spared no effort to support him behind the scenes. The Japanese word ‘ azamiuma’ means ‘ thrips’. Gender: feminine.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781241A501C64C5FB1FAEDDFE0C.taxon	description	(Figs 241 – 250, 257) Female (macroptera). Distended body length: about 1.7 mm. Body bicolored (Fig. 241). Head, prothorax, mesothorax and tube brown, tube with basal one-third yellowish; metathorax and abdominal segments I – IX largely yellowish, abdominal segments scarcely and partly shaded with pale brown. Fore femora brown, with apical half yellowish; the rest of legs yellow. Antennal segments I and VII – VIII brown; segments II – IV yellow, but IV shaded with pale brown at apex; segment V yellowish at basal half, pale brown at apical half; segment VI pale brown, a little paler than VII. Fore wings and major body setae pale. Head 1.23 times as long as wide (Fig. 242), widest across cheeks at a level of postocular setae, dorsal surface almost smooth, but with a few transverse wrinkles between postocular setae. Cheeks rounded, almost smooth. Postocular setae shorter than eyes, blunt or pointed, 142 µm apart from each other, 19 – 23 µm apart from eyes. Eyes about one-third of head length, about 70 µm apart from each other. Ocelli relatively small; posterior pair in contact with eyes, 48 µm apart from each other. Antennae (Fig. 249 & 250) about 1.5 times as long as head; segment VIII distinctly constricted at base, 0.76 times as long as segment VII; segment III a little longer than segment IV. Maxillary stylets reaching eyes, 21 µm apart from each other at middle; maxillary bridge distinct, a little narrower than one-fourth of head width, situated at the middle of head. Pronotum (Fig. 242) well-developed, 0.72 times as long as head, 1.31 times as wide as long, almost smooth; four pairs of major prothoracic setae developed, weakly expanded or blunt; ml longer than aa, pa a little shorter than ml. Prospinasternum rather small (Fig. 243). Mesonotum almost smooth, lateral setae short and pointed, about 15 µm long. Metanotum almost smooth (Fig. 245); median pair of setae short and acute, 35 µm apart from each other, 42 – 46 µm apart from anterior margin of metanotum. Fore wing (Fig. 257) with 4 duplicated cilia; sub-basal setae S 1 and S 2 weakly expanded, S 3 much longer than S 2, finely pointed. Pelta trapezoid (Fig. 246), sculptured weakly, about 1.1 times as wide as long. Tergites II – VIII S 1 setae weakly expanded. Tergites III – VIII S 2 setae: III – VI weakly expanded, VII and VIII pointed. Tergite IX S 1 setae very weakly expanded, much shorter than tube, S 2 setae on IX sharply pointed, almost as long as tube; intermediate setae about two-thirds of S 1. Tube about 0.5 times as long as head, about 2.0 times as long as wide; terminal setae 1.3 – 1.4 times as long as tube. Measurements (holotype female in µm). Body length 1720 (distended). Head length 200, from anterior margin of eyes 180, width across eyes 150, maximum width across cheeks 162, minimum width across base 135; eyes length 65; diameter of posterior ocelli 10 – 12; postocular setae 48 – 52. Maxillary bridge width 37. Antenna total length about 300, segments III – VIII length (width) as follows: 43 (27), 41 (27), 36 (25), 33 (23), 37 (20), 28 (13). Pronotum length 144, width 188. Setae on prothorax: aa 30 – 35, ml about 50, pa 45 – 48, epim 53 – 55. Fore wing length 640. Sub-basal wing setae: S 1 30 – 32, S 2 25 – 31, S 3 48 – 66. Pelta length 69, width 76. Tergite IX setae: S 1 60 – 61, S 2 100 – 103. Tube length 102, maximum width 50; terminal setae 135 – 142. Male. Unknown.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781241A501C64C5FB1FAEDDFE0C.taxon	materials_examined	Type material. Holotype: macropterous female, Thailand, nr. Chiang Mai, foot of Doi Suthep, on grass, 24. viii. 1992. SO.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
03E28781241A501C64C5FB1FAEDDFE0C.taxon	discussion	Comments. This species is described from Thailand based on a unique holotype female collected from grass together with Praepodothrips indicus. The fore tarsal armature of this taxon is characteristic.	en	Okajima, Shûji, Masumoto, Masami (2025): Some Poaceae-inhabiting Phlaeothripinae (Thysanoptera, Phlaeothripidae) in East Asia, with descriptions of six new genera and 18 new species. Zootaxa 5726 (1): 1-68, DOI: 10.11646/zootaxa.5726.1.1, URL: https://doi.org/10.11646/zootaxa.5726.1.1
