identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03EF1D07A713E71C5D25170D2E67FA5C.text	03EF1D07A713E71C5D25170D2E67FA5C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quadriacanthus aegypticus El-Naggar & Serag 1986	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Quadriacanthus aegypticus El-Naggar &amp; Serag, 1986</p>
            <p> Type-host and locality:  Clarias gariepinus ; Lake Manzala and the Demietta branch of the Nile River, Dakahlia Governorate, Egypt (El-Naggar and Serag 1986). </p>
            <p>Other localities: Ammar drainage canal, Bahr Mouis, Damroo drainage canal, Mansoura drainage canal, Nawasa El-Gheit drainage canal, Telbanah drainage canal (Egypt), Blue Nile, White Nile (Sudan), Krugerdrift Dam, Gariep Dam, Lake Tzaneen, Middle Letaba Dam, Nwanedi-Luphephe Dams, Phalaborwa Barrage, Vaal Dam (South Africa), Lake Tana (Ethiopia), Lake Turkana (Kenya) and Lake Kariba (Zimbabwe) (Truter 2022, Truter et al. 2023).</p>
            <p> New host:  Clarias ngamensis . </p>
            <p>New localities: Lake Lubanda, Luapula River.</p>
            <p>Prevalence: P = 6.7% (Lake Lubanda); P = 40% (Luapula River).</p>
            <p>Mean intensity: MI = 1 ± 1 (Lake Lubanda); MI = 2.7 ± 3.3 (Luapula River).</p>
            <p>Voucher specimens: 17 Voucher specimens XXII.3.”06; 15; 18; 23; 25; 26; 28; 29; 31; 35; 37–43” are deposited in the collection of the Research Group Zoology: Biodiversity &amp; Toxicology, at Hasselt University (Diepenbeek, Belgium).</p>
            <p>Diagnosis: Tubular and straight male copulatory organ (MCO), widest at base and narrowing towards distal extremity. Accessory piece ends in two distinctive lateral, club-shaped outgrowths projecting from its posterior half. Vagina partly sclerotized tube. Dorsal anchor without shaft nor guard, with broad base and short point. Dorsal bar with rectangular centre, funnel-like median process posteriorly directed and two lateral expansions. Large dorsal cuneus triangular. Ventral anchor without shaft nor guard, with regularly curved blade. Ventral bar with two lateral branches. Y-shaped ventral cuneus. Seven pairs of hooks: pairs IV, III, I (decreasing size) larger than pairs II, V, VI and VII, latter pairs almost equal (Fig. 3).</p>
            <p> Remarks: The specimens are reminiscent of the morphological characters of the MCO of the original description of  Q. aegypticus by El-Naggar and Serag (1986) (base of copulatory tube wider, anterior end pointed; accessory piece terminates in two hooks and possesses two distinctive, lateral club-shaped outgrowths, projecting from the posterior half) as well as the drawings/measurements provided by El-Naggar and Serag (1986), Kritsky and Kulo (1988), Douëllou and Chishawa (1995) and Francová et al. (2017). We consider </p>
            <p> AP them conspecific, as the morphometrical characteristics of the sclerotized structures correspond to those reported for  Q. aegypticus , i.e., AP (34–44) in this study vs (33–49), (36–44), (39–44) Kritsky and Kulo (1988), Douëllou and Chishawa (1995) and Francová et al. (2017) respectively; MCO (36–39) in this study vs (34–40), (36–40) following Douëllou and Chishawa (1995) and Francová et al. (2017). However the shafts of the ventral and dorsal anchors deviate from the original description outlined in El-Naggar and Serag (1986). </p>
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	https://treatment.plazi.org/id/03EF1D07A713E71C5D25170D2E67FA5C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Kasembele, Gyrhaiss K.;Vanhove, Maarten P. M.;Manda, Auguste Chocha;Jorissen, Michiel W. P.;Luus-Powell, Wilmien J.;Smit, Willem J.;Bilong Bilong, Charles F.;Bahanak, Dieu-Ne-Dort	Kasembele, Gyrhaiss K., Vanhove, Maarten P. M., Manda, Auguste Chocha, Jorissen, Michiel W. P., Luus-Powell, Wilmien J., Smit, Willem J., Bilong Bilong, Charles F., Bahanak, Dieu-Ne-Dort (2024): Diversity of Quadriacanthus (Monogenea: Dactylogyridae) in the Upper Congo Basin: new geographical records and description of five new species from the gills of Clarias ngamensis (Siluriformes: Clariidae). Zoologia (e 23090) 41: 1-19, DOI: 10.1590/S1984-4689.v41.e23090, URL: https://doi.org/10.1590/s1984-4689.v41.e23090
03EF1D07A715E71F5E9214EB2CFFFD6F.text	03EF1D07A715E71F5E9214EB2CFFFD6F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quadriacanthus allobychowskiella Paperna 1979	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Quadriacanthus allobychowskiella Paperna, 1979</p>
            <p> Type-host and locality:  C. gariepinus ; Uganda, Lake George (Paperna 1979). </p>
            <p>Other localities: Nile River basin: Ammar drainage canal, Damietta branch, Lake Manzala, Mansoura drainage canal, Nawasa El-Gheit drainage canal, Nile River barrage, Telbanah drainage canal (Egypt), and Middle Letaba Dam (South Africa) (Truter 2022, Truter et al. 2023).</p>
            <p> New host:  Clarias ngamensis . </p>
            <p>New localities: Lake Lubanda; Luapula River.</p>
            <p>Prevalence: P = 40% (Lake Lubanda); P = 60% (Luapula River).</p>
            <p>Mean intensity: MI = 1.5 ± 1.9 (Lake Lubanda); MI= 1.7 ± 1.1 (Luapula River).</p>
            <p>Voucher specimens: 24 voucher specimens XXII.3.”04; 05; 07–14; 16; 17; 19–22; 24; 27; 30; 32–34; 36;44”are deposited in the collection of the Research Group Zoology: Biodiversity &amp; Toxicology, at Hasselt University (Diepenbeek, Belgium).</p>
            <p>Diagnosis: Tube-shaped MCO wider at its base and gradually shrinking towards distal extremity. Accessory piece possessing two lateral outgrowths arising from outer surface of middle region of sclerite and ending with well-developed hook. Vagina relatively long tube, partly sclerotized duct wide at the basal extremity.Dorsal anchor without shaft nor guard, with broad base and short point. Dorsal bar with rectangular centre, small median process posteriorly directed and two lateral expansions. Large dorsal cuneus with bifid end. Ventral anchor regularly curved without shaft nor guard. Ventral bar V-shaped with two lateral branches. Y-shaped ventral cuneus. Seven pairs of hooks: pairs IV, III, I (decreasing size), bigger than pairs II, V-VII almost equal (Fig. 4).</p>
            <p> Remarks: This species,  Q. allobychowskiella (synonyms:  Quadriacanthus clariadis allobychowskiella Paperna, 1979 ;  Quadriacanthus kearni El-Naggar &amp; Serag, 1985 ) could be mixed up with  Q. clariadis Paperna, 1961 by the morphology of the ventral anchors with a curved shaft and long point, the dorsal anchor with an elongated bent shaft and short point, and the Y-shaped ventral cunei. It is however easily distinguishable as  Q. allobyschowskiella by possessing a large dorsal cuneus with a bifid end vs a triangular one for  Q. clariadis (Kritsky and Kulo 1988, Francová et al. 2017, Truter 2022, Truter et al. 2023). </p>
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	https://treatment.plazi.org/id/03EF1D07A715E71F5E9214EB2CFFFD6F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Kasembele, Gyrhaiss K.;Vanhove, Maarten P. M.;Manda, Auguste Chocha;Jorissen, Michiel W. P.;Luus-Powell, Wilmien J.;Smit, Willem J.;Bilong Bilong, Charles F.;Bahanak, Dieu-Ne-Dort	Kasembele, Gyrhaiss K., Vanhove, Maarten P. M., Manda, Auguste Chocha, Jorissen, Michiel W. P., Luus-Powell, Wilmien J., Smit, Willem J., Bilong Bilong, Charles F., Bahanak, Dieu-Ne-Dort (2024): Diversity of Quadriacanthus (Monogenea: Dactylogyridae) in the Upper Congo Basin: new geographical records and description of five new species from the gills of Clarias ngamensis (Siluriformes: Clariidae). Zoologia (e 23090) 41: 1-19, DOI: 10.1590/S1984-4689.v41.e23090, URL: https://doi.org/10.1590/s1984-4689.v41.e23090
03EF1D07A716E71E5D3611E82FF8FA9B.text	03EF1D07A716E71E5D3611E82FF8FA9B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quadriacanthus amakaliae Kasembele, Bahanak & Vanhove 2024	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Quadriacanthus amakaliae Kasembele, Bahanak &amp; Vanhove ,  sp. nov.</p>
            <p>https://zoobank.org/ 83453616-133F-4558-84E3-0EC86C687CC3</p>
            <p>
                 Type-host and locality:  C. ngamensis ;   DR Congo,  
                <a title="Search Plazi for locations around (long 27.916666/lat -11.066667)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=27.916666&amp;materialsCitation.latitude=-11.066667">Haut-Katanga</a>
                 , Lubanda village, Lake Lubanda 11°04’S, 27°55’E, November 2021, G.K. Kasembele leg. 
            </p>
            <p>Other locality: Luapula River.</p>
            <p>Prevalence: P = 6.7% (Lake Lubanda); P = 6.7% (Luapula River).</p>
            <p>Mean intensity: Lake Lubanda: 3 ± 0; Luapula River:1 ± 0.</p>
            <p>  Type-material: The holotype HU #888 and three paratypes HU #889; 890; 987 are deposited in the collection of the Research Group Zoology: Biodiversity &amp; Toxicology, at Hasselt University (  Diepenbeek , Belgium)  . </p>
            <p>Etymology: This species is named in honour of Annette Megameno Amakali, MSc, of the National Marine Information and Research Centre (NatMIRC), Ministry of Fisheries and Marine Resources, Swakopmund, Namibia, for her contribution in lab work.</p>
            <p>Description: Tube-shaped MCO (thick-walled) wide at its base and gradually shrinking towards the distal extremity. Accessory piece articulating with MCO, ending in point and having two filaments on either side of the terminal point. Funnel shaped tubular vagina. Dorsal bar with rectangular centre, median process posteriorly directed and two lateral expansions. Dorsal anchor without shaft nor guard, with broad base, shaft sharply curved, ending with short point. Dorsal cuneus triangular. Ventral bar V-shaped with two lateral branches. Ventral anchor without shaft nor guard, with base smaller than that of dorsal anchor, and curved blade. Ventral cuneus smaller than dorsal one. Seven pairs of hooks: pair IV bigger than the rest which are almost equal in length (Fig. 5).</p>
            <p> Remarks:  Quadriacanthus amakaliae sp. nov. is comparable to  Q. barombiensis Bahanak, Nack &amp; Pariselle, 2022 ,  Q. levequei Birgi, 1988 and  Q. anaspidoglanii Akoumba, Pariselle &amp; Tombi, 2017 , described in Cameroon from  C. maclareni ,  C. pachynema and  Anaspidoglanis macrostoma , respectively, by the shape of the dorsal bar median process, the tubular shape of the MCO which is enlarged at the basal zone and tapered at the distal end, and the shape of the accessory piece ending in a point. However, some differences differentiate them, namely: (i) the dorsal bar postero-median process without filaments at its end vs two filaments present in  Q. barombiensis and  Q. levequei ; (ii) the morphology of the distal extremity of the accessory piece with one small hook surrounded by two filaments vs simply one small hook in  Q. barombiensis and  Q. anaspidoglanii and two small hooks in  Q. levequei ; (iii) the difference in the size of the following sclerotized parts for  Q. amakaliae sp. nov. vs  Q. barombiensis : MCO (34–37.9) vs (25–29), AP (29–33) vs (20–26). </p>
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	https://treatment.plazi.org/id/03EF1D07A716E71E5D3611E82FF8FA9B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Kasembele, Gyrhaiss K.;Vanhove, Maarten P. M.;Manda, Auguste Chocha;Jorissen, Michiel W. P.;Luus-Powell, Wilmien J.;Smit, Willem J.;Bilong Bilong, Charles F.;Bahanak, Dieu-Ne-Dort	Kasembele, Gyrhaiss K., Vanhove, Maarten P. M., Manda, Auguste Chocha, Jorissen, Michiel W. P., Luus-Powell, Wilmien J., Smit, Willem J., Bilong Bilong, Charles F., Bahanak, Dieu-Ne-Dort (2024): Diversity of Quadriacanthus (Monogenea: Dactylogyridae) in the Upper Congo Basin: new geographical records and description of five new species from the gills of Clarias ngamensis (Siluriformes: Clariidae). Zoologia (e 23090) 41: 1-19, DOI: 10.1590/S1984-4689.v41.e23090, URL: https://doi.org/10.1590/s1984-4689.v41.e23090
03EF1D07A717E71E5E9714AE28F4FA9B.text	03EF1D07A717E71E5E9714AE28F4FA9B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quadriacanthus halajiani Kasembele, Bahanak & Vanhove 2024	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Quadriacanthus halajiani Kasembele, Bahanak &amp; Vanhove ,  sp. nov.</p>
            <p>https://zoobank.org/ C86AAF04-FA01-49CD-B24E-1B3BFCB39FA5</p>
            <p>
                 Type-host and locality:  C. ngamensis ;   DR Congo,  
                <a title="Search Plazi for locations around (long 27.916666/lat -11.066667)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=27.916666&amp;materialsCitation.latitude=-11.066667">Haut-Katanga</a>
                 , Lubanda village, Lake Lubanda 11°04’S, 27°55’E, November 2021, G.K. Kasembele leg. 
            </p>
            <p>Other locality: Luapula River.</p>
            <p>Prevalence: P = 46.7% (Lake Lubanda); P = 40% (Luapula River).</p>
            <p>Mean intensity: Lake Lubanda: 1.9 ± 1.8; Luapula River: 2.7 ± 2.3</p>
            <p>  Type-material: The holotype HU #985 and 27 paratypes HU #891; 894; 896–899; 902; 903; 906–909; 918; 919; 923; 9388– 940; 950; 951; 960; 967; 973; 986; 988; 992; 999 are deposited in the collection of the Research Group Zoology: Biodiversity &amp; Toxicology, at Hasselt University (  Diepenbeek , Belgium)  . </p>
            <p>Etymology: This species is named in honour of Dr. Ali Halajian for his valuable and kind contribution during field work and practice in fish parasitology in South Africa (University of Limpopo).</p>
            <p>Description: Tube-shaped MCO. Accessory piece simple, thicker in its distal part and with bulge and depression respectively on the external and internal sides of its median part and ending in well-developed point. Vagina not ob - served. Dorsal anchor without shaft nor guard, with broad base and short point. Dorsal bar with rectangular centre, medium median process posteriorly directed and two lateral expansions. Dorsal cuneus big and triangular. Ventral anchor without shaft nor guard, with regularly curved blade. Ventral bar V-shaped with two lateral branches. Y-shaped ventral cuneus. Seven pairs of hooks: pairs IV, III, I (decreasing size) larger than pairs II, V-VII almost equal (Fig. 6).</p>
            <p> Remarks: Based on the comparative morphology of the ventral and dorsal bars, ventral and dorsal cunei,  Q. halajiani sp. nov. is comparable to  Q. agnebiensis N’Douba, Lambert &amp; Euzet, 1999 and  Q. longifilisi N’Douba, Lambert &amp; Euzet, 1999 described from  Heterobranchus isopterus and  H. longifilis , in Ivory Coast. The new species differs from the latter two species by the shape and size of the MCO, slightly curved vs straight tube in  Q. longifilisi (28.3–34.8) vs (35–42), the AP, ending in a point and possessing a bulge at its external side vs two points at the distal part of the AP of  Q. agnebiensis , with two bulges at its external side (31.7–35.7) vs (38–43), and the vagina, non-sclerotized vs sclerotized in the two other species.  Quadriacanthus halajiani sp. nov. also differs in the length of the dorsal cuneus (22–28) vs (7–10) and (7–17) from  Q. agnebiensis and  Q. longifilisi respectively. </p>
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	https://treatment.plazi.org/id/03EF1D07A717E71E5E9714AE28F4FA9B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Kasembele, Gyrhaiss K.;Vanhove, Maarten P. M.;Manda, Auguste Chocha;Jorissen, Michiel W. P.;Luus-Powell, Wilmien J.;Smit, Willem J.;Bilong Bilong, Charles F.;Bahanak, Dieu-Ne-Dort	Kasembele, Gyrhaiss K., Vanhove, Maarten P. M., Manda, Auguste Chocha, Jorissen, Michiel W. P., Luus-Powell, Wilmien J., Smit, Willem J., Bilong Bilong, Charles F., Bahanak, Dieu-Ne-Dort (2024): Diversity of Quadriacanthus (Monogenea: Dactylogyridae) in the Upper Congo Basin: new geographical records and description of five new species from the gills of Clarias ngamensis (Siluriformes: Clariidae). Zoologia (e 23090) 41: 1-19, DOI: 10.1590/S1984-4689.v41.e23090, URL: https://doi.org/10.1590/s1984-4689.v41.e23090
03EF1D07A717E7125D3C14AE2EB8F9FD.text	03EF1D07A717E7125D3C14AE2EB8F9FD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quadriacanthus domatanai Kasembele, Bahanak & Vanhove 2024	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Quadriacanthus domatanai Kasembele, Bahanak &amp; Vanhove ,  sp. nov.</p>
            <p>https://zoobank.org/ 7F290D48-6835-4D0F-9E05-7BE97306AFF2</p>
            <p>
                 Type-host and locality:  C. ngamensis ;   DR Congo,  
                <a title="Search Plazi for locations around (long 27.916666/lat -11.066667)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=27.916666&amp;materialsCitation.latitude=-11.066667">Haut-Katanga</a>
                 , Lubanda village, Lake Lubanda 11°04’S, 27°55’E, November 2021, G.K. Kasembele leg. 
            </p>
            <p>Other locality: Luapula River</p>
            <p>Prevalence: P = 13.3% (Lake Lubanda); P = 20% (Luapula River).</p>
            <p>Mean intensity: Lake Lubanda: 1.5 ± 1.6; Luapula River: 2 ± 2.1</p>
            <p>MCO</p>
            <p>  Type-material: The holotype HU n° 892 and seven paratypes HU #900; 901; 928–930; 1001; 1004 are deposited in the collection of the Research Group Zoology: Biodiversity &amp; Toxicology, at Hasselt University (  Diepenbeek , Belgium)  . </p>
            <p>Etymology: This species is named in honour to Doma Tana Anicet, MSc, of the Unité de Recherche en Biodiversité et Exploitation durable des Zones Humides, Faculté des Sciences agronomiques, Université de Lubumbashi, DR Congo, for his valuable and kind assistance during our field campaigns.</p>
            <p>Description: MCO long, wide at proximal end and tapered at distal extremity. Accessory piece long, robust and thickening in its median part, and with bulge on the external and internal faces and ending in hook. Vagina not observed. Dorsal anchor without shaft nor guard, but with curved blade ending with short point. Dorsal bar with rectangular centre, one small median process posteriorly directed and two lateral expansions. Dorsal cuneus triangular. Ventral anchor without shaft nor guard, with thin and regularly curved blade. Ventral bar V-shaped with two lateral branches. Y-shaped ventral cuneus. Seven pairs of hooks: pairs IV, III, I (decreasing size) larger than pairs II, V, VI and VII, latter pairs almost equal in length (Fig. 7).</p>
            <p> Remarks: This species is similar to  Q. ndoubai Bahanak, Pariselle &amp; Bilong Bilong, 2017 and  Q. macrocirrus N’Douba, Lambert &amp; Euzet, 1999 , described respectively on  H. longifilis and  H. isopterus , considering the morphology of their haptoral structures (the robust shape of the ventral and dorsal bars). However, they can be distinguished by the shape of the MCO (large and slightly curved vs large and straight in  Q. macrocirrus ), and the shape of the AP (large, robust, thickening in its median part and ending in a point vs simple, long and tapered in  Q. macrocirrus , and ending in a fork in  Q. ndoubai ). </p>
            <p> Quadriacanthus lubandaensis Kasembele , </p>
            <p> Bahanak &amp; Vanhove,  sp. nov.</p>
            <p>https://zoobank.org/ F914590C-3987-466B-8FB0-470B7C280F7D</p>
            <p>
                 Type-host and locality:  Clarias ngamensis ;   DR Congo,  
                <a title="Search Plazi for locations around (long 27.916666/lat -11.066667)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=27.916666&amp;materialsCitation.latitude=-11.066667">Haut-Katanga</a>
                 , Lubanda village, Lake Lubanda 11°04’S, 27°55’E, November 2021, G.K. Kasembele leg. 
            </p>
            <p>Other locality: Luapula River.</p>
            <p>Prevalence: P = 26.7% (Lake Lubanda); P = 33.3% (Luapula River).</p>
            <p>Mean intensity: Lake Lubanda: 1 ± 0.8; Luapula River: 1.8 ± 1.4.</p>
            <p>Type-material: The holotype HU #954 and 12 paratypes HU #893; 895; 904; 905; 914; 920; 934; 956; 972; 978; 990;</p>
            <p>991 are deposited in the collection of the Research Group Zoology: Biodiversity &amp; Toxicology, at Hasselt University (Diepenbeek, Belgium).</p>
            <p> Etymology: The epithet  lubandaensis refers to the type locality. </p>
            <p>Description: MCO simple and straight tube. Accessory piece articulating with copulatory tube, with hook-like ending. Vagina not observed. Dorsal anchor without shaft nor guard, with broad base, curved shaft, medium point lenght. Dorsal bar with rectangular centre, medium median process posteriorly directed and two lateral expansions. Dorsal cuneus triangular. Ventral anchor without shaft nor guard, with regular curved blade. Ventral bar V-shaped with two lateral branches. Y-shaped ventral cuneus. Seven pairs of hooks: pairs IV, I, III (decreasing size) larger than pairs II, V, VI and VII. Four latter pairs are almost equal in length (Fig. 8).</p>
            <p> Remarks: This species is comparable to  Q. fornicatus Francová &amp; Řehulková, 2017 , described from  C. gariepinus from Sudan (Francová et al. 2017) by the morphology of the ventral bar, the ventral anchors, the ventral and dorsal cunei, and the MCO (straight tube, base simple). However, these species are different in the morphology of the dorsal anchors (with long points in  Q. fornicatus 12–13 vs 5–11 in  Q. lubandaensis sp. nov. ), the MCO (with a thickened wall) and the AP (with a well developed hook at the distal end) in  Q. fornicatus vs the MCO (with thin margin) and the AP (hook-like, not well developed, less curved) in  Q. lubandaensis sp. nov.</p>
            <p> Quadriacanthus curvicirrus Kasembele, Bahanak</p>
            <p> &amp; Vanhove,  sp. nov.</p>
            <p>https://zoobank.org/ 8DCFEA56-C3A9-426F-94E4-B620B0DE2F5B</p>
            <p>
                 Type-host and locality:  Clarias ngamensis ;   DR Congo,  
                <a title="Search Plazi for locations around (long 28.616667/lat -10.4)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=28.616667&amp;materialsCitation.latitude=-10.4">Haut-Katanga</a>
                 , Kasenga, Luapula River 10°24’S, 28°37’E, November 2021, G.K. Kasembele leg. 
            </p>
            <p>Prevalence: P = 66.7% (Luapula River).</p>
            <p>Mean intensity: Luapula River: 6.3 ± 5.8.</p>
            <p>  Type-material: The holotype HU n° 984 and 62 paratypes HU n° 910–913; 915–917; 921; 922; 924–927; 931–933; 935–937; 941–949; 952; 953; 955; 957 –959; 961–966; 968–971; 974–976; 979–984; 989; 993–998; 1000; 1002; 1003 are deposited in the collection of the Research Group Zoology: Biodiversity &amp; Toxicology, at Hasselt University (  Diepenbeek , Belgium)  . </p>
            <p> Etymology: the epithet  curvicirrus refers to the curved form of the male copulatory tube. </p>
            <p>AP</p>
            <p>Description: Tubular MCO, with simple base, curved at its distal end, hook shaped, with a swollen portion just before the curvy part. Accessory piece massive with outgrowths posteriorly and anteriorly, with the distal part smaller, ending in a point. Dorsal anchor without shaft nor guard, with broad base, shaft sharply bent, short point. Dorsal bar with rectangular centre, broad and long median process posteriorly directed and two lateral expansions. Dorsal cuneus triangular. Ventral anchor without shaft nor guard, with narrow base and curved blade. Ventral bar V-shaped with two lateral branches. Ventral cuneus triangular. Seven pairs of hooks: pairs IV larger than pairs I, II, III, V, VI and VII. Four latter pairs are almost equal in length (Fig. 9). Vagina not observed.</p>
            <p> Remarks:  Quadriacanthus curvicirrus sp. nov. is similar to  Q. barombiensis in the shape of the ventral and dorsal anchors and cunei. They can be distinguished by the median process of the dorsal bar without filaments at its end in  Q. curvicirrus sp. nov. vs with two filaments in  Q. barombiensis ; with respective median process length h= (41.1–50.2) vs (27–32). The MCO of  Q. curvicirrus sp. nov. is characteristic, tubular, curved at its distal end with a simple base and the accessory piece massive with outgrowths posteriorly and anteriorly directed. This differs from the situation in  Q. barombiensis with a tubular straight MCO and a simple, slightly S-shaped accessory piece, ending in one small point. </p>
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	https://treatment.plazi.org/id/03EF1D07A717E7125D3C14AE2EB8F9FD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Kasembele, Gyrhaiss K.;Vanhove, Maarten P. M.;Manda, Auguste Chocha;Jorissen, Michiel W. P.;Luus-Powell, Wilmien J.;Smit, Willem J.;Bilong Bilong, Charles F.;Bahanak, Dieu-Ne-Dort	Kasembele, Gyrhaiss K., Vanhove, Maarten P. M., Manda, Auguste Chocha, Jorissen, Michiel W. P., Luus-Powell, Wilmien J., Smit, Willem J., Bilong Bilong, Charles F., Bahanak, Dieu-Ne-Dort (2024): Diversity of Quadriacanthus (Monogenea: Dactylogyridae) in the Upper Congo Basin: new geographical records and description of five new species from the gills of Clarias ngamensis (Siluriformes: Clariidae). Zoologia (e 23090) 41: 1-19, DOI: 10.1590/S1984-4689.v41.e23090, URL: https://doi.org/10.1590/s1984-4689.v41.e23090
