taxonID	type	description	language	source
038187C1FFBA5D2AFF52FCE1A3ADF834.taxon	type_taxon	Type genus: Stephanacris Redtenbacher, 1908: 441.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFBA5D2AFF52FCE1A3ADF834.taxon	description	Hennemann & Conle, 2004: 48. Otte & Brock, 2005: 33. Hennemann & Conle, 2008: 54, Figs. 37, 43, 60 - 61, 80 - 82. Pharnaciini Günther, 1953: 555 (in part). Bradley & Galil, 1977: 193 (in part). Otte & Brock, 2005: 32 (in part).	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFBA5D2AFF52FCE1A3ADF834.taxon	discussion	Comments. Stephanacridini was established by Günther (1953: 555) as a tribe of the subfamily Phasminae (= Phasmatinae Bradley & Galil, 1977) and originally only contained the type-genus Stephanacris Redtenbacher, 1908. The fundamental differences of the morphology of the terminalia and eggs of Stephanacridini and Phasmatinae genera, that suggest the tribe cannot be a member of Phasmatidae s. str. (= Lanceocercata), were summarized and illustrated by Hennemann & Conle (2008: 55), who transferred several misplaced genera from Pharnaciini Günther, 1953 (Phasmatidae: Clitumninae) and provided a list of genera included in Stephanacridini. The five main morphological characters that readily separate Stephanacridini from Pharnaciini as well the whole of Clituminae and Phasmatidae s. str. (= Lanceocercata) are 1) the simple anal segment (abdominal tergum X) of ♂♂ that is not longitudinally tectate and split to consist of two movable hemi-terga that form a clasping apparatus to grasp the ♀ during copulation, 2) the presence of a well-developed and sclerotised vomer in ♂♂, 3) the often enormously elongated gonapophyses VIII in ♀♀, 4) the rather indistinct medioventral carina of the profemora that runs midways on the ventral surface of the profemur in both sexes, and 5) the closed internal micropylar plate of the eggs, which lacks a median line (Hennemann & Conle, 2008: 55). While Hennemann & Conle (2008: 57) already estimated close relation to the genus Platycrana Gray, 1835, Hennemann (2020: 174) supported this hypothesis based on morphological traits and attributed Stephanacridini to the newly established subfamily Platycraninae Hennemann, 2020, which in addition to Stephanacridini comprises the Platycranini. Presently, Platycraninae is considered as the sister taxon of Phasmatidae s. str. (= Lanceocercata), which is supported by molecular (Buckley et al., 2009; Robertson et al., 2018, Bank & Bradler, 2022; Forni et al., 2023) and morphological data (Buckley et al., 2009). While all of the abovementioned subsequent molecular studies support the splitting of Pharnaciini and Stephanacridini introduced by Hennemann & Conle (2008), the relationships and phylogenetic position of Stephanacridini has experienced somewhat contrary results throughout the different approaches. While Buckley et al. (2009) and Robertson et al. (2018) recovered Stephanacridini as sister to Phasmatidae s. str. (= Lanceocercata), Bank & Bradler (2022) hypothesized that Stephanacridini either is the sister taxon of Phasmatidae s. str. (= Lanceocercata) + Xenophasmina, or the sister taxon of Xenophasmina alone. Forni et al. (2022) in contrast revealed Stephanacridini as sister to the New World Haplopodinae. Yet, it appears that still more research is needed to finally clarify the exact placement of Stephanacridini with certainty. As for now and considering the synonymy recovered herein, Stephanacridini now comprises seven valid genera. The two New Guinean genera Stephanacris and Macrophasma Hennemann & Conle, 2006 have been revised by Hennemann & Conle (2006). A revision of Phasmotaenia Návas, 1907 at the species level was presented by Hennemann & Conle (2009) and a taxonomic review of Nesiophasma Günther, 1934 was provided by Hennemann (2021). The genus Sadyattes Stål, 1875 (= Eucarcharus Brunner v. Wattenwyl, 1907 syn. nov.) is reviewed herein. From the two remaining genera of the tribe Diagoras Stål, 1877 is monotypical and as for now only known from the ♀. For a taxonomic review of Hermarchus Stål, 1875, a more comprehensive integrative approach that includes broad molecular sampling to cope with the wide distributional range and numerous island populations seen within the genus is necessary and needs to be addressed by future studies.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFBA5D2AFF52FCE1A3ADF834.taxon	distribution	Distribution. Taiwan, Peninsular Malaysia, Nicobar Islands, Sumatra, Java, Borneo, Philippines, Wallacea, New Guinea, Solomon Islands, NE-Australia, Micronesia and Melanesia (Fiji, Tonga, Samoa, Vanuatu, New Caledonia & Western French Polynesia). Genera included: 1. Diagoras Stål, 1877: 66. Type species: Diagoras ephialtes Stål, 1877: 66, by original monotypy.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFBA5D2AFF52FCE1A3ADF834.taxon	distribution	Distribution: NW-Pacific (Caroline Islands, Palau). 2. Hermarchus Stål, 1875: 45. Type species: Phibalosoma pythonius Westwood, 1859: 73, pl. 12: 1 & 35: 3, by subsequent designation of Kirby, 1904: 361. Distribution: Melanesia (Fiji, Tonga, Vanuatu, New Hebrides, New Caledonia, Western French Polynesia and NE-Australia). 3. Macrophasma Hennemann & Conle, 2006: 3. Type species: Hermarchus biroi Redtenbacher, 1908: 445, by original designation. Distribution: New Guinea. 4. Nesiophasma Günther, 1934: 5. Type species: Nesiophasma eremothocus Günther, 1934: 6, fig. 1, by original designation.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFBA5D2AFF52FCE1A3ADF834.taxon	distribution	Distribution: Wallacea (Sulawesi, Selayar Island, Sangir Islands and Lesser Sunda Islands). 5. Phasmotaenia Návas, 1907: 10. Type species: Taeniosoma sanchezi Bolívar, 1897: 31, fig. 1, by indication. [replacement name for the preoccupied Taenionema Bolivar, 1906] = Taeniosoma Bolivar, 1897: 31. Type species: Taeniosoma sanchezi Bolívar, 1897: 31, fig. 1, by monotypy. [Praeoccupied by Taeniosoma Stimpson, 1857 (Nematoda)] = Taenionema Bolivar, 1906: 393. Type species: Taeniosoma sanchezi Bolívar, 1897: 31, fig. 1, by indication. [Replacement name for preoccupied Taeniosoma Bolivar, 1897 - preoccupied by Taenionema Banks, 1905 (Plecoptera)] = Taeniophasma Uvarov, 1940: 379. Type species: Taeniosoma sanchezi Bolívar, 1897: 31, fig. 1, by indication. [Homonym of Phasmotaenia Návas, 1907 & unnecessary replacement name for Taenionema Bolivar, 1906] = Phasmotaenionema, Günther, 1933: 155. [Misspelling of Phasmotaenia Návas, 1907]	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFBA5D2AFF52FCE1A3ADF834.taxon	distribution	Distribution: Lanyuh Island SE of Taiwan, Philippines, Micronesia, Solomon Islands, New Guinea and Fiji. 6. Sadyattes Stål, 1875: 44, 88. Type species: Sadyattes borrii Stål, 1875: 88, by original monotypy.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFBA5D2AFF52FCE1A3ADF834.taxon	distribution	Distribution: Sundaland (Java, Sumatra, Peninsular Malaysia, Borneo and Nicobar Islands) and Philippines. 7. Stephanacris Redtenbacher, 1908: 441. Type species: Stephanacris brevipes Redtenbacher, 1908: 441, pl. 21: 4 a-b, by subsequent designation of Bradley & Galil, 1977: 194. Distribution: New Guinea.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFB85D20FF52FF2AA01AF983.taxon	type_taxon	Type species: Sadyattes borrii Stål, 1875: 88, by original monotypy.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFB85D20FF52FF2AA01AF983.taxon	description	Redtenbacher, 1908: 444. [in Acrophyllini] Bradley & Galil, 1977: 193. [in Phasmatinae: Pharnaciini] Bragg, 2001: 643. Zompro, 2004: 321. Otte & Brock, 2005: 311, Hennemann & Conle, 2008: 55, 60 [in Stephanacridini] Seow-Choen, 2017: 150. [in Xeroderinae – in error] Seow-Choen, 2018: 452. Hennemann, 2020: 175. [in Stephanacridini] Seow-Choen, 2021: 772. Brock & Büscher, 2022: 559.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFB85D20FF52FF2AA01AF983.taxon	description	Zompro, 2004: 310. Otte & Brock, 2005: 136. Hennemann & Conle, 2008: 132 [in Stephanacridini] Bollens, Krijns & Bresseel, 2010 a: 10. Hennemann, 2020: 175. [in Stephanacridini] Brock & Büscher, 2022: 559. Seow-Choen, 2023: 516.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFB85D20FF52FF2AA01AF983.taxon	description	Seow-Choen, 2018: 421 (in part). Seow-Choen, 2020: 244 (in part). Seow-Choen, 2021: 734 (in part).	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFB85D20FF52FF2AA01AF983.taxon	diagnosis	Diagnosis (♂, ♀). Medium to large (body length ♂♂ 72.3 – 142.0 mm, ♀♀ incl. subgenital plate 127.8 – 245.0 mm) almost exceptionally slender and elongate, stick-like Stephanacridini (only ♀ of one species stocky with body notably broadened); females apterous, ♂♂ winged. Body of both sexes smooth and unarmed, often slightly shiny in ♂♂. Sexual dimorphism strongly developed with ♂♂ much smaller and slenderer than ♀♀. Head ovoid to globose, longer than wide with vertex smooth; no ocelli. Gula small, variable in shape and covering less than half of cervical membrane. Antennae of moderate length, filiform and laid back at least reaching to posterior of mesothorax; comparatively longer in ♂♂ and may reach to abdominal segment V; scapus small, slender and not notably dilated and just moderately compressed dorsoventrally. Mesothorax> 3.5 x longer than prothorax, usually much longer and slenderer in ♂♂; cylindrical, slender and ± uniform in diameter (only in one species with lateral margins notably deflexed and rounded); mesonotum with a ± distinct transverse ridge or bulge at anterior margin. Meso- and metapleurae simple, unarmed. Meso- and metanotum with a ± distinct medio-longitudinal carina or keel. Median segment in ♀♀ at best three-quarters the length of metanotum, much longer than metanotum in ♂♂. Tegmina in ♂♂ elongate and spatulate in shape with the basal half strongly narrowed and the central protuberance shallow. Alae variable in length and moderately developed, at best and only in one species reaching to abdominal segment V, mostly much shorter; anal fan transparent grey or brown. Abdomen excluding median segment longer than head and thorax taken together. Abdominal segments II-VII slender, much longer than wide and uniform in diameter in ♂♂; in ♀♀ less elongate and slightly sub-uniform in diameter (only in one species with lateral margins of terga weakly expanded and rounded). Sterna II-VII with a ± distinct medio-longitudinal carina or keel (more pronounced in ♂♂). Sternum VII in ♀♀ with a moderate praeopercular organ that is formed by a posteromedian pit and a variably shaped granule or tubercle in front. Terminalia of ♀♀: Anal segment weakly tectate and with a shallow concave to triangular posteromedian notch, the lateral margins with a distinct sometimes semi-circular excavation at base of cerci with the basal portion ± deflexed and occasionally lobe-like. Epiproct small, scale-like and scarcely projecting underneath posterior margin of anal segment; keel medio-longitudinally. Cerci small, round in cross-section and ± tapering towards the tip. Gonapophyses VIII elongated, rather filiform and up-curved but never considerably projecting beyond anal segment. Subgenital plate bulgy in basal portion, variable in length and shape and always projecting beyond tip of abdomen by at least two-thirds the length of anal segment and as much as slightly more than the length of the three terminal terga taken together; the flattened posterior half ranging from broad, spatulate and roundly angular to strongly tapering, pointed and lanceolate. Terminalia of ♂♂: Tergum VIII slightly to distinctly inflated and widened posteriorly, but three terminal segment usually not distinctly broader than preceding segment. Anal segment simple, gently tectate longitudinally with the posterior margin ± emarginated and the outer angles ± expanded and triangular and with a distinct and large thorn-pad ventrally. Epiproct very small and fully concealed under anal segment. Vomer a strongly sclerotised, variably shaped plate that ranges in shape from triangular over heart- or pear-like to having the basal portion sub-circular; always with a single ± arched terminal hook. Phallus enlarged, elongated and often projecting over upper lateral margin of poculum. Cerci fairly small, elongate, round in cross-section, ± curved and with the apex ± club-like. Poculum small and at best scarcely projecting beyond posterior margin of tergum IX; basal portion weakly to moderately bowl-shaped and only in two species with a small, obtuse central protuberance; the posterior margin ± labiate and occasionally with a slight median indention. Legs all long and slender (only ♀♀ of two species have noticeably incrassated meso- and metafemora), even more so in ♂♂; all carinae to a variable degree dentate or serrate and usually no considerably enlarged teeth or lobes present on mid and hind legs except for an enlarged apical tooth on two outer ventral carinae of meso- and metafemora. Medioventral carina of meso- and metafemora dentate to spinose. Armature of limbs generally less developed in ♂♂. Probasitarsus with dorsal carina unarmed; ventral carinae of all protarsi minutely dentate. Dorsal carina of basitarsi in ♀♀ ± raised or crested (occasionally strongly deflexed in probasitarsus); basitarsi always slender in ♂♂; in both sexes notably elongated and at least as long as following three tarsomeres taken together. Eggs (Fig. 21). Of average size if compared to the eggs of other genera of the tribe and comprising a wide range of different chorion shapes and surface textures; mostly not noticeably longer than wide (2 x longer than wide only in one species) and more or less compressed laterally (strongly globose with a smooth surface only in one species). Dorsal surface often with two longitudinal carinae or bulges. Polar area mostly rounded but may be flattened (one species) or with a central protrusion (one species). Surface of chorion often with a mesh-work of ridges, keel or bulges, occasionally pitted to a variable degree. Micropylar plate less than three-fifth the length of chorion and spearhead-shaped to rhomboidal; the lower portion more or less distinctly narrowed; closed internally. Micropylar cup fairly central within the borders of the micropylar plate. Operculum oval and flattened; often arched with the two ends facing the dorsal and ventral egg surfaces distinctly downward-directed and in these species with a conspicuous pit or impression in the portion facing the dorsal egg surface. Capitulum not stalked, variable in size and shape. Differentiation. Sadyattes is morphologically closest to Nesiophasma, which is distributed throughout most of Wallacea (Hennemann, 2021: 103), and presumably the sister taxon. Based on the available data there seems to be no distributional overlap between these two genera, which means that they are biogeographically separated by the Wallace’s Line that runs between Borneo and the Philippines in the west and north, and Sulawesi in the east. The islands east of Wallace’s Line are termed Wallacea which is principally regarded a transition zone between the Oriental and Australian regions (Dickerson, 1928). Although the Philippines are now also considered as part of Wallacea (e. g. Vallejo, 2011), which is in accordance with the interpretation of Wallace’s Line after Huxley (1868) separating the Philippines from Borneo, the distribution of Sadyattes however reflects the faunal boundary originally intended by Wallace that includes the Philippines on the west side of the line (Wallace, 1876). Such, the Oriental Sadyattes has seven of its fourteen known species distributed in the Philippines, whereas the Wallacean Nesiophasma has no representatives in the Philippines (Hennemann, 2021: 104). Whereas ♀♀ of both genera are virtually identical in means of morphology and very difficult to separate from another (Hennemann, 2021: 104), ♂♂ of Sadyattes readily differ from the consistently apterous ♂♂ of Nesiophasma by the presence of tegmina and alae and therefore much longer median segment, that is considerably longer than the metanotum. In ♀♀ on Nesiophasma the anal segment shows a deep and narrow posteromedian incision or slit, whereas in Sadyattes there merely is a shallow concave to triangular excavation. The mostly distinct lateral excavation of the anal segment at the base of the cerci on the other hand is shared between both genera. Females have the gonapophyses VIII always hidden under and never projecting beyond the tip of the anal segment like in most Nesiophasma - species, but there is one species of Nesiophasma, namely N. plateni (Dohrn, 1910) that also has short gonapophyses VIII (Hennemann, 2021: 117, Figs. 45 P-R) and therefore renders this possible distinguishing character not fully reliable. This is also the case with the variable subgenital plate, which is fairly short and broadly spatulate in several species, a condition never seen in Nesiophasma, but other species have it long and lanceolate like in Nesiophasma - species. The known eggs of Sadyattes are remarkably polymorphic and show a wide range of different shapes and chorion surface textures that also render a reliable delimitation and distinction from eggs of Nesiophasma impossible. All eggs, except for those of the Bornean S. decoris (Seow-Choen, 2016) comb. nov. which is here transferred from the Pharnaciini genus Phobaeticus Brunner v. Wattenwyl, 1907, have the chorion more bulgy and less elongate than in Nesiophasma being not considerably longer than wide. The eggs of several Philippine species have a conspicuous pit or impression in the portion of the operculum that faces towards the dorsal egg surface and moreover have the operculum notably arched with the two ends facing the dorsal and ventral egg surfaces distinctly downward-directed. Both features are not seen in eggs of Nesiophasma. Altogether, reliable morphological differences between Sadyattes and Nesiophasma, other than the presence of absence of wings in the ♂♂ and minor differences mentioned above, are very hard to define why it is hoped that molecular data can assist with the delimitation of these two genera in due course.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFB85D20FF52FF2AA01AF983.taxon	discussion	Remarks. Stål (1874: 44, 88) originally described Sadyattes based on a unique ♂ without locality in the collection of RBINS, namely S. borrii, and the true identity of the genus has long been questionable although subsequent authors linked it with taxa that now belong in Clitumninae: Pharnaciini. Hennemann & Conle (2008: 55, 60) examined the holotype specimen in detail and transferred Sadyattes from Pharnaciini to the tribe Stephanacridini Günther, 1953, mainly based on morphological characteristics of the ♂ terminalia, like the presence of a well-developed vomer and simple, non-split anal segment. Still however, the identity of the genus has remained unsolved because the ♀ and eggs were not known. It was until now that the availability of new material of the type-species S. borrii Stål, 1875 has rendered a clarification of the taxonomic position and delimitation of Sadyattes within Stephanacridini possible and proven this species to be distributed on Java and Sumatra. It also became obvious that Stål’s borrii is congeneric to the type-species of Eucarcharus Brunner v. Wattenwyl, 1907, the Philippine E. feruloides (Westwood, 1859), why Eucarcharus needs to be synonymised under Sadyattes (syn. nov.). This has already been suggested previously by Hennemann (2021: 104). Moreover, further investigation of the two type specimens of Westwood’s feruloides, one from the Philippines (lectotype) and one from Java (paralectotype), has shown that these represent two distinct species, which has already been suggested by Seow-Choen (Seow-Choen, 2023: 516). In fact, the Javanese paralectotype is a specimen of borrii, the type-species of Sadyattes. Comprehensive investigation of species with questionable generic affiliations within Hermarchus Stål, 1875 (tribe Stephanacridini) and Phobaeticus Brunner v. Wattenwyl, 1907 (Clitumninae: Pharnaciini) has uncovered three further species that actually belong in Sadyattes and are here transferred (comb. nov.). It seems worth mentioning that members of Sadyattes are remarkably similar in overall appearance to species of Phobaeticus Brunner v. Wattenwyl, 1901, a genus of Clitumninae: Pharnaciini, that has frequently been shown to be not closely related (e. g. Hennemann & Conle, 2008; Bradler, 2009; Buckley et al., 2009; Robertson et al., 2018, Bank & Bradler, 2022; Forni et al., 2023). Representatives of both genera have widely overlapping distributions, are large to very large slender stick insects with wingless ♀♀ and much smaller, more colourful winged ♂♂ and often have a more or less elongated subgenital plate in the ♀♀. It may be for these reasons why, several of the species here transferred to Sadyattes have previously been misplaced in Phobaeticus. One species originally described in the genus Phobaeticus and the world’s longest known insect, the Bornean Phobaeticus chani Bragg, 2008, has even been erroneously attributed to Sadyattes subsequently but was re-transferred to Phobaeticus (Hennemann, 2021: 104). As argued by Hennemann (2021: 104) Phobaeticus chani shows all the distinctive morphological traits that separate members of Clituminiae: Pharnaciini from the tribe Stephanacridini, this the split anal segment in ♂♂ that consists of two hemi-terga, lack of a vomer, short gonapophyses in ♀♀ and the lamellate, displaced medioventral carina of the profemora.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFB85D20FF52FF2AA01AF983.taxon	distribution	Distribution (Fig. 1). Java, Sumatra (including the surrounding islands of Banka, Enggano and Nias), Peninsular Malaysia, Nicobar Islands, Borneo and Philippines. Species included: 1. Sadyattes annulatus (Redtenbacher, 1908: 451) [Pharnacia]. (Fig. 2) Distribution: Borneo. 2. Sadyattes banwaon sp. nov. (Fig. 3) Distribution: Philippines (Mindanao). 3. Sadyattes borrii Stål, 1875: 88. (Figs. 4 – 5)	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFB85D20FF52FF2AA01AF983.taxon	distribution	Distribution: Java and Sumatra. 4. Sadyattes decoris (Seow-Choen 2016: 267, figs.) [Phobaeticus]. comb. nov. Distribution: Borneo. 5. Sadyattes enganensis (Redtenbacher, 1908: 451) [Pharnacia]. Distribution: Enggano Id. and Nias Id. East of Sumatra and Peninsular Malaysia. 6. Sadyattes fallax (Brunner v. Wattenwyl, 1907: 186) [Eucarcharus]. comb. nov. (Fig. 6) Distribution: Philippines (Luzon, Samar and Panay). 7. Sadyattes feruloides (Westwood, 1859: 45, pl. 6: 5) [Lonchodes]. comb. nov. (Fig. 7) Distribution: Philippines (Luzon). 8. Sadyattes incertus (Brunner v. Wattenwyl, 1907: 185) [Phobaeticus]. comb. nov. (Figs. 8 – 9) = Nearchus grubaueri Redtenbacher, 1908: 448. syn. nov. Distribution: Sumatra, Banka Id., Peninsular Malaysia and Nicobar Islands. 9. Sadyattes leytensis (Zompro, 1997: 38, figs.) [Hermarchus]. comb. nov. (Figs. 10 – 11) Distribution: Philippines (Leyte, Mindanao and Panay). 10. Sadyattes maganda sp. nov. (Figs. 12 – 13) Distribution: Philippines (Mindoro). 11. Sadyattes matipuno sp. nov. (Figs. 14 – 15) Distribution: Philippines (Luzon). 12. Sadyattes mindanaense sp. nov. (Figs. 16 – 18) Distribution: Philippines (Mindanao). 13. Sadyattes nigricornis (Redtenbacher, 1908: 451) [Pharnacia]. Distribution: Borneo. 14. Sadyattes panayense sp. nov. (Fig. 19) Distribution: Philippines (Panay) 15. Sadyattes tubaense sp. nov. (Fig. 20) Distribution: Philippines (Luzon)	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFB35D22FF52F9C7A63EF9A2.taxon	description	(Figs. 2, 22 A)	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFB35D22FF52F9C7A63EF9A2.taxon	description	Seow-Choen, 1998: 89. Otte & Brock, 2005: 268. Baculolonga annulata, Bragg, 2001: 378 Sadyattes annulatus, Hennemann & Conle, 2008: 132. Seow-Choen, 2017: 151, figs. [Illustrations of HT] Seow-Choen, 2019: 268. Brock & Büscher, 2022: 559.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFB35D22FF52F9C7A63EF9A2.taxon	materials_examined	Further material: 1 ♂: Borneo, W-Kalimantan, Prov. Kalimantan Barat, Mount Bawang, 00 ° 53,5 ’ N 109 ° 22,2 ’ O, ca. 250 m, local collector II-IX. 2009 [FH, No. 1390 - 1]. Differentiation. The ♂ of this Bornean species (the only sex known) is morphologically and chromatically very similar to that of incertus comb. nov., a species found in Peninsular Malaysia, islands in the Strait of Malacca and on the Andamans, with which it shares the long alae that reach as far back as to abdominal segment V. It can however be separated by the somewhat more globose head (Fig. 2 B), which is less narrowed posteriorly and has the posterior margin less angulated if seen in lateral aspect, less pronounced medio-longitudinal carina of the mesosternum, shorter and rather gradually tapering vomer (Fig 2 G; almost reaching to epiproct and with a slight widening preapically in incertus), generally being more colourful but with the meso- and metafemora almost uniformly green with the exception of a black apex (Fig. 2 D; with a faint yellowish pre-apical transverse band in incertus).	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFB35D22FF52F9C7A63EF9A2.taxon	discussion	Remarks. This species is likely to be the opposite sex of S. mjoebergi (Günther, 1935), which is here removed from the genus Phobaeticus Brunner v. Wattenwyl, 1907 and transferred to Sadyattes (comb. nov.). However, specimens of both sexes from the same locality or from captive breeding need to become available for examination to prove or disprove this assumption. Hennemann & Conle (2008: 132) removed annulatus from the genus Pharnacia Stål, 1877 and placed it in Sadyattes. Seow-Choen (2017) provided detailed figures of the holotype ♂ in the collection of MNHU. Female and egg unknown. The ♀ from Bako National Park-NW-Sarawak illustrated by Seow-Choen (2016: 274, Fig. 618) as Phobaeticus mjoebergi is misidentified and undoubtedly a species of Sadyattes (Fig. 22 A). Judging from the fairly large size (body length including subgenital plate 206.0 mm, without subgenital plate 178.0 mm) it is not unlikely to be the opposite sex of annulatus. Phobaeticus mjoebergi (Günther, 1935) keys out as a member of the tribe Pharnaciini (subfamily Clitumninae) because Günther (1935 c: 11) explicitly mentions a praeopercular organ on abdominal sternum VII that consists of two peg-like appendages, and also the strongly serrated “ kräftig gesägt ” (Günther, 1935 c: 10) anterodorsal carina of the profemora and unarmed medioventral carinae of the meso- and metafemora, as well as a median segment that is about three-quarters the length of the metanotum affiliate mjoebergi with the genus Phobaeticus Brunner v. Wattenwyl, 1907 or at least the tribe Pharnaciini respectively. Unfortunately, the holotype from Mount Tibang, Kalimantan in the collection of NHRS could not be examined for the present study. The unique holotype is strongly discoloured due to former preservation in ethanol, but the ♂ in the author’s collection allows a description of the natural colouration of this pretty insect. Therefore, illustrations and a description of the colouration are here provided as follows (Fig. 2): General colour dark green with the head, prothorax and three basal abdominal segments buff; the lateral areas of pronotum dark brown and the meso- and metapleurae dark yellowish. Abdominal terga IV – VII with a washed transverse posterior band and a black posterolateral marking and yellowish anterolaterally; sterna II-VII green with the posterior portion yellow and with a small black marking posterolaterally. The terminal three terga mostly dark yellow; VII and VIII each with a cream-colored median marking laterally and with a small black marking antero- and posterolaterally (Figs. 2 E – F). Sternum VII with a black posterior transverse band (Fig. 2 G). Tegmina and costal region of alae light cream-coloured with a yellowish stripe along anterior margin and a broad black medio-longitudinal streak (Fig. 2 C). Meso- and metafemora dark green with the apex black (Fig. 2 D); the profemora black with irregular orange mottling that form transverse bands in apical half. Tibiae black with three orange transverse annulae. All teeth of limbs black. Basitarsi black with basal half orange. Antennae black. Measurements in table 1 below.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFB35D22FF52F9C7A63EF9A2.taxon	distribution	Distribution: Borneo. Central Kalimantan (Kalimantan Tengah, East Kotawaringin Regency, Sampit [MNHU – type locality]; West Kalimantan (Kalimantan Barat, Mount Bawang, ca. 250 m [FH]).	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFB05D3DFF52FD65A356FB9E.taxon	description	(Fig. 3) HT, ♀: Philippinen, Mindanao Id., Prov. Agusan del Sur, San Francisco Munip., Borbon, local collector 24. IV. 2012 [RBINS, ex coll. FH]. Differentiation. The ♀ of this new species (the only sex known) is morphologically very close to the ♀♀ of S. panayense n. sp, from the island of Panay and S. maganda sp. nov. from the island of Mindoro. With both species it shares the overall shape and long, lanceolate subgenital plate (Figs. 3 G – I) but may be separated by the smaller size, proportionally shorter limbs, that have the all the dentations comparatively smaller and less developed, but instead possess a distinctly enlarged apical tooth on the two outer ventral carinae of the mesofemora (Fig 3 D) and bear a distinct, rounded apical crest on the meso- and metatibiae. Moreover, the mesopleurae are reddish in colour (Fig. 3 E) and distinguish this ♀ from that of maganda this species also differs by the slenderer meso- and metafemora and from panayense it may also be separated by the relatively longer mesothorax (5.5 x vs. 4.5 x longer than prothorax) and lack of a definite medio-longitudinal carina on the meso- and metanotum Moreover, also the structure of the praeopercular organ and shape of the anal segment as well as the comparatively larger epiproct distinguish banwaon from the other two ♀♀.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFB05D3DFF52FD65A356FB9E.taxon	etymology	Etymology. Named after the Banwaon people, an indigenous ethnic tribe that is located in Agusan del Sur Province in Mindanao, the type-locality of this new species. They are traditionally nomadic forest dwellers but now typically live in settled villages, relying on agriculture and fishing for their livelihood. Neuter.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFB05D3DFF52FD65A356FB9E.taxon	description	Description. The unique holotype lacks most of the antennae and has suffered from damage by dermestid beetles, which have destroyed considerable parts of the posterior portion of the head capsule as well as the left eye (Figs. 3 E-F). ♀ (Fig. 3). Small (body length incl. subgenital plate 139.0 mm) and of average form for the genus with a long and lanceolate subgenital plate. General colour of the holotype ochraceous buff with an olive hue on the abdominal terga and base of subgenital plate. Transverse anterior bulge of mesonotum and mesopleurae reddish (Fig. 3 E), the latter with the upper area green in anterior half. Genae with a few irregular washed blackish speckles posteriorly and around compound eye (Fig. 3 E). Dentations of the legs tipped with black (ventral dentations in particular). Eyes ochre and the antennae dark greyish brown with the apex of each joint dark yellowish. Head (Figs. 3 E – F): Oval, about 1.45 x longer than wide, broadest at the eyes with the genae slightly narrowing; vertex moderately convex with a fine coronal line and the lateral furrow weakly indented posteriorly. Frons with a distinct, short and gently arched furrow just behind bases of antennae. Eyes moderately convex and their diameter corresponding to about 0.5 x length of genae. Antennae broken in the holotype with the left antennae present only until the sixteenth joint; the joints slightly unequal in length and somewhat decreasing in length towards the front. Scapus rectangular, somewhat compressed dorsoventrally and about 1.8 x longer than wide; medio-dorsal portion black longitudinally. Thorax: Pronotum roughly rectangular and weakly constricted pre-medially, slightly shorter but much narrower than head; the lateral margins gently concave. Anterior margin gently concave and slightly inflated with a distinct transverse furrow behind; the median transverse sulcus weakly arched and just not reaching lateral margins of segment; in front with a pair of shallow median swellings (Figs. 3 E – F). Meso- and metanotum with a weakly indicated medio-longitudinal line, otherwise wholly smooth. Mesothorax somewhat constricting in anterior onequarter, somewhat inflated pre-medially and 5.5 x longer than prothorax; the anterior margin of mesonotum with a fairly well-developed transverse bulge and two low granules posterior of it. Metanotum parallel-sided, about half as long as mesonotum and 2.7 x longer than wide. Pleurae simple, the sterna with a very shallow medio-longitudinal ridge (Fig. 3 C). Abdomen: Median segment almost 2 x longer than wide, trapezoidal in outline with anterior margin notably narrower than posterior margin and only 0.45 x length of metanotum. Segments II – VII slightly sub-uniform in diameter, II-IV slightly increasing in length and V – VII decreasing in length; II 1.3 x longer than median segment and VII slightly longer than II, all on average 2.3 x longer than wide. Terga smooth except for a shallow and obtuse longitudinal carina close to lateral margins; V with posterior margin somewhat inflated. Sterna with a moderately developed medio-longitudinal ridge. Praeopercular organ formed by an irregularly rugulose swollen posteromedian area on sternum VII, which has a small verrucose swelling in front and a shallow impression posteromedially (Fig. 3 I). Terga VIII – X uniform in width, VIII about two-thirds as long as VII and IX about quadrate in outline and half as long as VIII. Anal segment roughly rectangular in dorsal aspect and somewhat longer than IX, rather weakly tectate longitudinally and only with a shallow concave post-median lateral excavation; the posterior margin obtusely angular laterally and slightly rounded medially with a small and shallow median indention (Fig. 3 H). Epiproct large, scale-like with a distinct medio-longitudinal keel, almost semi-circular and distinctly projecting over anal segment (Fig. 3 H). Cerci small, conical and tapering towards a narrow and pointed tip, slightly projecting over posterior margin of anal segment. Gonapophyses VIII moderately elongated, and not reaching tip of anal segment (Fig. 3 G). Subgenital plate lanceolate with apical portion slightly but gradually narrowing towards a pointed, triangular apex (Figs. 3 H – I); projecting beyond tip of abdomen by roughly the combined length of two terminal terga; basal portion with a prominent ledge-like longitudinal lateral carina (Figs. 3 G, I). Legs: Moderately slender and fairly short fort the genus with all carinae rather minutely dentate; the ventral dentations generally more pronounced than the dentations of dorsal carinae. Profemora about as long as mesothorax, mesofemora notably shorter than mesothorax, metafemora reaching three-quarters along abdominal segment IV and metatibiae only reaching to posterior of segment VI. Posteroventral carina of profemora with about 15 – 16 acutely pointed and sharp teeth, the anteroventral carina with about 20 broader and more closely as well as regularly spaced saw-like teeth; posterodorsal carina supplied with about 15 – 17 small denticles. Dentations of mid and hind legs small and rather spinose in shape; the two outer ventral carinae of meso- and metafemora with a prominently enlarged and broad triangular apical tooth, which is accompanied by a smaller tooth anteriorly on anteroventral carina (Fig. 3 D; these notably more distinct on mesofemora). Medioventral carina of meso- and metafemora distinct and armed with eleven (mesofemora) or 12 – 13 moderately distinct spines, the three or four sub-basal ones of which are the most prominent; medioventral carina of meso- and metatibiae spinose as well. Posterodorsal carina of meso- and metafemora elevated into an almost semi-circular crest apically. Basitarsi with all carinae minutely denticulate (teeth least distinct on probasitarsus) and the dorsal carina slightly raised and rounded apically; slightly longer than following three joints taken together. Measurements [mm]: Body (incl. subgenital plate) 139.0, body 132.7, pronotum 4.6, mesonotum 26.7, metanotum 13.5, median segment 6.2, profemora 29.0, mesofemora 21.2, metafemora 27.0, protibiae 29.5, mesotibiae 18.4, metatibiae 25.8, antennae> 8.5. Reamrks. So far only known from the unique ♀ holotype. Male and egg unknown.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFB05D3DFF52FD65A356FB9E.taxon	distribution	Distribution. Philippines: Mindanao (Agusan del Sur, San Francisco Municipality, Borbon [RBINS – type locality]).	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFAE5D39FF52FB3BA131F882.taxon	description	(Figs. 4 – 5)	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFAE5D39FF52FB3BA131F882.taxon	materials_examined	Further material: 1 ♂: Rajamandala, Mts. Djampang, W. Java; Le Moult vend. via Reinbek, Eing. Nr. 1, 1957 [ZMUH]; 1 ♀: Java, H. Fruhstorfer [ZMPA]; 1 ♂: W-Sumatra, Prov. Sumatera Barat, Padang area, local collector, V. 2002 [FH, No. 1523 - 1]; 1 ♀ (immature): W-Sumatra, Prov. Sumatera Barat, Sijunjung Region, local collector, VI. 2012 [FH, No. 1523 - 2]; 1 ♂: no data [ZSM]. Differentiation. Both sexes of this, the type-species of Sadyattes, are morphologically closest to S. enganensis (Redtenbacher, 1908) from Peninsular Malaysia, Sumatra and the islands of Enggano and Nias east of Sumatra, with which they share the distinctive pale cream-coloured protarsus of both sexes (Figs. 4 C, 5 F). Females of both species also have in common the distinct posterolateral lobes of abdominal terga VIII – X (Fig. 4 E), conspicuous shape of the anal segment, which has the lateral margins deeply and almost semi-circularly excavated and the posterior margin broadly bi-lobed (Figs. 4 E, G), as well as the rounded dorsal lobe of the probasitarsus (Fig. 4 C). However, ♀♀ of borrii (Fig. 4) are less robust in overall shape, have comparatively slenderer legs, abdominal segments III – VI not swollen as in enganensis, the subgenital plate has the apex more obtuse and the dorsal lobe of the probasitarsus is not as high as in enganensis. At first glance ♀♀ appear similar to those of S. incertus (Brunner v. Wattenwyl, 1907) from Peninsular Malaysia, Bangka Island in the Strait of Malacca and the Andamans, but readily differ by the stockier shape and more ovoid head as well as the distinctive lateral lobes of abdominal terga VIII – X, conspicuous shape of the anal segment and contrastive pale cream-coloured protarsus, which has the basitarsus distinctly lobed dorsally. Males of borrii (Fig. 5) are easily distinguished from those of enganenesis by the slightly slenderer form and notably longer abdominal segments II – VI (6 x vs. 4.5 x longer than wide in enganensis), less globose head which has the vertex notably less convex (Fig. 5 E), not distinctly club-shaped apex of the abdomen (Figs. 5 H – J), slightly more elongated and isosceles triangular vomer (Fig. 5 J; with a slight median widening in enganensis), lacking the dark brown medio-longitudinal streak seen on the tegmina of enganensis, having all the dentations of the limbs more acutely pointed and distinctly black as well as the characteristic orange outer ventral carinae of the mesofemora (Figs. 5 C – D).	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFAE5D39FF52FB3BA131F882.taxon	description	Description. Since this is the type-species of Sadyattes, and only a very brief original description of the ♂ was provided by Stål (1875: 88), a detailed re-description is here provided. The three non-type examples at hand match very well with the holotype and show no significant variability. The holotype has suffered from some damage since it’s description in 1875 with three of its legs uncarefully glued to the body in very awkward positions and the left front leg glued to a piece of cardboard. It however is almost complete and merely lacks the right profemur. Measurements in table 2 below. ♂ (Fig. 5). Fairly large (body length 121.0 – 131.0 mm) for the genus, form slender, with distinctive orange outer ventral carinae of the meso- and metafemora and contrastive cream-coloured protarsi. Overall colour of body and legs olive, the abdomen genera generally more brownish to drab. Head pale ochraceous, the vertex sometimes of a slight greenish hue (Figs. 5 E – F). Eyes russet. Antennae greyish brown or reddish dark brown (specimen in ZMUH) with scapus and pedicellus dark ochre; following five or so antennomeres blackish ventrally. Meso- and metapleurae with a faint greyish stripe along lower margin; the prosternum and very posterior part of meso- and metasternum orange brown. Tegmina and costal region of alae light ochre with the main longitudinal veins brown and the anterior margin broadly cream-coloured, the tegmina with the radial vein marked by a washed but broad brown streak throughout basal half; anal fan transparent grey with greyish brown veins. Abdominal sterna II-VI each with a partly fused pair of faint whitish pro-posterior margin. All femora olive and slightly brownish at the apex with the two outer ventral carinae of the meso- and metafemora orange except for the basal portion (Figs. 5 C-D); profemora gradually becoming brown from the middle onwards and with irregular dark brown speckles and annulae. Tibiae dark ochre with three faintly orange annulae. All dentations contrastive black and marked by a black spot at the base (Figs. 5 C – D). Protarsi cream-coloured. Head (Figs. 5 E – F): Sub-globose, somewhat flattened, vertex moderately convex and with a shallow but large impression in front; broadest at eyes and notably narrowing towards the posterior. Coronal line distinctly, lateral furrow weakly indented. Frons with a small but distinct transverse impression between bases of antennae. Eyes very large, projecting sub-spherically and their diameter corresponding to as much as 0.8 x length of gena. Antennae long and reaching to posterior margin of abdominal segment III; scapus rectangular and 1.5 x longer than wide, pedicellus less than half as long as scapus und round in cross-section; following antennomeres first unevenly increasing then decreasing in length towards apex of antennae. Thorax: Pronotum slightly shorter and noticeably narrower than head, roundly rectangular and 1.8 x longer than wide; anterior margin concave, raised, swollen medially and followed by three impressions, the median one rather shallow and triangular the two outer ones near anterolateral angles of notum oval and deep (Figs. 5 E – F). Median line impressed over whole length of segment, transverse median sulcus short and almost straight. Mesothorax elongate, 6.5 x longer than prothorax and just weakly widened at anterior margin and in posterior portion. Mesonotum with a weakly indicated medio-longitudinal line and a fairly distinct transverse ridge, which has the median part somewhat labiate and curved towards the anterior. Meso- and metasternum simple; the medio-longitudinal line just scarcely indicated in anterior portion of mesosternum. Tegmina spatulate in shape with the basal two-thirds gradually constricting and the apical oner-third rounded; central hump small and obtusely gibbose (Fig. 5 D). Alae reaching to posterior margin of abdominal segment III. Abdomen: Segments II – IV slightly increasing and V – VII decreasing in length, all uniform in diameter; IV longest and about 6 x longer than wide, VII shorter than all preceding and less than 5 x longer than wide. Tergum V with a prominent transversely gibbose posterior swelling (Fig. 5 G). Sterna II – VII with a shallow medio-longitudinal line. Tergum VIII slightly widening towards posterior, trapezoidal in outline and a little less than two-thirds the length of VII; IX rectangular and about three-quarters the length of VIII. Anal segment moderately tectate longitudinally, with posterior portion somewhat constricted; the posterior margin with a shallow median indention, the outer angles rounded in later aspect, the ventral surface supplied with several minute denticles facing each other at an angle of about 70 – 80 ° (Fig. 5 I). Vomer elongate, basically triangular in outline with the base somewhat widened and then gradually narrowing towards a fairly long, slightly up-curved terminal hook; the base with a narrow, curved transverse furrow, the central portion of ventral surface forming an indented triangular area and the outer lateral margins notably inflated (Fig. 5 J). Cerci round in cross-section with the apex slightly club-shaped and notably projecting beyond anal segment. Phallus projecting slightly over tight lateral margin of poculum (Figs. 5 H – J). The poculum large, bulgy with an obtuse central protrusion at the base (Fig. 5 H); the posterior margin slightly labiate, broad, widely bi-lobed and slightly projecting over posterior margin of tergum IX (Fig. 5 J). Legs: Long, slender and with basically all carinae supplied with rather widely spaced flat, saw-tooth like teeth at fairly regular intervals; the spaces between the teeth gradually decreasing towards apex of each femur or tibia and teeth generally less pronounced on front legs. Two outer ventral carinae of meso- and metafemora with about 12 – 14 teeth, medioventral narrow and supplied with only about ten minute spines. Profemora about as long as head, pro- and mesothorax combined, mesofemora as long as combined length of pro- and mesothorax, metafemora reaching about halfway along abdominal segment VI and metatibiae projecting beyond tip of abdomen by more than combined length of five terminal abdominal segments. Meso- and metabasitarsi about as long as remaining tarsomeres taken together and serrate ventrally with the dorsal carina smooth. Probasitarsi wholly unarmed and notably longer than remaining joints combined.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFAE5D39FF52FB3BA131F882.taxon	discussion	Remarks. The identity and distribution of this species has long been a mystery, which has also had effect on the understanding of the genus Sadyattes, of which it is the type species. Thus, no subsequent author understood the true taxonomic position of Stål’s genus until Hennemann & Conle (2008: 132) recognised the congeneric nature of several previously misplaced species, which they transferred to Sadyattes. But still there was no proper diagnosis of the genus. Stål (1875: 88) described Sadyattes borrii, and therefore the genus Sadyattes, from a unique ♂ holotype without locality in the collection of RBINS. Meanwhile two further ♂♂ have been identified as being conspecific with borrii and provide the first definite records for Stål’s species, which in combination with the previously unrecognised ♀♀ render Java as the most likely type-locality of borrii. A complete and nicely preserved ♀ is housed in the collection of ZMPA, which was recorded as “ Eustygera feruloides ” by Dohrn (1910: 405), who believed that this specimen would confirm Java as a locality for Westwood’s feruloides. However, the penultimate instar ♀ paralectotype of feruloides from Java in the collection of NHMUK is specifically distinct from the Philippine lectotype and actually is a specimen of Stål’s borrii. That this Javan specimen was most certainly a distinct species, has already been suggested by Seow-Choen (2023: 516) who provided detailed photographs of both of Westwood’s type specimens. Body length of the paralectotype ca. 125.0 mm. Nesiophasma zanus Hennemann, 1999 was described from a unique ♀ in the collection of RBINS, which bears a label that states “ Nouvelle Guinee, Fruhstorfer ”. This locality information prompted Hennemann (1999) to assume the species belonged in the genus Nesiophasma Günther, 1934, which the author however doubted subsequently and suspected the locality to be wrong (Hennemann, 2021: 124). Actually, careful re-examination of the holotype has supported the falsity of the locality and shown the specimen be a ♀ of borrii, therefore most likely to be from Java. Therefore, zanum is here synonymised under borrii (syn. nov.). Comparison with the holotype of borrii in the same collection even suggests these two specimens might originate from the same source, collecting event and maybe even locality. Both are preserved on the same type of pin, which has been placed through the posterior portion of the mesothorax, appear to be of similar age and both show similar damage with several of the legs glued to the insect with apparently the same type of glue. The holotype of zanus has been repaired and re-set with the legs aligned and the front legs folded when it was described by the author in 1999, but previously it also had the legs arranged in a similar manner to the holotype of borrii. Egg unknown.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFAE5D39FF52FB3BA131F882.taxon	distribution	Distribution. Java: W-Java (Jawa Barat, Sukabumi, Distr. Djampang Kulon, Rajamandala [ZMUH]; Jawa Barat, Gung Gede-Pangrango N. P., Gede Volcano, Tjibodas [USNM?]); “ Java ” [ZMPA]. W-Sumatra: (Sumatera Barat, Padang area [FH]; Sumatera Barat, Sijunjung area [FH]).	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFA95D3BFF52FC73A6BAFBBA.taxon	description	(Figs. 21 K – L, 23 A)	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFA95D3BFF52FC73A6BAFBBA.taxon	description	Brock & Büscher, 2022: 554. Differentiation. The ♀ (Fig. 23 A) of this Bornean species (the only sex known) most closely resemble the Philippine S. panayense sp. nov. with which they share the overall appearance and shape, as well as the size and elongated, lanceolate subgenital plate. This species however is geographically well separated from any Philippine congenerics and moreover differs from panayense by the more elongate head, much smaller serrations of the limbs and notably longer subgenital plate, which extends beyond the tip of the abdomen by the combined length of abdominal terga VIII – X. The only similar Bornean species is S. mjoebergi (Günther, 1935), which however is much larger (body length of the holotype including subgenital plate 249.0 mm), has the medioventral carina of the meso- and metafemora unarmed and the subgenital plate even longer. At first glance ♀♀ of decoris also strongly resemble those of S. incertus (Brunner v. Wattenwyl, 1907) from Peninsular Malaysia, Bangka Island east of Sumatra and the Andaman Islands. They can however be separated by the smaller size (body length including subgenital plate> 189.0 mm in incertus) and somewhat stockier overall shape and stockier limbs, more ovoid head, smaller and rather serrate than spinose dentations of the extremities, shorter subgenital plate, which extends beyond the tip of the abdomen by no more than the length of the three terminal abdominal terga taken together, and less pronounced praeopercular organ. Egg (Figs. 21 K – L). The following description of the eggs is presented because Seow-Choen (2016: 269, Fig. 609) only presented a picture and based on colour photographs kindly provided by Francis Seow-Choen (Singapore). Unfortunately, none of the pictures is scaled, no measurements were taken and no samples were saved for further study since these eggs laid by the paratype were used for a trying to breed the species (personal communication with F. Seow-Choen). Size unknown. Shape very elongate for the genus, chorion 1.8 x longer than high, flattened laterally and distinctly oval in cross-section; polar-area with a deep crater-like central indention and the dorsal and ventral surfaces each with a central impression, which is more pronounced on dorsal surface; posterior portions of dorsal and ventral surfaces bulgy, the posterodorsal swelling with an indention below. Surface slightly shiny, minutely coriaceous and unevenly pitted; an arched transverse row of 5 – 7 deep and enlarged pits ventrally on lateral surfaces and a transverse rim of notably enlarged pits also below anterior margin; the latter raised and densely covered by small, uneven, longitudinally directed rugulae. Micropylar plate positioned roughly in centre of medio indention of dorsal surface, small, roughly one-third as long as capsule with the broadened upper portion drop-shaped and the posterior portion strongly narrowed; outer margin somewhat bulgy and the surface indented and concave. Micropylar cup just above posterior narrowing of plate and distinctly bowl-shaped. Operculum sub-circular, slightly ascending towards the centre and with fine radially directed ridges. Capitulum a large, basally cylindrical protrusion. That has the apex constricted with the upper margin irregularly crenate, down-folded and roughly star-shaped. Colour reddish ochre to dark russet and the micropylar cup straw-coloured. Operculum and capitulum dark orange, the latter with the folded upper margin greyish brown.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFA95D3BFF52FC73A6BAFBBA.taxon	discussion	Remarks. This species was described from the ♀ and egg. Characteristic morphological traits such as the low but minutely dentate medioventral carina of the profemora, which is roughly central on the ventral surface of the profemur, indistinct pit-shaped praeopercular organ and bowl-like basal swelling of the subgenital plate as well as the distinctive egg morphology clearly place decoris in the genus Sadyattes, hence it is here removed from its original genus Phobaeticus Brunner v. Wattenwyl, 1907 (comb. nov.). Moreover, decoris is not unlikely to be the opposite sex of S. nigricornis (Redtenbacher, 1935) if the distribution in Borneo and the size relations of ♂♂ and ♀♀ within Sadyattes into account. The holotype of nigricornis was said to have a body length of 104.0 mm (Redtenbacher, 1908: 451), which is fairly small for the genus, thus the ♀ of this species should be an insect that measures about 15 – 16 cm in length. However, specimens of both sexes from the same locality or from captive breeding are needed for examination to prove or disprove the assumption that nigricornis and decoris are conspecific. Therefore, the ♂ is to be regarded as unknown. Since Seow-Choen (2016) only presented a picture of the eggs of decoris but no description, a formal description is here presented. Body length of ♀♀ (including subgenital plate) 158.0 – 165.0 mm.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFA95D3BFF52FC73A6BAFBBA.taxon	distribution	Distribution. NW-Borneo, Sarawak (Mount Serapi [LKCNH]; Bako National Park [SMSM]).	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFA85D34FF52FBDFA705F876.taxon	description	(Figs. 21 O – P)	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFA85D34FF52FBDFA705F876.taxon	materials_examined	Further material: 1 ♂: Kleiwegde Zwaas, Nias 1911; Pharnacia acanthopus Burm. K. Günther det. [RMNH]; 1 ♂: Penang Hills, 1800 - 2500 ft. S. S. Flower, 99 - 248 [NHMUK]; 1 ♀: West Malaysia, Perak, Tapah Hills ca. 800 m, local collector V. 2013 [FH, No. 1458 - 1]; 1 ♀: W. Indie [NHMM]. Differentiation. Both sexes of this distinctive species are morphologically closest to the type-species S. borrii Stål, 1875 from Java and Sumatra with which they share the distinctive pale cream-coloured protarsus of both sexes. Females of these two species also share the distinct posterolateral lobes of abdominal terga VIII-X, shape of the anal segment, which has the lateral margins deeply and almost semi-circularly excavated and the posterior margin broadly bi-lobed, as well as the rounded dorsal lobe of the probasitarsus. However, this species is more robust in overall shape with comparatively stockier legs, has abdominal segments III – VI distinctly swollen, the subgenital plate more acutely pointed apically and the dorsal lobe of the probasitarsus higher than in borrii. Males of enganensis are readily separable from those of borrii by the slightly stockier form, which includes notably shorter abdominal segments II – VI (4 x vs. 6.5 x longer than wide in borrii), more globose head which has the vertex notably more convex, more club-shaped apex of the abdomen with abdominal segments VIII and IX notably stronger inflated, shorter and gently arched vomer, having a dark brown medio-longitudinal streak on the tegmina, having all the dentations of the limbs more obtuse and not as distinctly black as in borrii and lacking the distinctively orange outer ventral carinae of the meso- and metafemora seen in borrii. From the known eggs of the genus, those of enganensis (Figs. 21 O – P) most closely resemble the eggs of incertus but differ by the punctured chorion surface, lack of the antero-ventral and dorsal and posteroventral impressions of the chorion, flattened polar-area, presence of two prominent sub-parallel longitudinal dorsal ridges and the much longer spearhead-shaped micropylar plate.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFA85D34FF52FBDFA705F876.taxon	description	Egg (Figs. 21 O – P). The following description is based on scaled colour photos taken of eggs from Peninsular Malaysia by Francis Seow-Choen (Singapore) the late Michael K. P. Yeh (Ipoh, Malaysia) and is meant to accomplish the treatment of this species. Unfortunately, no samples were secured (personal communication with F. Seow-Choen). Fairly large, chorion just slightly longer than high, somewhat flattened laterally and angularly oval in cross-section; polar-area flattened to slightyl concave and rectangular with rounded lateral surfaces in outline and surrounded by a ridge; dorsal and ventral surface each with a sub-parallel pair of distinct, longtitudinal ridges (those of the dorsal surface more prominent than those of the ventral surface. Capsule surface sparsely but distinctly punctured and with a weakly raised network of unevenly granular bulges; polar area somewhat raised and granulose in centre and with a circle of about ten pits. Micropylar plate placed between the two longitudinal ridges, somewhat displaced towards the posterior, rather large and almost two-thirds as long as chorion; spearhead-shaped with the posterior portion strongly narrowed and gradually tapering towards a narrow tip; outer amrgin somewhat raised the surface even, slightly concave and in centre with an oval, granular swelling. Micropylar cup positioned just above posterior narrowing of plate, flat and slightly bowl-shaped. Operculum slightly oval and flat with the outer margin somewhat raised. Capitulum large, conical and star-shaped in dorsal aspect with a distinct ridge running towards the centre from each of the six tips. Colour ranging from orangey ochre over russet to dark brown; the granules yellowish. Capitulum dark straw-coloured. Measurements [mm]: Length incl. operculum 4.8, length 4.2, width 3.5, height 3.8, length of micropylar plate 2.8.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFA85D34FF52FBDFA705F876.taxon	discussion	Remarks. This species was originally described from two ♂♂ from the island of Enggano, southwest off Sumatra in the collection of MSNG and Redtenbacher (1908: 451) stated “ This species differs from all other known species [of Pharnacia] by the almost club-shaped apex of the abdomen ”. Seow-Choen (2018: 452) presented a translation of Redtenbacher’s original description and illustrations of the lectotype ♂ in the collection of MSNG. Seow-Choen (2021) described the previously unknown ♀ and provided detailed figures of both sexes and the eggs based on specimens from Peninsular Malaysia, as well as figures of the ♂ paralectotype in MSNG and ♂ from Penang Hills, Peninsular Malaysia in the collection of NHMUK. Since this author missed out to give a description of the eggs, this is given above along with measurements based on scaled photos provided to the author by Francis Seow-Choen (Singapore) the late Michael K. P. Yeh (Ipoh, Malaysia). Seow-Choen (2021: 772) mentioned Psidium guajava (Myrtaceae), Mangifera indica (Anarcadiaceae) and Syzygium aqueum (Myrtaceae) as foodplants. Hennemann & Conle (2008: 194) have pointed out that the penultimate instar ♀ paralectotype of Phobaeticus sobrinus Brunner v. Wattenwyl, 1907 from the island of Nias was specifically distinct from the ♂ lectotype from Si- Rambe, Sumatra (both in MSNG) and is not even congeneric. This was confirmed by Seow-Choen (2018: 426), who provided a description of the ♀ as well as illustrations of both sexes and the intraspecific variability of Phobaeticus sobrinus, which actually is a fairly common species on mainland Sumatra. The now more comprehensive knowledge of S. enganensis and examination of a ♂ of enganensis from the Island of Nias in the collection of RMNH show the paralectotype of sobrinus to be a specimen of enganensis. Body length of the specimen is 125.0 mm. Females show considerable variability in colouration, which was illustrated by Seow-Choen (2021, figs. 727 – 732). The great majority of specimens appear to be whitish or cream coloured and variably flecked with different tones pf grey, brown and black. Occasionally the limbs possess a slight greenish to olive wash. So far, only one almost plain green specimen has become known, which only has some dark posterior spots on abdominal terga II-V. A ♀ from the Tapah Hills, Perak, Peninsular Malaysia an examined from photos taken by the late Michael K. P. Yeh (Ipoh, Malaysia) is basically pale olive with light greyish mottling. Body lengths ♂♂ 117.0 – 129.5 mm, ♀♀ (incl. subgenital plate) 158.0 – 183.0 mm.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFA85D34FF52FBDFA705F876.taxon	distribution	Distribution. Indonesia: Enggano Island (Bua Bua [MSNG – type locality]); Nias Island [MSNG, RMNH]. Peninsular Malaysia: Perak (Pulau Penang, Penang Hills 1800 – 2500 ft. [NHMUK]; Tapah Hills [FH, FSC]).	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFA65D32FF52FF2AA33AFECE.taxon	description	(Figs. 6, 21 C – D) Eucarcharus fallax Brunner v. Wattenwyl, 1907: 186. HT, ♀: Coll. Br. V. W., Luzon, Mus. Berlin; der. Br. v. W., Eucarcharus fallax; 11.332; Holotypus [NHMW, No. 310]. Bruner, 1915: 231. Brock, 1998: 28. [Type data] Otte & Brock, 2005: 136. Brock & Büscher, 2022: 559.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFA65D32FF52FF2AA33AFECE.taxon	materials_examined	Further material: Luzon: 1 ♂: Philippinen, South Luzon, II. 1986 local collector [FH, No. 1522 - 1]; 1 ♂: Philippinen, South Luzon Island, Prov. Quezon, Infanta Municipality, local collector [FH, No. 1522 - 2]; ♀: Philippinen, Luzon, leg. Jager [MNHU]; ♂ Philippinen. Luzon, leg. Jager " Lonchodes feruloides Westwood det. ", Nr. 3228 [MNHU]; 1 ♀: Vinai Pl., III. 10.1916 (W. Boettcher), Eucarcharus feruloides (Westw.) ♀ Det. Rehn 1934, Hebard Collection [ANSP]; 1 ♀: II. 14.1916 [ANSP]; ♀: Philippines (Ledyard), Eucarcharus feruloides (Westw.) ♀ Det. Rehn 1934, A. N. S. P. [ANSP]. Samar: 2 ♀♀, 1 ♂, 4 ♀♀ (immatures), 1 ♂ (immature n 4), 4 immatures: Philippines, Samar Id., Samar Prov., Hinubangan, San Isidro, San Isidro River, 90 – 200 m, 31. III. – 6. IV. 1997, leg. et Coll. R. a. Müller [coll. OZ, No. 330]; 1 ♂: Philippines, Samar Id., Samar Prov., Hinubangan, Arizona, Mining C., 280 m, 29. III. – 6. IV. 1997, leg. et Coll. R. A. Müller [coll. OZ, No. 330]. Panay: 1 ♂, 2 ♀♀, 1 ♀ (immature): Philippines, Panay Island, Mt. Madja-as, 1000 m, VIII. 2005, R. Cabale legit [MG, No’s 0222 - 2225]. No data: 3 ♀♀: no data (presumably Luzon, leg. Jager) [MNHU]. Doubtful data: 1 ♂: Dapi, Siargao, K. Günther det. [MHNG]. Differentiation. Females of this species show the morphologically closest affinity to the Luzonian S. feruloides (Westwood, 1859) but may be separated by the stockier overall form (mesothorax just 6.8 x longer than prothorax vs. 8 x in feruloides) and more robust limbs, which lack the distinctive dorso-apical tooth on the meso- and metatibiae seen in feruloides, have the cristate dorso-apical lobe of the meso- and metatibiae smaller and the basitarsi less distinctly crested dorsally. Moreover, distinction is rendered possible by the broadly triangular posteromedian notch of the anal segment (Fig. 6 I; shallow and concave in feruloides), the somewhat shorter and tapering subgenital plate, that protrudes beyond the tip of the abdomen by less than the combined length of the two terminal terga and has the apex obtusely triangular (Figs. 6 H – J; spatulate with the lateral margins roughly parallel-sided and the apex broad and roundly angular in feruloides) as well as the less distinct median node of the praeopercular organ. Males resemble those of the Luzonian S. matipuno sp. nov., Mindanaoan S. leytensis (Zompro, 1997) and S. maganda sp. nov. from the island of Mindoro, with which they share the general colour pattern and contrastive yellowish head (Figs. 6 D – E) but differ by the notably slenderer overall shape, more delicate limbs and trapezoidal anal segment, that strongly widens towards the posterior and has the outer angles somewhat protruded and triangular (rather parallel-sided or narrowing posteriorly in matipuno and maganda). From matipuno they may also be separated by the more globose head, which bears a distinct black postocular streak (a faint green streak in matipuno), green mesonotum that lacks the black lateral streaks seen in matipuno, shorter alae that do not reach the posterior margin of abdominal segment III and the distinctly annulated meso- and metatibiae. From the ♂♂ of leytensis they may be also be differentiated by the notably larger size (body length <95.0 mm in leytense), relatively larger head and more contrastive and defined postocular streak, shorter alae (reaching about half the way along abdominal segment IV in leytense), that have the costal region not distinctly striped longitudinally, lack of the black apex of the meso- and metafemora seen in leytense, as well as the smaller and narrower, rather triangular vomer (heart-shaped in leytense). Finally, from maganda these ♂♂ are also distinguishable by the relatively larger and much more globose head and presence of a black postocular streak (Figs. 6 E – F; ovoid and lacking a postocular streak in maganda), not longitudinally striped costal region of the alae, lack of the black lateral markings and blotches of abdominal terga II-VIII seen in maganda, lack of the black apex of the femora and much smaller and triangular vomer (broad with base almost circular in maganda). Although the colour pattern is essentially similar, the colouration in fallax is notably less striking and contrastive if compared to maganda. From the known eggs of Sadyattes, those of fallax (Figs. 21 C – D) most strongly resemble the ones of mindanaense sp. nov. but differ by the more fine-meshed network of ridges and bulges of the chorion, lack of the posteroventral indention seen in the eggs of mindanaense as well as the somewhat smaller and rather pear-shaped micropylar plate, which has the polar extension notably shorter (rather spearhead-shaped in mindanaense).	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFA65D32FF52FF2AA33AFECE.taxon	description	Description. Since this species has so far only been known from the ♀ and the very brief and insufficient original description was based on the strongly shrivelled and discoloured holotype only, a detailed redescription of the ♀ as well as a first description of the ♂ are here provided. Since all ♀♀ examined were provisionally stored in spirits, the colouration has certainly faded and care should be taken because the colour of live specimens may be different, for instance might also comprise green tones that are discoloured to yellow in the dried examples at hand. Measurements in table 3 below. ♀ (Figs. 6 A, F – J, N). Medium-sized (body length incl. subgenital plate 146.5 – 172.0 mm), form average for the genus with a short subgenital plate, that projects beyond the tip of the abdomen by less than length of anal segment (Figs. 6 H – J). Colour various tones of ochre, buff and brown and to a variable degree with darker brown and pale greyish to whitish speckles; the meso- and metafemora and tibiae to a variable intensity with three irregular, pale annulae. The holotype rather uniformly ochre with the head, prothorax and posterior portions of the mesonotum and median segment sepia and with a fine but uncontinuous dark brown medio-longitudinal line on the meso- and metanotum and median segment. Moreover, the holotype lacks the annulae of the legs seen in all the other specimens. Genae white in ventral portion and with a faintly indicated brown postocular streak (Fig. 6 F). Head (Fig. 6 F): Sub-oval but somewhat compressed dorso-ventrally and broadest just behind the eyes with the posterior portion narrowed; the vertex roundly convex with a shallow median impression in front, the coronal line fairly distinctly, the lateral furrow slightly indented. Frons with two closely spaced, small and C-shaped impressions between bases of antennae. Eyes rather small and just weakly projecting, round in outline and their diameter corresponding to 0.4 x length of gena. Antennae reaching about three-quarters the way along mesonotum; scapus small, rectangular and about 1.5 x longer than wide; pedicellus much shorter and round in cross-section. Thorax: Pronotum noticeably shorter and narrower than head, basically rectangular with the lateral margins gently concave and about 1.7 x longer than wide; the anterior margin concave, weakly bi-gibbose medially and followed by a furrow; the transverse median sulcus rather shallow, gently arched and almost expanding over entire width of notum. Mesothorax almost uniform in width and just slightly widened posteriorly, 6.3 x longer than prothorax. Mesonotum with a very fine medio-longitudinal carina and at anterior margin with a narrow transverse ridge, which has the median portion somewhat anterior-directed. Metanotum almost parallels-sided, 2.7 x longer than wide and somewhat less than half as long as mesonotum; the medio-longitudinal carina scarce. Meso- and metasternum with a weak and shallow medio-longitudinal keel. Abdomen: Median segment rectangular, 2.3 x longer than wide and scarcely more than half the length of metanotum. Segment II about 1.25 x longer than median segment, II – IV increasing in length, V as long as IV and VVII decreasing in length with VII shorter than all preceding; IV and V about 2.5 x longer than wide. II – VII just very slightly sub-uniform in diameter. Tergum V with a fairly distinct gibbose transverse posterior swelling (Fig. 6 N). Sterna II – VII smooth and each at best only with a slightly indicated medio-longitudinal carina in anterior portion. Praeopercular organ on VII formed by a short, granular longitudinal ridge some distance off the weakly medially notched posterior margin; the whole area unevenly verrucose and including the ridge black in colour (Fig. 6 J). Tergum VIII about half as long as VII and slightly widened posteriorly, IV roughly half as long as VIII and scarcely wider than long. Anal segment with a large, almost semi-circular posterolateral emargination (Fig. 6 H); the anterior portion of lateral margins somewhat deflexed and the posterior margin widely bi-lobed with a small triangular median indention (Fig. 6 I); dorsal surface with a shallow medio-longitudinal carina. Epiproct small, wider than long with a distinct medio-longitudinal keel and a posteromedian notch; filling the median gap of posterior margin of anal segment but not protruding beyond it (Fig. 6 I). Cerci fairly large, conical and tapering towards a narrow tip; roughly reaching to posterior margin of anal segment. Gonapophyses VIII up-curved, filiform and notably projecting under anal segment (Fig. 6 I). Subgenital plate scoop-shaped, obtusely keeled medio-longitudinally and with a prominent ledge-like keel along lateral margins in the basal one-third; apex obtuse, rounded and with the tip somewhat protruded, projecting beyond tip of abdomen by less than length of anal segment (Figs. 6 H – J). Legs: Moderately long and fairly stocky with essentially all carinae minutely denticulated; the meso- and metafemora very slightly incrassate sub-basally with the two outer ventral carinae somewhat deflexed. Profemora somewhat longer than mesothorax, mesofemora about four-fifths the length of mesothorax, metafemora almost reaching to posterior margin of abdominal segment IV and metatibiae reaching about halfway along abdominal segment VII. Dentations on anterodorsal carina of profemora prominent and serrate, those of the posteroventral carina slightly smaller and rather tooth-like; posterodorsal carina supplied only with very small denticles. The apical tooth of two outer ventral carinae of the mesofemora strongly enlarged and triangular, often of contrastive black colour. Medioventral carina of meso- and metafemora supplied with about 15 small spines; that of the meso- and metatibiae raised and forming a distinct, rounded lobe sub-basally. Meso- and metabasitarsus about as long as following three tarsal joints taken together with all three carinae minutely but densely denticulate; probasitarsus notably longer than following three tarsomeres combined and wholly unarmed; all with dorsal carina slightly raised. ♂ (Figs. 6 B – E, G, K – M). Of average size (body length 116.3 – 118.5) and typical form for the genus, with a sub-spherical, yellow head with a dark postocular streak, a black transverse band on the abdominal terga II-VI and short alae, that on average only reach only halfway along abdominal segment III (length 24.5 – 26.6 mm). Colour basically olive with the abdomen ochre to buff. Head straw-coloured to yellow with a rather slender and washed brown postocular streak that vanishes towards the posterior of gena (Fig. 6 E). Antennae brown with the basal eight or so joints black ventrally, the basal three joints blackish brown also dorsally. Prothorax greyish buff. Mesonotum occasionally with a slight brownish was dorsally. Tegmina and costal region alae greyish green to cream with the anterior margin marked by a broad creamy yellow band, which is interiorly bordered by a narrow brown longitudinal stripe; longitudinal veins green; anal fan hyaline with a slight greyish hue towards the outer margin and with all veins dark olive. Abdominal terga and sterna II-VI with a black transverse marking at posterior margin, which is broadest on II and becomes gradually narrower towards VI. Tergum VIII with a washed cream-coloured posterolateral marking, IX with a similar but more elongate marking. Profemora ochre with the compressed and curved base dark green, meso- and metafemora olive with the apex slightly brownish (Fig. 6 G). Protibiae orangey ochre, meso- and metatibiae greyish brown with three distinct, broad yellow annulae. Head (Figs. 6 D – E): Strongly globose, sub-spherical with vertex convex; broadest at the eyes and noticeably narrowing towards posterior; coronal line and lateral furrow fine and weakly indented. Vertex with two shallow indentions in front, the frons with a small, but deep transverse pit between bases of antennae. Eyes very large, projecting sub-spherically and their diameter corresponding to 0.66 x length of gena. Antennae reaching about halfway along abdominal segment III; scapus roundly rectangular and 1.6 x longer than wide, pedicellus roughly two-thirds the length of scapus and round in cross-section. Thorax: Pronotum basically as in ♀ but the lateral margins less concave and the anterior margin with the two median swellings much smaller (Figs. 6 D – E). Mesothorax slender, elongate and just weakly widened posteriorly; 7.5 x longer than prothorax. Mesonotum with a very fine medio-longitudinal line that is only clearly recognized in the anterior portion; the anterior margin with a low and almost straight transverse ridge (Figs. 6 D – E). Meso- and metasternum with a fairly distinct medio-longitudinal keel, which is most prominent in the anterior section of mesosternum. Tegmina elongate, spatulate in shape with the basal two-thirds gradually narrowing and the apical one-third rounded; central hump very shallow. Alae somewhat variable in length, reaching no more than two-thirds the way along abdominal segment III (occasionally just reaching to posterior margin of II). Abdomen: Segments II – V slightly increasing, VI and VII decreasing in length with VII shorter than all preceding, all except for VII uniform in diameter; V about 6.3 x longer than wide, VII somewhat constricted medially and about two-thirds the length of V. Terga V and VI with a slight posteromedian swelling. Sterna with a distinct medio-longitudinal keel. Tergum VIII distinctly trapezoidal in outline with posterior margin about 1.3 x wider than anterior margin, about 0.6 x length of VII and broadest of all segments. IX constricted medially with posterior portion noticeably narrower than anterior portion and scarcely longer than VIII; dorsal surface strongly convex longitudinally. Anal segment much shorter than IX, the anterior portion broad with lateral surface convex and the posterior portion narrowed and strongly tectate longitudinally; posterior margin separated by a distinct transverse furrow, the outer angles rounded and inflated (Figs. 6 K – L) and the ventral surfaces minutely denticulate and facing each other at an angle of no more than 40 °. Vomer small, roundly triangular in outline, longer than width across base, the lateral portions inflated and the central portion impressed longitudinally; terminal hook rather strong and up-curved (Fig. 6 M). Cerci round in cross-section with apex very slightly club-shaped, gently up-curved and projecting notably beyond anal segment. Phallus projecting notably over posterior and left lateral margin of poculum, rather broad and pointed apically (Figs. 6 K – M). The poculum of moderate size, bowl-shaped with base bulgy and rounded (Fig. 6 K), the posterior margin somewhat labiate, narrowed and scarcely reaching to posterior of tergum IX (Fig. 6 M). Legs: Long, slender and with all carinae minutely and fairly regularly dentate; the medioventral carina of meso- and metafemora narrow and only supplied with about ten (mesofemora) or fifteen (metafemora) very minute spines. Profemora notably longer than head, pro- and mesothorax taken together, mesofemora about as long as pro- and mesothorax combines, metafemora reaching roughly halfway along abdominal segment VI and metatibiae projecting beyond tip of abdomen by combined length of six terminal abdominal segments. Tarsi elongate, meso- and metabasitarsi as long as remaining joints taken together, probasitarsus slightly longer; all destitute of teeth. Egg (Figs. 21 C – D). Two eggs were extracted from the abdomen of the holotype in NHMW and can serve for a brief characterisation. Care should be taken however, because these eggs may not be fully developed. This in particular concerns to the sculpturing of the chorion as well as the colouration. Medium-sized, chorion longer than high, flattened laterally and distinctly oval in cross-section; polar-area rounded; on the dorsal surface these ridges indicate a pair of arched, longitudinal bulges. Anterior margin bulgy. Surface wholly covered by an irregular meshwork of shallow keels that leave impressed areas inbetween them; surface of the indented areas very minutely granular. Micropylar plate very small with the broaded anterior portion oval and somewhat narrowed anteriorly and the lower portion strongly narrowed with the raised outer margin fused. Internal surface concave and with a small bowl-shaped micropylar cup at lower end of widened anterior area. Operculum elliptical and noticeably arched with the dorsal and ventral portions downward-directed. Anterior portion with a small oval pit. Capitulum an irregular, mushroom-like protrusion that consists of about seven irregular ridges that fuse in centre. Colour yellowish ochre with the raised portions of the chorion straw-coloured. Micropylar plare, operculum and capitulum orangey brown. Measurements [mm]: Length incl. operculum 3.9, length 3.4, width 2.3, height 2.8, length of micropylar plate 1.3.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFA65D32FF52FF2AA33AFECE.taxon	discussion	Remarks. This species was originally described upon a unique ♀ from the island of Luzon, Philippines in the collection of NHMW. Since, the data label states “ Luzon, Mus. Berlin ” it is most likely that the specimen is part of the series of specimens labelled “ Luzon, leg. Jager ” in the collection of MNHU. All have provisionally been preserved in spirits and the right mid leg glued to the holotype in NHMW is apparently a hind leg, that does not belong to the specimen. Frequently, specimens of this species have been misidentified as S. feruloides (Westwood, 1859). In fact, fallax is the more commonly encountered species throughout Luzon and exhibits a remarkable distributional pattern within the Philippines, since it is also found in the Eastern Visayas (Samar) and Western Visayas (Panay). On the island of Luzon, fallax appears to be restricted to the southern half of the island, although many more records will be needed for a delimitation of its distribution. The collection of MHNG houses a ♂ that is stated to be from Dapa on the island of Siargao northeast of Mindanao. This distribution however is doubtful because it violates biogeographic data, that regard Siargao as part of the Mindanao Region, which essentially has no common species with the Greater Luzon Region or the Visayas. Therefore, this record is here excluded from the known distribution of fallax and deserves further evaluation.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFA65D32FF52FF2AA33AFECE.taxon	distribution	Distribution. Philippines. Luzon: “ Luzon ” [MNHU]; Province Quezon (Infanta Municipality [FH]). Samar: Province Samar (Hinubangan, San Isidro, San Isidro River, 90 – 200 m [OZ]). Panay: Province Antique (Mt. Madja-as, 1000 m [MG]).	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFA15D0CFF52FE0BA72AFEB6.taxon	description	(Figs. 7, 21 E – F)	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFA15D0CFF52FE0BA72AFEB6.taxon	materials_examined	Further material: 24 eggs: ex Zucht R. Krijns (Maastricht), 2012, Herkunft: Philippinern, O-Luzon, Infanta [coll. FH, No. 1524 - E]. Differentiation. The ♀ of this species is morphologically closes to the Luzonian S. fallax (Brunner v. Wattenwyl, 1907) but may be differentiated by the slenderer overall form (mesothorax 8 x longer than prothorax vs. 6.8 x in fallax) and more delicate limbs, which have a distinctive dorso-apical tooth on the meso- and metatibiae, rounded and cristate dorso-apical lobe on the meso- and metatibiae (Fig. 7 D) and more prominently lobed basitarsi. Moreover, the posteromedian notch of the anal segment is shallowly concave (Fig. 7 B; rather triangular in fallax), the relatively longer subgenital plate is spatulate with the lateral margins roughly parallel-sided and has the apex broad and roundly angular (Figs. 7 B – C; notably tapering in fallax) and the praeopercular organ has the median node less pronounced (Fig. 7 C). Males are easily recognized and distinguished from all other known Philippine congenerics by the distinctly black abdominal terga II – VI, black longitudinal lateral stripe of the mesonotum and mesopleurae and distinctly longitudinally striped costal region of the alae which are blackish brown with all longitudinal veins ochre. The very distinctive eggs (Frigs. 21 D – F) differ from all other known eggs of Sadyattes by their conspicuous five-pointed star-like shape if seen in lateral aspect, which is generated by an obtuse polar protrusion, a dorsal and ventral indention that each produce a dorsal and ventral protrusion as well as a median protrusion of the anterior margin.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFA15D0CFF52FE0BA72AFEB6.taxon	description	Description. Only brief characterizations of both sexes are provided below. ♀ (Fig. 7). The ♀ is so far only known from the unique holotype, for which a set of measurements is given below. Since the original description by Westwood (1859: 45) is fairly detailed only illustrations and a differential diagnosis are here provided. This very slender species is well recognized by the broad roundly angular and spatulate subgenital plate, that extends beyond the tip of the abdomen by almost the combined length of the two terminal abdominal terga, distinct rounded and cristate dorso-apical lobe of the meso- and metatibiae, enlarged triangular dorso-apical tooth of the meso- and metafemora and strongly dorsally cristate basitarsi. A specimen from Infanta photographed by Thierry Heitzmann (Philippines) matches very well with the holotype in all aspects including the colouration. Measurements of HT in NHMUK [mm]: Body (incl. subgenital plate) 165.5, body 160.0, pronotum 4.6, mesonotum 37.5, metanotum 15.7, median segment 12.0, profemora 44.0, mesofemora 36.0, metafemora 42.5, protibiae 51.5, mesotibiae 34.8, metatibiae 46.2, antennae -. ♂. The ♂ could only be examined from two colour photographs of a specimen from Infanta, taken by Thierry Heitzmann (Philippines). Unfortunately, no preserved sample is at hand for detailed examination and the measurements are not known. Thus, only a brief diagnosis based on the two available photos can be presented here. Form very slender for the genus with long and slender, minutely denticulated limbs and distinctive colouration. General colour buff with abdominal terga II – VI wholly black. Head pale cream with a faint washed greyish brown postocular and coronal streak; antennae blackish except for a brown scapus. Mesonotum and mesopleurae each with a black longitudinal line along the lateral margins. Tegmina and costal region of alae with anterior margin broadly white; tegmina with a broad washed blackish brown medio-longitudinal streak; alae distinctly striped longitudinally, being blackish brown with all longitudinal veins ochre. Abdominal terga VII – X irregularly flecked with dark brown. Profemora russet with base dark green; meso- and metafemora green with apical one-third gradually turning to drab. All tibiae drab with a weakly indicated sub-basal pale transverse band. Head elongate-ovoid with vertex moderately convex; notably wider than pronotum. Antennae reaching to posterior margin of abdominal segment II. Mesothorax very elongate and slightly more than 6 x the length of prothorax. Alae reaching to posterior margin of abdominal segment III. Abdominal tergum V with the typical posteromedian swelling seen in other species of the genus. Anal segment notably widening towards posterior and trapezoidal in dorsal aspect; the posterior margin weakly notched medially. Profemora longer than head, pro- and mesothorax taken together. Metafemora reaching to posterior margin of abdominal segment V and metatibiae projecting strongly beyond tip of abdomen. Anterodorsal carina of meso- and metatibiae weakly rounded and cristate apically. Tarsi slender with all basitarsi longer than following tree tarsomeres combined. Egg (Figs. 21 E – F). Large and of a very distinctive shape. Overall shape in lateral aspect resembling a five-pointed star. Chorion strongly compressed laterally and slightly higher than long; dorsal and ventral surfaces indented medially with the anterior and posterior portions bulging; the anterior margin raised medially and distinctly lowered dorsally and ventrally and the polar-area with a prominent obtusely rounded central protrusion. Surface minutely and very densely granular and wholly covered by an irregular and rather incontinuous network of shallow bulges that leave impressed areas inbetween them. Micropylar plate small, elongate and roundly rhomboidal with the anterior end obtuse and the posterior portion gradually narrowing towards a narrow tip.; outher margin somewhat inflated and the internal area strongly concave in anterior half; micropylar cup very small, bowl-like and roughly in centre of plate at lower end of indented area. Operculum notably indented into chorion, roughly elliptical in shape with the lateral margins almost parallel-sided; strongly arched with the dorsal and ventral portions noticeably lowered. Surface with many radially directed ridges and dorsal portion with a large elongate impression. Capitulum an irregular and complex protrusion that essentially consists of four ridges that are aaligned at right angles to each other and merge in centre. Colour greyish ochre with the raised bulges slightly darker in colour. Operculum blackish brown; the capitulum black dorsally and reddish brown at the base. Micropylar cup black. Measurements [mm]: Length incl. operculum 4.7, length 4.1, width 2.4, height 3.5, length of micropylar plate 1.8.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FFA15D0CFF52FE0BA72AFEB6.taxon	discussion	Remarks. Westwood (1859: 45) described this species upon two specimens, an adult ♀ from the Philippines and a penultimate instar ♀ from Java, both of which are in the collection of NHMUK. The description and illustrations were based on the Philippine specimen, why Seow-Choen (2023: 516) designated this as the lectotype to ensure stability of Westwood’s taxon. Furthermore, Seow-Choen stated that already from geographic aspects alone it was most unlikely that the paralectotype from Java is the same species and presented detailed illustration of both type specimens (Seow-Choen, 2023: 516, Figs. 468 – 469). Indeed, the immature paralectotype shows several morphological characters that differ from the Philippine lectotype, which include the slightly stockier overall shape, more bi-lobed posterior margin of the anal segment and notably stronger limbs, which have all the dentations more pronounced with the mesofemora and four posterior tibiae lacking the dorsal apical lobe seen in the lectotype of feruloides. Comparison with other Javanese specimens that have been examined in course of this study shows the specimen to be the type species of the present genus, S. borrii Stål, 1875. This species is similar to S. fallax (Brunner v. Wattenwyl, 1907), which is widely distributed throughout the southern half of Luzon, and thus almost all specimens of fallax that were recorded from Luzon by former authors have been misidentified as feruloides (see comments on fallax above). Another reason for these frequent misidentifications has certainly been, that the very distinctive ♂ of feruloides was not known previously. In fact, the much slenderer feruloides is much less common than fallax and hitherto only known from the unique ♀ holotype. The ♂ is here briefly described the and egg is described and illustrated for the first time to render possible a proper distinction of feruloides from related species. Stock from Infanta, Quezon Province was introduced to Europe by Rob Krijns (Maastricht, Netherlands) in 2012 based on eggs laid by the abovementioned specimens collected and photographed by Thierry Heitzmann (Philippines). Breeding has however proven very difficult and did not succeed. Distribution. Philippines (type locality [NHMUK]). Luzon: Province Quezon (Infanta Municipality [FH]).	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF9F5D09FF52FAE4A767FD96.taxon	description	(Figs. 8 – 9, 21 M – N)	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF9F5D09FF52FAE4A767FD96.taxon	materials_examined	Further material: Peninsular Malaysia: 2 ♀♀: Perak (Malacca), A. Grubauer leg., B. Jachan vend. 15. XII. 1901 [ZMUH]; 1 ♂: Perak II. - III. 1900 (Kwala-Kangsar) B. Jachan vend. 15. VII. 1900 [ZMUH]; 1 ♂: Malay Penin., Pinding, Tanjong, Hauti, light-House, 26. VIII. 1929, F. N. Chasen [NHMUK]; 1 ♀: Malacca, Selangor, Ulu Gombak, N. C. E. Miller 1928 [NHMUK]. Nicobar Islands: 2 ♀♀, 2 eggs: Nicobar Islands, Kamorta Id., I. 2001 [coll. PDB, to bedeposited in NHMUK]. Differentiation. Females (Figs. 8 A – B) of this large species most closely resemble those of the Bornean S. decoris (Seow-Choen, 2016) but may be separated by the larger size (body length including subgenital plate> 189.0 mm) and somewhat slender overall shape and limbs, more elongated and notably more flattened head (Figs. 8 C – D), larger, black and rather spinose than serrate dentations of the extremities, longer subgenital plate, which extends beyond the tip of the abdomen by more than the combined length of the terminal three abdominal terga (Figs. 8 F-H), and the more distinct praeopercular organ (Fig. 8 H). The ♂♂ (Fig. 9 A) most closely resemble those of S. enganensis (Redtenbacher, 1908) from Peninsular Malaysia, Sumatra and the islands of Enggano and Nias east of Sumatra, S. borrii Stål, 1875 from Java and Sumatra and the Bornean S. annulatus (Redtenbacher, 1908). From the two first species they are however easily separable by the much more elongated and flattened head (Figs. 9 B – C), which is notably longer than the prothorax, and the longer alae, which almost reach to the posterior margin of abdominal segment IV. From enganensis they moreover differ by the slenderer shape, less inflated abdominal segments VIII – IX, as well as the more numerous and acutely pointed black teeth of the mid and hind legs (obtuse and dark green in enganensis). From borrii these ♂♂ also differ by the dark medio-longitudinal streak of the tegmina and costal region of the alae, not distinctly contrastive pale cream-coloured protarsi, not orange outer ventral carinae of the meso- and metafemora and smaller poculum, which lacks the obtuse basal protrusion seen in borrii (Fig. 9 D) and much slenderer and elongated, irregularly parallel-sided vomer (Fig. 9 F; triangular in borrii). The long alae are shared with annulate, but in this species the alae reach as far back as to abdominal segment V, the head has the vertex weakly convex, the medio-longitudinal carina of the mesosternum is much less pronounced, the meso- and metafemora are plain green except for a blackish apex, and the vomer is more distinctly and gradually tapering towards the terminal point and lacks the median widening seen in incertus. From the known eggs of the genus, those of incertus (Figs. 21 M – N) most closely resemble eggs of enganensis but differ by the minutely verrucose chorion surface, presence of antero-ventral and dorsal and posteroventral impressions of the chorion, impressed polar-area, lack of the sub-parallel longitudinal dorsal ridges and the much shorter round micropylar plate.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF9F5D09FF52FAE4A767FD96.taxon	description	Description. The following description of the previously unknown ♂ is based on the unique example in the collection of ZMUH (Fig. 9 A). The specimen is fairly complete except for the left mid leg, tips of the metatibiae and both metatarsi as well the right antenna and cerci. The colouration does not seem to be notably diverted by preservation or a preliminary storage in ethanol. Measurements in table 4 below. ♂ (Fig. 9). Large (body length 142.0 mm), form slender for the genus, with an elongated head and fairly long alae that reach about halfway along abdominal segment IV (length 58.0 mm). Meso- and metathorax greyish olive, rest of body essentially drab to ochre, the abdomen becoming brown towards the apex. Meso- and metapleurae with a washed black line along lower margin. Head buff, the genae cream-coloured ventrally and with a faint and rather narrow cream-coloured stripe anteriorly, that roughly runs at upper margin of compound eye (Figs. 9 B-C). Antennae dark reddish brown. Pronotum slightly russet. Tegmina and alae cream-coloured with a broad blackish medio-longitudinal streak and the anterior margin broadly light cream (possibly whitish when alive); anal fan hyaline with veins pale brown. Abdominal terga V-VII with an elongate washed cream anterolateral marking and the posterior margin broadly black. Terga VIII and IX with a whitish medio-lateral marking. Sterna II-VII with a washed dark medio-longitudinal streak, black posterolateral angles and a bold white posteromedian spot. Poculum brown basally with the remainder straw-coloured (Fig. 9 F). Profemora green at the base (Figs. 9 B – C), otherwise irregularly annulated and flecked with dark brown and pale cream, the apex black; protibiae mostly black with four whitish annulae. Meso- and metafemora green throughout the basal half, then gradually becoming buff, then with a washed greyish ochre transverse band and the apex black; meso- and metatibiae black with three broad, dark yellow annulae. All dentations of the limbs black. Basitarsi whitish to pale cream with the apex black. Head (Figs. 9 B – C): Elongate-oval, somewhat depressed dorsoventrally, only 1.5 x longer than wide, broadest at eyes and notably narrowing towards posterior with the posterior portion of capsule narrowed and somewhat angular in lateral aspect; vertex weakly convex with coronal line weakly and lateral furrow distinctly impressed. Frons with a closely spaced pair of shallow impression between the bases of the antennae. Eyes very large, projecting more than hemispherical and their diameter corresponding to almost 0.7 x length of gena. Antennae long and reaching to anterior of abdominal segment IV; scapus slender and slightly narrowed towards base; pedicellus round in cross-section; following joints slightly and irregularly unequal in length and essentially first increasing, then decreasing in length. Thorax: Pronotum shorter and noticeably narrower than head, basically rectangular in outline and about 1.8 x longer than wide, the lateral margins gently convex; the anterior margin somewhat inflated and with a pit near each outer angle of segment; transverse median sulcus distinct, notably impressed, very weakly arched and almost reaching to lateral margins of segment; the medio-longitudinal line slightly indented (Figs. 9 B – C). Mesothorax very slender, elongate and uniform in diameter with only a slight widening in posterior portion and at anterior margin; almost 7.2 x longer than prothorax. Mesonotum with a distinct transverse bulge at anterior margin (Fig. 9 C) and a weakly indicated medio-longitudinal line. Meso- and metasternum with a fine and low medio-longitudinal carina, which however is quite prominent in the anterior portion of the mesosternum. Tegmina spatulate with basal half gradually narrowing and the central hump shallow and weakly convex. Alae reaching about halfway along abdominal segment IV. Abdomen: Abdominal segment II and III equal in length, IV – VII slightly gradually decreasing in length but all uniform in diameter; III about 6 x and VII only 5 x longer than wide. Terga V and VI with an obtusely rounded posteromedian swelling, which is comparatively more pronounced on VI. Sterna II – VII all with a shallow medio-longitudinal carina. Tergum VIII slightly trapezoidal in outline, gradually widening towards the posterior and about half as long as VII; IX just scarcely shorter than VIII and somewhat constricted medially. Anal segment strongly tectate with posterior half distinctly narrowing (Fig. 9 E); the lateral surfaces slightly gibbose pre-basally; posterior margin somewhat inflated with the ventral surfaces minutely denticulate and distinctly facing each other (Fig. 9 G). Vomer very elongate, somewhat irregularly digitiform with an up-curved terminal point; the ventral surface with a deep medio-longitudinal furrow and the lateral margins roundly inflated (Figs. 9 F – G). Phallus enlarged and projecting over posterior margin of poculum (Figs. 9 D – G). Cerci missing in the unique specimen at hand. Poculum rather small, bowl-shaped with the base moderately bulgy and angular (Fig. 9 D) and the remainder portion with a distinct medio-longitudinal keel; posterior margin narrowly rounded and hardly reaching posterior of tergum IX (Fig. 9 F). Legs: Long, slender and with all carinae rather minutely but acutely dentate; the ventral dentations basically more pronounced that those of the dorsal carinae and the latter somewhat erratic in their distribution. Medioventral carina of meso- and metafemora only supplied with a few very minute spines. Profemora notably longer and mesofemora about as long as head, pro- and mesothorax taken together, metafemora almost reaching posterior margin of abdominal segment V and metatibiae projecting beyond tip of abdomen. Probasitarsi strongly elongated, unarmed and notably longer than all remaining joints taken together. Mesobasitarsi minutely denticulate ventrally and roughly equal in length to combined length of remaining tarsomeres.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF9F5D09FF52FAE4A767FD96.taxon	discussion	Remarks. This species was originally described based on a ♀ from the island of Banka in the Strait of Malacca some kilometres off the north-eastern coast of Sumatra. The unique holotype has provisionally been preserved in spirits, why it has lost its natural colours and has most of the body but the abdomen in particular considerably shrivelled. The notably larger holotype ♀ of grubaueri from Kuala Kangsar, Perak, Peninsular Malaysia in the collection of NHMW is very well preserved. While an egg could be removed from the ovipositor of the holotype of grubaueri, some eggs were extracted from the abdomen of the holotype of incertus in order to provide certainty about its taxonomic position. Since, nor the ♀♀ neither the eggs provided any significant differences from each other Hennemann & Conle (2008: 195) synonymised grubaueri under incertus. This was doubted by Seow-Choen (2018: 421), who re-established grubaueri as a valid species, and postulated that the holotype of incertus was smaller, slenderer and has the posterior margin of abdominal sterna V – VII rounded and slightly raised posteromedially, whereas it is medially incised and bi-lobed in the holotype of grubaueri as well as further specimens the author had examined from Pahang and Johor, Peninsular Malaysia. Re-examination of the holotype of incertus and comparison with the Malayan ♀♀ listed above and taking the shrivelled condition of the type of incertus into account leaves no doubt incertus and grubaueri are the same species. The difference in size is not unusual between specimens of an individual species from the mainland and circumjacent islands, and within an expectable range. As already stated by Hennemann & Conle (2008: 197) also the eggs of both taxa do not show any significant differences that would separate them as distinct species. Therefore, Redtenbacher’s grubaueri is here re-synonymised with incertus (rev. stat.). Seow-Choen (2021: 735) assumed it was highly likely that grubaueri was the opposite sex of Phobaeticus pinnipes (Redtenbacher, 1908), however these two species are not closely related, since the latter belongs in Clitumninae: Pharnaciini and is not a member of Platycraninae: Stephanacridini. Moreover, the collection of ZMUH houses a ♂ from Kuala-Kangsar, Perak, Peninsular Malaysia, the type-locality of grubaueri, which undoubtedly represents the previously unknown ♂ of the present species. In addition to this ♂ there is another Malayan example in the collection of NHMUK. A description of the ♂ of incertus is here presented based on these two specimens and detailed illustrations of both sexes are presented. Hennemann & Conle (2008: 197, Figs. 167 – 168) have provided a description and illustrations of the egg but a photographic documentation of the eggs is here presented to accomplish the treatment of this species.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF9F5D09FF52FAE4A767FD96.taxon	distribution	Distribution. Banka Island (northeast off Sumatra) [MNHU – type-locality]. Peninsular Malaysia: Perak (Kuala Kangsar [NHMW, ZMHU]); Selangor: (Ulu Gombak [NHMUK]); Penang (Tanjong [NHMUK]); Pahang [Seow-Choen, 2018: 421]; Johor [Seow-Choen, 2018: 421]. Nicobar Islands: Kamorta Island [PDB (NHMUK)].	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF9A5D04FF52FD33A6B1FEB6.taxon	description	(Figs. 10 – 11, 21 G – H, 24)	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF9A5D04FF52FD33A6B1FEB6.taxon	description	Otte & Brock, 2005: 155. Gottardo, 2008: 11, Figs. [Life history, description of egg] Bollens, Krijns & Bresseel, 2010 b: 23. Gottardo & Vallotto, 2014: 258. [Description and microanatomy of ♂] Harman, 2015: 26. [Note on culture stock] Brock & Büscher, 2022: 559, Fig.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF9A5D04FF52FD33A6B1FEB6.taxon	materials_examined	Further material: 17 ♀♀, 4 ♀♀ (immature): Coll. I. R. Sc. N. B., Philippines, Mindanao, Leg: R. Cabale 2011, Gift: J. Bresseel, IG: 32.386 [RBINS]; 1 ♂: Philippinen, Mindanao Id., Mount Busa, III. 2007, leg. R. Kabale [coll. FH, No. 0601 - 1]; 1 ♀: ex Zucht: Ingo Morisse 2009, Herkunft: Philippinen, Mindanao, Mount Apo, leg. Bresseel et al. 2008 [coll. FH, No. 0601 - 9]; 50 eggs: ex Zucht: M. Gottardo (Rovigo, Italien) 2007, Herkunft: Philippinen, Mindanao Id. [coll. FH, No. 0601 - E]; 2 ♀♀: Philippinen, Mindanao Island, Mount Matutum, 2007 [0601 - 2 & 3]; 1 ♀: Philippinen, Mindanao Island, South Cotabato Province, Mount Apo, local collector III. 2010 [coll. FH, No. 0601 - 4]; 1 ♀: Philippinen, Mindanao Id., South Cotabato Prov., Kidapawan, ca. 30 m, via I. Lumawig, local collector IX. 2011 [coll. FH, No. 0601 - 5]; 2 ♀♀: Philippinen, Mindanao Island, Prov. Cotabato, Municipality Magpet O of Davao, local collector X. 2011 [coll. FH, No’s 0601 - 6 & 7]; 1 ♀: Philippinen, Mindanao Island, Prov. Davao del Sur, Talomo Mountain Range, local collector V. 2011 [coll. FH, No. 0601 - 8]; 1 ♀: Philippines, Panay Island, Mt. Madja-as, 1000 m, VIII. 2005, R. Cabale legit [MG, No. 0226]. Differentiation. The very distinctive ♀♀ of this species are easily separated from all other members of the genus by the very stocky shape and broadened body, having the mesonotum strongly laterally dilated with the lateral margins forming a broadly rounded flange and the lateral margins of abdominal terga II – VII roundly deflexed, and all body segments relatively shorter with the mesonotum less than 4 x the length of the pronotum (Figs. 10 A – B). Moreover, there is a very distinct medio-longitudinal carina on the meso- and metasternum, which is also present but notably less pronounced in the other species. The strong legs of these ♀♀ are only seen to a similar degree in matipuno sp. nov., but in this species the spines on the medio-ventral carina of the meso- and metafemora are much stronger. Males (Fig. 11 A – B) come morphologically closest to those of fallax and the possibly locally sympatric mindanaense sp. nov. but may be easily separated from both species by the relatively smaller and less globose head that lacks a dark postocular streak (Fig. 11 C), having the main longitudinal veins of the alae marked by dark brown stripes (almost unicoloured in fallax and mindanaense sp. nov.), longer and much slenderer, gradually tapering vomer and black apex of the meso- and metafemora (Fig. 11 D). The eggs also most closely resemble those of these two species but can be differentiated by the more globose overall shape and differently shaped capitulum, which bears an orangey irregularly cross-shaped bulge (Figs. 21 G – H).	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF9A5D04FF52FD33A6B1FEB6.taxon	discussion	Remarks. This species was originally described in the genus Hermarchus Stål, 1875 based upon a unique ♀ holotype from the island of Leyte, Philippines, which unfortunately is still deposited in the personal collection of its author (coll. OZ). The original generic placement is apparently wrong, since Hermarchus is restricted to Melanesia (Hennemann & Conle, 2006; Gottardo, 2008; Gottardo & Vallotto, 2014). When Zompro (1997) described leytensis, he linked his new species to the New Guinean Macrophasma lyratus (Redtenbacher, 1908) due to an overall resemblance of the ♀♀, a species that previously was in the genus Hermarchus. Hennemann & Conle (2006) have accommodated all New Guinean taxa formerly in Hermarchus in the newly described genus Macrophasma Hennemann & Conle, 2006. The egg was described in detail and illustrated by Gottardo (2008: 11, fig. 2) and a superb and exhaustively illustrated macro- and microanatomical study of the previously unknown ♂ was presented by Gottardo & Vallotto (2014). Despite the striking ♀ morphology, which includes a very stocky general shape, a remarkably dilated mesonotum and laterally dilated abdominal terga, the morphology of the ♂ and egg as well as the distribution in the Philippines clearly links leytensis with the genus Sadyattes. The ♂ is a very typical representative of that genus and the egg strongly resembles that of the Philippine S. fallax (Brunner v. Wattenwyl, 1907) and S. mindanaense sp. nov. Therefore, leytensis is here removed from Hermarchus and transferred to Sadyattes (comb. nov.). The characteristic traits of Hermarchus and Macrophasma, that morphologically differentiate these genera from Sadyattes are highlighted in the keys to the genera of Stephanacridini above. Illustrations of both sexes and the eggs are presented herein. Information on the life cycle, captive breeding and dietary was provided by Gottardo (2008), who stated Hypericum spp. (Hypericaceae), Quercus spp. (Fagaceae), Psidium guajava (Myrtaceae) and Rubus ulmifolius (Rosaceae) to be accepted as alternative foodplants in captivity in Europe. In addition to these, also Gaultheria shallon (Ericaceae), Corylus avellana (Betulaceae), Rubus idaeus (Rosaceae) and Fagus sylvatica (Fagaceae) have proven to be taken as alternative food (personal communication with Bruno Kneubühler). Two culture stocks have been introduced from Mount Apo, Mindanao, a parthenogenetic stock in 2007 and a second sexual stock in 2009, but unfortunately, both stocks were lost after only a few generations. Body lengths ♂♂ 72.3 – 93.9 mm, ♀♀ (incl. subgenital plate) 127.8 – 149.0 mm.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF9A5D04FF52FD33A6B1FEB6.taxon	distribution	Distribution. Philippines. Leyte (Mahaplag, Hilusig, Mount Balocaue, 700 m [OZ – type locality]. Panay (Mount. Madja-as, 1000 m [MG]). Mindanao: “ Mindanao ” [RBINS]; Province South Cotabato (Mount Apo [FH, MG, OC, MCFS, MSNG]; Mount Matutum [FH, OC]; Kidapawan [FH]); Province Cotabato (Magpet [FH]); Province Davao del Sur (Talomo Mountain Range [FH]); Province Sarangani (Mount Busa [FH]).	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF975D01FF52FED3A28DFF52.taxon	description	(Figs. 12 – 13) HT, ♂: Philippinen, Mindoro Isl., Mt. Halcon, leg. Mohagan 27.4.1996 [RBINS, ex coll. coll. FH, No. 0278 - 1]. PT, ♂: Philippinen, Mindoro Id., Prov. Mindoro Oriental, Naujan Munip., Arangin, local collector, 17. VII. 2004 [coll. FH, No. 0278 - 2]. PT, 9 ♀♀: 15 ♂♂: Philippinen, Mindoro Island, Prov. Oriental Mindoro, Mount Halcon, leg. N. Mohagan 6.2006 [coll. OC, No’s 0178 - 1 to 24]. PT, 8 ♀♀: 31 ♂♂: Philippinen, Mindoro Island, Prov. Oriental Mindoro, Mount Halcon, leg. N. Mohagan 6.2004 [coll. OC, No’s 0178 - 25 to 63]. Differentiation. The pretty ♂ (Fig. 13) of this new species is easily recognised by its distinctive and contrastive colouration, which includes an unicoloured greyish green head, glaucous to greyish blue thorax, meso- and metafemora and bases of the profemora as well as an orange abdomen, profemora and tibiae, the latter of which are distinctly annulated with black. The lack of a distinct and defined postocular streak (Figs. 13 M – N), longitudinally striped costal region of the alae and black apex of the meso- and metafemora are shared with leytensis, but in this new species the costal region of the alae is dark brown with the longitudinal veins pale cream-coloured (Fig. 13 O; reverse in leytensis) and the vomer is much shorter, broader and basically oval in outline with a small terminal point (Figs. 13 I, L). Besides, the ♂ of this new species also resembles that of S. fallax (Brunner v. Wattenwyl, 1907) from Luzon and Samar but is distinguishable by the relatively smaller and less globose head (Figs. 13 M – N) and lack of the definite black postocular streak seen in fallax, longitudinally striped costal region of the alae (Fig. 13 O), presence of the distinctive but variable black lateral markings and blotches of abdominal terga II-VIII (Figs. 13 E – F), black apex of the femora (Fig. 13 P) and much larger broadly oval base of the vomer (Figs. 13 I, L; smaller and triangular in fallax). The ♀ (Fig. 12) is morphologically closest to that of S. panayense sp. nov. from the island of Panay with which it shares the overall shape and long, lanceolate subgenital plate (Figs. 12 G – I). It may however be separated from panayense by the somewhat more globose head and black postocular speckles (Fig. 12 D; both missing in maganda), more incrassate basal portion of the meso- and metafemora (Fig. 12 E), distinctly serrate rather than spinose ventral dentations of the meso- and metafemora, which are comparatively more pronounced but fewer in number and distinctly black (Fig. 12 E), large triangular pical tooth of the two outer ventral carinae of the meso- and metafemora (Fig. 12 F), as well as the broader posterior margin of the anal segment (Fig. 12 H). In several aspects, the ♀ of this species also resembles that of S. matipuno sp. nov. from the island Luzon, e. g. the somewhat incrassated basal portion and red ventro-basal area of the meso- and metafemora. However, maganda has the meso- and metafemora comparatively much slenderer and in general is much less stocky than matipuno and has the anterior of the mesothorax not conspicuously constricted like in matipuno.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF975D01FF52FED3A28DFF52.taxon	etymology	Etymology. The name maganda (filipino = pretty, nice, beautiful) refers to the vibrant colouration of the ♂ of this new species. Feminine.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF975D01FF52FED3A28DFF52.taxon	description	Description. The considerable colour variability of this species is summarised in the species description below and not provided separately because no noteworthy morphological variability can be seen in the numerous specimens at hand. ♀ (Fig. 1 2). Medium-sized (body length incl. subgenital plate 140.0 – 159.5 mm), form typical for the genus, with a long and lanceolate subgenital plate and basally incrassate meso- and metafemora that bear very prominent saw-like teeth on the two outer ventral carinae. Colour remarkably variable (Figs. 12 A – C) and ranging from olive over various tones of ochre or buff to brown. Occasionally specimens possess irregularly dispersed white dots on the meso- and metanotum or more rarely have a bold more or less continuous white medio-longitudinal streak running along the meso- and metanotum and median segment (Fig. 12 C). Often a pair of washed blackish posterolateral markings is seen on the meso- and metanotum (Fig. 12 C). Meso- and metapleurae occasionally flecked with black and light grey or white. Area surrounding the compound eye blackish and genae with a variable and washed diagonal but often uncontinuous black postocular streak that runs from the eye towards the posteroventral end of head capsule (Fig. 12 D). Antennae greenish brown to drab and blackish ventrally except for the scapus and pedicellus. Base of subgenital plate often notably darker in colour than rest of body, sometimes black (Fig. 12 I). Meso- and metafemora and tibiae each with two more or less distinct, sometimes very faint lighter coloured annulae; all dentations of legs tipped with black. Base of meso- and metafemora red ventrally and ventro-laterally (Fig. 12 E). Basitarsi with base lighter in colour. Head (Fig. 12 D): Sub-globose, about 1.3 x longer than wide and broadest just behind the eyes; vertex convex with only the lateral furrow slightly indented and the anterior portion very shallowly concave; front with a rather shallow and slightly arched transverse impression between bases of antennae. Eyes moderately projecting and their diameter corresponding to about half the length of gena. Antennae reaching three-quarters the way along mesothorax; scapus compressed dorsoventrally, rectangular and about 1.6 x longer than wide, pedicellus round in cross- section and much shorter than scapus. Thorax: Pronotum noticeably shorter and much narrower than head, almost rectangular in outline and 1.5 x longer than wide; near anterolateral angles with a V-shaped pit; the transverse median sulcus rather shallow, gently arched and expanding almost over entire width of segment; the lateral margins with a slight indention at transverse sulcus. Mesothorax 5.5 x longer than prothorax and slightly sub-uniform in diameter with the anterior one-third slightly constricting. Mesonotum with a very faint medio-longitudinal line and only a low and obtuse, gently arched ridge at anterior margin. Metanotum rectangular, 2.4 x longer than wide and about 0.4 x length of mesonotum. Meso- and metapleurae with a shallow medio-longitudinal carina. Medio-longitudinal carina of meso- and metasternum almost indiscernible. Abdomen: Median segment weakly trapezoidal, 1.5 x longer than wide and about 0.6 x length of metanotum. Segments II – V slightly increasing, VI and VII decreasing in length; V about 2 x longer than wide, VII roughly 0.8 x length of V. Segments slightly sub-uniform in diameter with II-V somewhat inflated and VII narrower than all preceding. Sterna II – VII with a fine but fairly distinct medio-longitudinal carina; praeopercular organ formed by a slightly swollen area of irregular, roughly longitudinally directed ridges, which have a somewhat enlarged Y-shaped bulge in centre; this region blackish brown and shiny. Tergum VIII slightly narrowing towards the anterior, 1.75 x longer than wide and about half as long as VII; IX almost quadrate in dorsal aspect about three-fifth the length of VIII. Anal segment scarcely longer than IX and narrowing towards the posterior with a fine medio-longitudinal keel and somewhat tectate posteriorly; the posterior margin narrowed, rounded and with a minute median indention; lateral margins very weakly concave in posterior half. Epiproct broad, transverse and scarcely projecting over posterior margin of anal segment (Fig. 12 H). Cerci fairly small, conical and almost reaching to tip of anal segment. Gonapophyses VII hidden underneath anal segment. Subgenital plate somewhat variable in length, lanceolate, bulgy in basal portion with the elongated apical portion naviculate and with a more or less acute tip; projecting beyond tip of abdomen by more than length of terminal two terga taken together (usually much longer although and extreme cases projecting by more than combined length of terga VIII – X; Figs. 12 G – I). Legs: Of moderate length and stock for the genus with the meso- and metafemora notably incrassate in basal one-third with the two outer ventral carinae expanded; all carinae dentate; essentially all ventral dentations notably more pronounced than dorsal dentations. Profemora about as long as pro- and mesothorax combined, mesofemora slightly shorter than mesothorax, metafemora reaching roughly reaching to posterior margin of abdominal segment Iv and metatibiae reaching posterior of VII. Dentations on posteroventral carina of profemora more triangular in shape, slightly more numerous, more closely spaced and more distinctly black that the lower and rather saw-like teeth of anterodorsal carina; posterodorsal carina supplied with about 20 small denticles. Dentations of two outer ventral carinae of meso- and metafemora very prominent, acutely triangular, almost wholly black and closely spaced; teeth of the dorsal carinae small, less in number and irregularly spaced (Fig. 12 E). Medioventral carina of meso- and metafemora supplied with some rather small spines. The apical tooth of two outer ventral carinae noticeably enlarged (mesofemora in particular; Fig. 12 F). Meso- and metabasitarsus somewhat longer than following three tarsomeres taken together with the two ventral carinae dentate; probasitarsus unarmed and slightly longer than all remaining joints combined. Measurements of ♀♀ PT [mm]: Body (incl. subgenital plate) 140.0 – 159.5, body 128.9 – 146.2, pronotum 5.3 – 5.6, mesonotum 25.8 – 30.2, metanotum 11.8 – 13.0, median segment 7.3 – 7.8, profemora 30.9 – 38.1, mesofemora 23.9 – 28.1, metafemora 29.0 – 35.2, protibiae 33.8 – 42.6, mesotibiae 22.3 – 26.5, metatibiae 29.1 – 35.8, antennae> 32.0. ♂ (Fig. 13). Of average size (body length 95.1 – 113.6 mm) and typical form for the genus, prettily multicoloured with fairly short alae that reach no further back than two-thirds the way along abdominal segment III (length 21.9 – 27.4 mm). Body surface slightly glossy. Meso- and metathorax as well as meso- and metafemora mid to dark green and to a variable degree of a slight bluish wash; the femora all with apex black (Fig. 13 P). Pronotum green laterally, ochre medially and with two closely spaced almost parallel black medio-longitudinal lines (Fig. 13 M). Head dark yellow to orange with an almost indiscernible washed darker postocular streak (Figs. 13 M – N); the frons with a washed greyish central marking. Antennae dark reddish brown, except for scapus and pedicellus, which are greenish ochre. Tegmina and costal region of alae reddish mid to dark brown with all longitudinal veins ochraceous yellow and the anterior margin marked by a broad white band (Fig. 13 O); anal fan transparent grey with brown veins. Abdomen basically orange and very variably furnished with black markings, which are mostly restricted to the posterior portion of terga II-VII (e. g. holotype), or terga almost wholly black with only the lateral margins and posterior margin orange. Sterna II – VII green with washed black marking anteriorly and posterior margin broadly orange. Terga VIII and IX only with a variably shaped elongate black lateral marking, VIII with lateral margin broadly white (Figs. 13 G – H, J – K). Anal segment destitute of black markings. Profemora bright orange to red with compressed base turquoise to blue (Figs. 13 M – N). All tibiae orange with three black annulae. Basitarsi orange with apex black. It is noteworthy that the holotype has all the colours somewhat faded with the green portions more bluish than in the numerous paratypes. Head (Figs. 13 M – N): Basically, as in ♀ but the coronal line and lateral furrow more indented and the impression of frons rather oval. Eyes much larger, projecting hemispherical and their diameter corresponding to as much as 0.7 x length of gena. Antennae very long and reaching to posterior of abdominal segment V; otherwise as in ♀. Thorax: Pronotum basically as in ♀. Mesothorax elongate, 7.2 x longer than prothorax and uniform in diameter with a slight widening at anterior margin and in posterior portion. Mesonotum with a very fine medio-longitudinal line and anterior margin with the transverse ridge indistinct. Medio-longitudinal carina of mesosternum rather indistinct and almost indiscernible on metasternum. Tegmina moderately elongate, spatulate with basal two-thirds gradually narrowing and the apical one-third rounded; central hump rather shallow and obtusely rounded (Fig. 13 O). Alae somewhat variable in length and ranging from merely reaching to posterior margin of abdominal segment II to reaching two-thirds the way along segment III (e. g. holotype). Abdomen: Segments II-V slightly increase, VI and VII decreasing in length with VII only about 0.6 x length of V and V about 5.5 x longer than wide; all uniform in diameter, VII notably constricted medially. Sterna II – VII all with a very distinct and acute medio-longitudinal carina. Tergum VIII trapezoidal in outline with posterior margin about 1.25 x wider than anterior margin and a little less than half the length of VII. IX narrowed in posterior half, strongly convex longitudinally roughly as long as VIII. Anal segment shorter than IX, slightly narrowing towards posterior, moderately tectate longitudinally; lateral surfaces with a shallow gibbose swelling sub-basally (Figs. 13 H, K); the posterior margin strongly inflated, labiate and arched outwards, the ventral surfaces minutely denticulated and facing each other at an angle of about 40 °. Cerci round in cross-section, very gently up-curved with apex weakly club-shaped and considerably projecting beyond anal segment. Vomer broadly heart-shoed with base large and oval and the terminal hook short and distinctly up-curved; ventral surface impressed interiorly with a central swelling and the outer margins broad and inflated (Figs. 13 I, L). Phallus not distinctly projecting over upper margin of poculum, Poculum moderately sized, bowl-shaped with the base somewhat protruded centrally and obtusely bulgy (Figs. 13 G, J); the posterior margin noticeably narrowed and roughly reaching to posterior of tergum IX (Figs. 13 I, L). Legs: Long, slender and with all carinae minute but fairly uniformly dentate; ventral teeth comparatively more pronounced than those of dorsal carinae. Profemora noticeably longer than head, pro- and mesothorax combines, mesofemora a little longer than combined length of pro- and mesothorax, metafemora slightly projecting over posterior of abdominal segment V and metatibiae projecting beyond tip of abdomen by more than combined length of terminal four abdominal segments. Medioventral carina of meso- and metafemora supplied only with a few minute spines. Meso- and metabsitarsi about as long as remaining tarsomeres combined, probasitarsus very elongate and considerably longer than remaining tarsomeres taken together; all destitute of teeth. Measurements of ♂ HT [mm]: Body 102.0, pronotum 2.7, mesonotum 23.8, metanotum -, median segment -, tegmina 5.4, alae 26.0, profemora 39.6, mesofemora 30.5, metafemora 39.0, protibiae 50.0, mesotibiae 34.8, metatibiae 43.3, antennae 80.0. Measurements of ♂♂ PT [mm]: Body 95.1 – 113.6, pronotum 2.6 – 2.9, mesonotum 21.6 – 27.3, metanotum -, median segment -, tegmina 5.4 – 6.8, alae 21.9 – 27.4, profemora 31.8 – 40.3, mesofemora 26.0 – 32.7, metafemora 31.3 – 39.8, protibiae 38.9 – 49.3, mesotibiae 26.3 – 33.3, metatibiae 33.1 – 44.2, antennae> 57.0.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF975D01FF52FED3A28DFF52.taxon	discussion	Remarks. Egg unknown.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF975D01FF52FED3A28DFF52.taxon	distribution	Distribution. Philippines. Mindoro (Mount Halcon – type locality [RBINS, OC]; Naujan Municipality, Arangin [FH]).	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF925D1EFF52FEF7A2B3FECE.taxon	description	(Figs. 14 – 15, 21 I – J, 22 B) HT, ♀: Philippinen, O-Luzon, Sierra Madre, Provinz Aurora, Dingalan, local collector VII. 2011 [RBINS, ex coll. FH No. 1266 - 1]. PT, egg (ex ovipositor of HT): same data [RBINS, ex coll. coll. FH No. 1266 - E]. PT, ♂: Philippinen, O-Luzon Id., Provinz Nueva Ecija, Aurora N. P., Gabaldon Munip., Mingan Mts., local collector 2013 [coll. FH, No. 1266 - 2]. Differentiation. The large ♀ of this new species is readily separated from all other Philippine members of the genus by combining a long and lanceolate subgenital plate with remarkably stocky limbs, that have the femora strongly incrassated (Fig. 14). The stocky limbs and long median segment, which is almost equal in length to the metanotum is shared with leytensis, but this species is much larger and slenderer and has the lateral margins of the mesonotum and abdominal terga not deflexed. The lanceolate subgenital plate (Figs. 14 G – I) is shared with the ♀♀ of S. maganda sp. nov. from Mindoro and S. panyense sp. nov. from the island of Panay, but both species are notably smaller, slenderer and geographically separated from matipuno sp. nov. From those two species, these ♀♀ more closely resemble maganda, but in addition to the abovementioned features they clearly differ by the even more incrassate meso- and metafemora, which however have the ventral dentations of the two outer ventral carinae less prominent and rather spinose (Fig. 14 F; distinctly serrate and black in maganda), and relatively longer median segment. The ♂ (Fig. 15) most closely resembles that of the Luzonian S. fallax (Brunner v. Wattenwyl, 1907) from which however it is readily distinguishable by the notably slenderer overall shape, more delicate limbs, distinctively trapezoidal anal segment (Fig. 15 G), that strongly widens towards the posterior and has the outer angles somewhat protruded and triangular (narrowed posteriorly in fallax), less globose head (Fig. 15 D), which lacks the black postocular streak seen in fallax, ochraceous mesonotum that has irregularly shaped black lateral streaks, black longitudinal dorsal streak in the posterior half of the mesopleurae (Figs. 15 D – E), longer alae that reach as far back as to the posterior margin of abdominal segment III and the lack of definite annulae of the meso- and metatibiae. The fairly unusual egg of this new species is readily separated from all other known eggs of congenerics by the almost uniformly sub-spherical chorion wholly smooth chorion surface and large, knob-like capitulum (Figs. 22 I – J).	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF925D1EFF52FEF7A2B3FECE.taxon	etymology	Etymology. The name matipuno (filipino = robust, hardy, muscular) emphasizes on the robust and stocky habitus and strong limbs of the ♀ of this large and eye-catching new species. Masculine.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF925D1EFF52FEF7A2B3FECE.taxon	description	Description. The colouration is described upon colour photos of a live ♀ taken by Albert Kang (Malaysia) at Real, Quezon, Luzon (Fig. 22 B). ♀ (Fig. 14). Large (body length incl. subgenital plate 185.0 mm), form very stocky for the genus, with strong legs that have the meso- and metafemora prominently incrassate, and with a long, lanceolate subgenital plate that considerably extends beyond the tip of the abdomen. The holotype is discoloured due to a provisional storage in ethanol, being plain yellowish ochre with the mesonotum and metanotum as well as abdominal segment II with a dull orange hue. Colour of the life insects unevenly olive to mid green and partly with some buffy portions. Some off-white spots along lateral margins of mesonotum and the meso- and metapleurae with irregular dark brown markings. Head including eyes pale green, the genae with a faint washed greyish dark green postocular streak. Pronotum with a pair of closely spaced, sub-parallel, slender black medio-longitudinal lines; mesonotum with a pair of sepia brown markings that consist of ana aggregations of small speckles. Abdominal sterna mostly greyish with some white and faintly flecked with brown. Legs mid green; the bases of all femora dark red to purple and the meso- and metafemora and tibiae each with three faint, washed greyish dark green annulae. All teeth of the legs tipped with black. The antennae green dorsally and gradually becoming dark grey towards the tip and mostly black ventrally. Head (Figs. 14 D – E): Sub-globose, 1.36 x longer than wide and broadest just behind the eyes; vertex convex with a very shallow impression anteriorly, the coronal line and lateral furrow just very weakly indented. Eyes rather small, moderately projecting and their diameter corresponding to 0.6 x length of gena. Antennae reaching to posterior margin of mesothorax; scapus compressed, roundly rectangular and about 1.6 x longer than wide; pedicellus much shorter, round in cross-section and barrel-shaped. Thorax: Pronotum shorter and considerably narrower than head, almost rectangular in outline with the lateral margins very gently concave, 1.6 x longer than wide. Anterior margin somewhat inflated medially and followed by a deep, slightly arched transverse furrow that laterally terminates in a distinct pit some distance off the outer angle of notum; median line indented of entire length and transverse median sulcus impressed, almost straight and fairly short (Fig. 14 E). Mesothorax fairly short and only 4.5 x longer than prothorax, distinctly constricted in anterior portion, somewhat inflated pre-medially and slightly widened at posterior margin. Mesonotum with a very fine medio-longitudinal line that is only clearly visible in the narrowed anterior portion but soon almost vanishes towards middle of segment; the anterior margin with a low medially notched and curved transverse carina. Metanotum very weakly tectate medio-longitudinally, 2 x longer than wide, constricted medially and less than half the length of mesonotum. Meso- and metapleurae with an obtuse longitudinal carina close to upper margin. Meso- and metasternum with a prominent but obtuse medio-longitudinal keel. Abdomen: Median segment about 0.65 x length of metanotum, sub-rectangular and only 1.2 x longer than wide. Segments II – VII sub-uniform in width with II – IV slightly increasing and V – VII slightly decreasing in diameter. II – IV increasing in length, V as long as IV and VI – VII decreasing in length; V about 1.6 x longer than wide and VII about as long but narrower than II. Sterna II and II with a distinct medio-longitudinal carina, which is only discernible in the very anterior portion on IV-VII. Praeopercular organ formed by a verrucose posteromedian swelling on sternum VII and surrounded by an area of minute, irregularly longitudinal directed rugulae (Fig. 14 G). Tergum VIII constricted medially and strongly convex longitudinally with posterior margin slightly wider than anterior margin and roughly half as long as VII. IX sub-quadrate in dorsal aspect and some two-thirds as long as VIII. Anal segment notably longer than IX and slightly gradually narrowing towards a rounded and weakly bi-lobate posterior margin that has a small median indention; dorsal surface with a fine medio-longitudinal carina and lateral margins with a rather shallow concave emargination posteriorly (Fig. 14 H). Epiproct small, distinctly transverse and just weakly projecting under anal segment (Fig. 14 H). Cerci conical and slightly projecting over anal segment. Gonapophyses VIII strong, up-curved and roughly reaching to posterior of anal segment, therefore fully concealed under it. Subgenital plate elongate, lanceolate with the elongated apical portion distinctly keeled medio-longitudinally, gradually narrowing towards a fairly pointed tip and projecting beyond abdomen by about the combined length of terminal three terga (Figs. 14 G – I); basal portion with a prominent pre-basal narrowing (Fig. 14 G) and a distinct arched carina along lateral surfaces. Legs: Legs fairly short and very stocky for the genus, the meso and metafemora in particular very strong and distinctly incrassate in basal half with the two outer ventral margins expanded; all carinae to a variable degree dentate with the ventral dentations of mid and hind legs much more prominent than those of the dorsal carinae. Profemora longer than pro- and mesothorax combined, mesofemora scarcely longer than mesothorax, metafemora reaching somewhat beyond posterior margin of abdominal segment IV and metatibiae roughly reaching to posterior of VII. Posteroventral carina of profemora with about 25 teeth, the teeth of anterodorsal carina slightly less in number and comparatively somewhat lower but broader; posterodorsal carina only supplied with about 12 – 13 minute teeth. Two outer ventral carinae armed with prominent teeth, that are largest and most closely spaced in the incrassate basal half and gradually become smaller towards the apex with only the apical tooth somewhat enlarged; dorsal carinae only with minute and more widely spaced teeth; the medioventral carina distinctly raised and armed with 10 – 12 (mesofemora; Fig. 14 F) or 13 – 15 (metafemora) fairly strong spines. Ventral teeth of meso- and metatibiae also notably more closely spaced and numerous than the comparatively smaller teeth of dorsal carinae. Meso- and metabasitarsi longer than following three tarsomeres combined with all four carinae denticulate, probasitarsus as long as remaining tarsomeres taken together except claw and only the anteroventral carina supplied with a few minute denticles. Measurements of ♀ HT [mm]: Body (incl. subgenital plate) 185.0, body 170.0, pronotum 6.8, mesonotum 33.0, metanotum 14.0, median segment 8.5, profemora 44.0, mesofemora 34.8, metafemora 37.5, protibiae 48.0, mesotibiae 32.0, metatibiae 41.5, antennae 50.0. ♂ (Fig. 15). Fairly large for the genus (body length 124.0 mm), form very slender, with distinctive black lateral markings on mesonotum and a trapezoidal anal segment that is strongly widening towards the posterior. General colour ochraceous with black markings on mesonotum and abdominal terga. Head dark yellow with the frons orangey (Fig. 15 E) and the genae with a faint, washed green postocular streak that vanishes some distance before posterior end of head capsule (Fig. 15 D); scapus and pedicellus greenish ochre, rest of antennae blackish brown. Pronotum olive. Mesonotum with lateral margins cream-coloured and with an irregularly shaped black lateral streak that starts narrow at the anterior and gradually becomes wider and finally disperses towards back of segment (Figs. 15 D – E). Tegmina and costal region of alae cream-coloured with all longitudinal veins dark brown and the anterior margin broadly light creamish; the anal fan transparent grey with brown veins. Abdominal terga II-VII dark brown anteriorly and posteriorly with two pairs of black spots, the second pair of which is much larger. Sterna with similar posterior black spots, II wholly green, III mostly green and IV with a green longitudinal median streak, V-VII ochre (Fig. 15 C). Profemora dark green basally (Figs. 15 D – E) and the rest orangey brown with slight greenish wash. Meso- and metafemora green with the apex somewhat darkened; all tibiae greenish ochre with three faint darker brown annulae. Head (Figs. 15 D – E): Oval, about 1.2 x longer than wide, broadest at the eyes and distinctly narrowing towards the posterior; vertex moderately convex with a notably indented coronal line and the lateral furrow only indented at posterior margin of head capsule; in front with a shallow triangular impression. Frons with a distinct, small Wshaped pit between bases of antennae. Eyes very large, projecting slightly more than hemispherically and their diameter corresponding to 0.75 x length of gena. Antennae long and reaching slightly beyond posterior margin of abdominal segment III; scapus and pedicellus as in ♀. Thorax: Pronotum basically as in ♀. Mesothorax very elongate and 7.2 x longer than prothorax, uniform in diameter with only a slight widening at anterior margin and somewhat expanded posteriorly. Mesonotum with a very fine medio-longitudinal line but the transverse bulge at anterior margin more pronounced than in ♀ with median portion weakly bi-gibbose and indented medially. Meso- and metasternum with a prominent medio-longitudinal keel. Tegmina elongate with the basal half strongly narrowed and the apical half roundly angular; the central hum fairly well developed and obtusely conical. Alae projecting somewhat behind posterior margin of abdominal segment III. Abdomen: Segments II – VII uniform, in diameter, II – IV slightly decreasing in length, V as long as IV and VI- VII decreasing in length with VII only two-thirds the length of V; VI and V 7 x and VII only 5.5 x longer than wide. Sterna II – VII all with a distinct and acute medio-longitudinal keel. Tergum VIII slightly constricted just behind anterior margin with posterior half gradually widening and posterior margin 1.3 x wider than anterior margin; less than half as long as VII. IX rectangular, just scarcely narrower than posterior margin of VIII and about 1.25 x longer than wide. Anal segment shorter than IX, distinctly trapezoidal in dorsal aspect (Fig. 15 G), being widened towards the posterior with the posterolateral angles strongly expanded and roundly triangular, in dorsal aspect the posterior margin with a shallow triangular emargination; ventral surfaces densely set with minute denticles to form a pair of large thorn-pads (Fig. 15 H); these just slightly facing each other at an angle of about 120 °. Vomer roundly triangular in outline with the base rather parallel-sided and the terminal hook strong, short and up-curved; lateral margins strongly inflated and ventral surface with a deep medio-longitudinal furrow (Fig. 15 H). Cerci fairly small, round in cross-section and the apex somewhat club-like and arched inwards; scarcely projecting over anal segment. Phallus digitiform with a narrow apex that project noticeably over left upper margin of poculum (Figs. 15 F – H). The poculum small, bowl-shaped with the medio-basal portion somewhat bulgy and protruded (Fig. 15 F) and the posterior margin rounded and just not reaching to posterior of tergum IX (Fig. 15 H). Legs: Long, slender and with all carinae minutely but acutely dentate; the teeth on ventral carinae comparatively larger than those on dorsal carinae. Profemora considerably longer than head, pro- and mesothorax combined, mesofemora roughly equal in length to pro- and mesothorax combined, metafemora almost reaching abdominal segment V and metatibiae projecting beyond tip of abdomen by about the combined length of terminal four abdominal segments. Apical tooth on two outer ventral carinae of meso- and metafemora slightly enlarged; the medioventral carina distinct and armed with about 22 – 25 somewhat unevenly spaced but fairly distinct spines. Tarsi all elongate, meso- and metabasitarsi almost as long as remaining joints excluding claw taken together and with the three ventral carinae minutely denticulated; probasitarsus unarmed and slightly longer than combined length of remaining tarsomeres. Measurements of ♂ PT [mm]: Body 124.0, pronotum 3.8, mesonotum 27.2, metanotum 4.9, median segment 9.0, tegmina 6.1, alae 35.8, profemora 40.6, mesofemora 32.8, metafemora 41.5, protibiae 51.7, mesotibiae 34.0, metatibiae 45.0, antennae 73.0. Egg (Figs. 21 I-J). Fairly large for the genus, chorion almost circular in lateral aspect, scarcely more than 1.2 x longer than wide and oval in cross-section; the anterior portion slightly constricted and the anterior margin weakly inflated; polar-area rounded. Chorion surface even and almost wholly smooth with only a few very minute scattered granules; dull. Micropylar plate rather large and more than half as long as chorion; spearhead-shaped with the posterior portion fairly broad and gradually tapering towards a narrow tip; surface even and like chorion. Micropylar cup slightly displaced towards the anterior, small and bowl-shaped. Operculum slightly oval and flat with the outer margin somewhat raised and set with some small granules. Operculum large roundly knob-like, slightly constricted at base and with a central pit. Colour plain orangey mid brown with the anterior portion and area around micropylar plate washed dark brown. Outer margin of micropylar plate russet, micropylar cup black. Outer margin of operculum ochre, the capitulum russet and black at the base. Measurements [mm]: Length incl. operculum 3.8, length 3.6, width 2.9, height 3.1, length of micropylar plate 2.1.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF925D1EFF52FEF7A2B3FECE.taxon	discussion	Comments. A photo of a live ♀ in situ taken by Albert Kang (Singapore) shows the specimen to be domestic to a guava tree (Psidium guayava, Myrtaceae), which shows considerable feeeding damage. Thus, guava can be assumed to be one of the host-plants of this stunning species (Fig. 22 B).	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF925D1EFF52FEF7A2B3FECE.taxon	distribution	Distribution. Philippines. Luzon: Province Aurora (Dingalan [RBINS – type locality]); Province Nueva Ecija (Gabaldon Municipality, Mingan Mountains [FH]); Province Quezon, Real [photographic record by A. Kang; Fig. 22 B]).	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF8D5D15FF52FE0BA773FD02.taxon	description	(Figs. 16 – 18, 21 A – B, 23 B – C)	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF8D5D15FF52FE0BA773FD02.taxon	description	PT, 19 ♀♀, 12 ♂♂, 2 ♀♀ (immature): Coll. I. R. Sc. N. B., Philippines, Mindanao, Leg: R. Cabale 2011, Gift: J. Bresseel, IG: 32.386 [RBINS]. PT, 3 ♀♀, 4 ♂♂, 1 ♀ (immature), 1 ♂ (immature), 75 eggs: ex Zucht F. Hennemann 2007, Herkunft: Philippinen, Mindanao, Bukidnon Prov., Tampakan [coll. FH, No’s 0630 - 1 to 9 & E]. PT, 2 ♂♂: Philippinen, Mindanao Island, Mount Matutum 2007 [coll. FH, No’s 0630 - 10 & 11]. PT, 11 ♀♀, 17 ♂♂, 2 ♀♀ (immature), 1 ♂ (immature): Philippinen, Mindanao Island, S-Cotabato Prov., Tupi, Mount Matutum, V. 2011 [coll. FH, No’s 0630 - 13 to 43]. PT, ♀: Philippinen, Mindanao Id., Davao Reg., Compostela Valley, nr. Diwalwal (Mount Diwata), I. 2011 [coll. FH, No. 0630 - 44]. PT, 3 ♀♀, 3 ♂♂: Philippinen, Mindanao Island, Prov. Cotabato, Municipality Magpet O of Davao, local collector X. 2011 [coll. FH, No’s 0630 - 45 to 50]. PT, ♂: Philippinen, NW Mindanao Island, Provinz, Llanao del Norte, Kapatagan, local collector VIII. 2011 [coll. FH, No. 0630 - 51]. PT, 3 ♀♀, 1 ♂: Philippinen, Mindanao Island, Prov. Agusan del Sur, Sibagat Munip., ca. 50 m, local collector VIII. 2012 [coll. FH, No’s 0630 - 52 to 55]. PT, ♀: Philippinen, Mindanao Island, Prov. Agusan del Sur, San Agustin, local collector IV. 2003 [coll. FH, No. 0630 - 56]. PT, 23 ♀♀, 11 ♂♂, 1 egg: Philippinen, Prov. Cotobato, Mount Matutum, 03.2006 [coll. OC, No’s 0128 - 1 to 34 & E] PT, 29 ♀♀, 10 ♂♂: Philippinen, Mindanao, Prov. Cotabato, Mount Apo, local collector 03.2010 [coll. OC, No’s 0128 - 35 to 74].	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF8D5D15FF52FE0BA773FD02.taxon	materials_examined	Further material: ♂: Philippinen, Mindanao Id., North Cotabato Prov., Mt. Apo, Marbel River, 1100 – 1400 m, 19.724. II. 1994, Alex Buenafe leg., coll. R. A. Müller [coll. OZ, No. 220 - 1] ♂: Philippinen, Mindanao, Bukidnon Prov., Mt. Imbayao, Baungon, Tanalaong River, 25. IX. - 01. X. 1989, Th. Borromeo leg., ex Coll. R. A. Müller [coll. OZ, No. 220 - 2]; ♀ (immature n 4): Philippines, Mindanao Id., North Cotabato, Mt. Apo, Lake Venado, 2280 m, 01. IV. 1995, leg. A. Bunafe, ex coll. R. A. Müller [coll. OZ No. 220 - 3]; 14 ♀♀, 2 eggs: Philippines, Mindanao, Sultan Kudawat Palimbang, Brgy, Milbuk, 12.30 - 20.30 h, on Psidium guayava L., 25. viii. 1999, leg. O. Zompro [coll. OZ 330 - 1 to 14, 330 - E]; 1 ♂: Surigao, Mindanao, Baker, Pharnacia annulata Redt.? 451 [ANSP]. Differentiation. Females of this new species are very similar and morphologically closest to S. fallax (Brunner v. Wattenwyl, 1907) from Luzon, Samar and Panay with which they share the general appearance, form, size and shape of the subgenital plate and essentially are difficult to distinguish. These ♀♀ may however be separated by the more globose and strongly convex vertex (Figs. 17 F – H), notably more pronounced dentations of the limbs, which are distinctly spinose particularly in the basal portions of the two outer ventral carinae of the meso- and metafemora and includes two comparatively larger apical teeth on the same carinae of the mesofemora (Figs. 17 I – L), more shallow lateral excavation of the anal segment (Figs. 17 A – B), and differently shaped praeopercular organ (Fig. 17 E). Males show close affinity to those of fallax and S. leytensis (Zompro, 1997), the latter of which is also found throughout Mindanao and may be sympatric in certain localities. The distinct postocular streak is shared with the very similar fallax but it is broader in this species and either black or dark green (Figs. 18 I, K). Besides, these ♂♂ are smaller (body length at best 100.0 mm) and slightly more robust in shape with stockier legs, that have all the dentations comparatively more pronounced, the black transverse posterior bands of abdominal terga II-VI are much broader, and the vomer is basically heart-shaped with the base broader (Fig. 18 H; almost gradually tapering and basically triangular in fallax). From leytensis the ♂♂ of mindanaense can be distinguished by the dark postocular streak (Figs. 18 I – K; head almost unicoloured in leytensis), shorter alae that only reach about halfway along abdominal segment III (projecting over posterior margin of segment III on leytensis), and slightly less widened basal portion of the vomer (Fig. 18 H). From the known eggs of Sadyattes, those of mindanaense (Figs. 21 A – B) most strongly resemble the ones of fallax but differ by the more wide-meshed network of ridges and bulges of the chorion, presence of a shallow posteroventral indention near the polar-area as well as the somewhat larger and rather spearhead-shaped micropylar plate, which has the polar extension notably longer than in fallax (micropylar plate rather pear-shaped in mindanaense).	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF8D5D15FF52FE0BA773FD02.taxon	etymology	Etymology. Named after the distribution on the island of Mindanao, where it is endemic. Neuter.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF8D5D15FF52FE0BA773FD02.taxon	description	Description. The colouration is described upon colour photographs of live insects as well as dried specimens. Measurements in table 5 below. ♀ (Fig. 16). Medium-sized to fairly large (body length incl. subgenital plate 141.0 – 183.0 mm), form typical for the genus, with a strongly globose head and a broad, spatulate subgenital plate that projects over the tip of the abdomen by no more than the combined length of terminal two abdominal terga. Colour very variable (see summary of variability below), ranging from grey and drab over various tones of ochre, buff and brown to olive and mid green; often species are prettily flecked and mottled with white and dark brown and occasionally possess either large white or cream-coloured markings on single or various dorsal body segments (Figs. 16 C, F) or are more or less continuous medio-longitudinal dorsal streak of the same colour (Figs. 16 D – E). Head at least with a broad dark green postocular streak in green specimens but mostly with various blackish speckles on genae that more or less definitely shape an arrangement that resembles a postocular streak; lower portion of genae often whitish. Antennae drab with all segments of basal half black; occasionally scapus not wholly black ventrally. Legs more or less distinctly annulated and flecked with white and dark grey or black; more rarely plain green. Base of profemora red interiorly, base of meso- and metafemora almost always slightly pink to reddish. Head (Figs. 17 F – H): Globose with the vertex almost spherical posteriorly and shallowly impressed anteriorly; the coronal line and lateral furrow weakly indented; just slightly longer than wide and broadest directly behind eyes. Frons with a deep transverse impression just behind bases of antennae. Eyes of moderate size but distinctly projecting; their diameter corresponding to about 0.8 x length of gena. Antennae reaching one-third the way along metanotum; scapus small and slightly constricted basally but basically rectangular, about 1.6 x longer than wide; pedicellus round in cross-section weakly narrowing towards apex and about two-thirds the length of scapus. Thorax: Pronotum shorter and scarcely more than half as wide as head, rectangular in outline with a gentle pre-median narrowing and a concave medially somewhat protruded anterior margin; transverse furrow behind anterior margin laterally terminating in narrow but distinct pit close to outer angles of notum; transverse median sulcus distinct, gently arched and expanding over entire width of notum, the median line indented and occasionally with a pair of low median granules just in front and / or behind the transverse sulcus; a few minute granules may also be present along median line throughout the posterior portion of pronotum. Mesothorax 6.6 – 6.8 x longer than prothorax, slightly constricted at anterior margin slightly widened posteriorly. Mesonotum with medio-longitudinal line almost indiscernible and at anterior margin with a distinct, medially indented and weakly bi-granular transverse ridge. Metanotum with medio-longitudinal line like on mesonotum, almost rectangular, 2.6 x longer than wide and about 0.45 x length of mesonotum. Meso- and metasternum with a shallow medio-longitudinal keel. Abdomen: Median segment scarcely more than half the length of metanotum, very slightly narrowing towards anterior and roughly 1.5 x longer than wide. Segments II – VII somewhat sub-uniform in diameter, II – IV increasing in length V roughly as long as IV and VI – VII decreasing in length with IV – V longest and about 2.5 x longer than wide; VII narrower than all preceding, scarcely longer than median segment and only 2.3 x longer than wide. Medio-longitudinal carina only visible on sterna II – III and anterior portion, sterna V – VII smooth. Praeopercular organ formed by an elongate-oval verrucose and ridge-like glossy black median swelling some distance in front of the posterior margin; the surrounding area unevenly granular to rugulose (Fig. 17 E). Terga-VIII and IX roughly uniform in width, VIII about 0.6 x length of VIII and almost rectangular; IX about half the length of VIII and sub-quadrate. Anal segment as long as IX with basal portion of same width and only the posterior half slightly narrowing with the posterior margin widely bi-lobate and with a small, triangular median indention (Figs. 17 C – D); lateral margins with a moderate concave emargination (Figs. 17 A – B); dorsal surface with a fairly acute medio-longitudinal keel. Epiproct small, scale like with a median keel and a minute posteromedian indention, scarcely projecting beyond anal segment (Figs. 17 C – D). Cerci slender, conical and slightly reaching beyond posterior margin of anal segment. Gonapophyses VIII slightly projecting over tip of abdomen. Subgenital plate granular basally and rather weakly keeled longitudinally in central portion; the elongated apical section flattened, spatulate and either weakly narrowing to almost parallel-sided with the posterior margin broadly rounded to roundly angular and extending beyond tip of abdomen by at most he combined length of two terminal abdominal terga (Figs. 17 A – E). Legs: Moderately long and fairly stocky with all carinae dentate; profemora slightly shorter than pro- and mesothorax combined, mesofemora somewhat shorter than mesothorax, metafemora reaching to posterior margin of abdominal segment IV and metatibiae reaching about halfway along segment VII. Teeth on anteroventral carina of profemora moderately distinct, saw-like and broader than teeth on posteroventral carina; the posterodorsal carina supplied with only 12 – 18 minute black teeth, that are rather unevenly spaced. Meso- and metafemora slightly incrassate sub-basally with the two outer ventral carinae weakly expanded; the dentations of two outer ventral carinae prominent, especially in the widened sub-basal section where they are elongate and rather spiniform, and the two apical teeth more or less prominently enlarged and forming acutely triangular lobes (Figs. 17 J – L); dorsal teeth much smaller; medioventral carina distinct and supplied with about 10 – 15 rather small spines. Medioventral carina of meso- and metatibiae rounded and raised sub-basally and the posterodorsal carina deflexed to form a small 4 - dentate apical lobe. Meso- and metabasitarsi with all four carinae minutely denticulated, the dorsal carina slightly rounded and almost as long as remaining tarsomeres except clow taken together; probasitarsus roughly equal in length to remaining joints taken together, slender and with only the ventral carinae indistinctly denticulate. ♂ (Fig. 18). Moderately sized (body length 82.0 – 100.0 mm), form stocky for the genus, with a globose head and a distinct dark postocular streak and short alae that reach no further back than half way along abdominal tergum III (length 18.0 – 21.0 mm). General colour green to olive, the abdominal terga gradually becoming more brownish towards tip of abdomen, II – VI each all with a broad and transverse black posterior marking, the corresponding sterna only with a narrow transverse-sub-posterior band. Head yellow to pale orange with a distinct dark green to black postocular streak (Figs. 18 I, K). Antennae brown with ventral surface of segments in basal half black. Pronotum with lateral portion darker in colour than median section. Tegmina and costal region of alae with anterior margin broadly off-white to pale yellow and interiorly bordered by a fine brown line, otherwise greyish green with the more interior section gradually turning to purplish brown; anal fan hyaline to transparent grey with dark green veins. Profemora drab with the anterior surface slightly russet and compressed base green. Meso- and metafemora greyish green to pale greyish brown with three faint light cream-coloured annulae and somewhat darkened at the apex; base occasionally slightly pinkish. Profemora ochre to yellow with about five faint and variable brown annulae. Meso- and metafemora dark yellow to orange with two black annulae and the apex brown (Fig. 18 D). Basitarsi creamy basally. Head (Figs. 18 I – K): Strongly globose, shape basically as in ♀, but the longitudinal lines less indented and the anterior portion of vertex with two shallow, transverse diagonal directed impressions; the indention on frons less distinct. Eyes very large, projecting sub-spherically and their diameter corresponding to about 0.75 x length of gena. Antennae reaching to posterior of abdominal segment III; scapus and pedicellus like in ♀. Thorax: Pronotum essentially as in ♀, notably shorter and narrower than head, but the anterior margin less protruded (Fig. 18 J). Mesothorax uniform in diameter except for being weakly widened posteriorly. Mesonotum with a very fine medio-longitudinal line that is almost indiscernible in the posterior three-quarters, the anterior margin with an arched transverse ridge that is notched medially. Meso- and metasternum with a fine medio-longitudinal carina. Tegmina elongate and very slender, spatulate with the basal two-thirds notably narrowing and the apical one-third roundly angular; central hump small but pronounced and obtusely conical. Alae reaching about halfway along abdominal segment III. Abdomen: Segments II – VI just very slightly sub-uniform in length and on average 5 x longer than wide; VII shorter than all preceding, constricted medially, somewhat widened posteriorly and only about 0.7 x length of VI. Tergum V with a low and obtuse posteromedian swelling. Sterna distinctly carinate medio-longitudinally. Tergum VIII strongly inflated, broadest of all abdominal terga, roundly trapezoidal in dorsal aspect with posterior margin almost 1.5 x wider than anterior margin, slightly more than half as long as VII. IX much narrower, also trapezoidal, scarcely longer than VIII and gradually narrowing towards the posterior; roundly tectate dorsally. Anal segment wider and shorter than IX with lateral surfaces convex and obtusely gibbose sub-basally; the posterior margin noticeably inflated, somewhat labiate and with the obtuse rounded outer angles deflexed and angled outward; anteriorly the posterior margin is bordered by a distinct transverse groove on each side of the obtuse medio-longitudinal keel (Fig. 18 F); ventral surfaces densely denticulated and facing each other at an angle of about 45 °. Cerci slightly club-shaped, round in cross-section, gently up- and in-curved and notably projecting beyond anal segment. Phallus slightly projecting over left lateral margin (Fig. 18 H). Vomer rather small, basically heart-shaped in outline with a deep medio-longitudinal furrow on ventral surface and the lateral portion strongly inflated; the terminal hook short but string and weakly up-curved (Fig. 18 H). Poculum bowl-shaped but rather weakly bulgy with a small and obtuse rounded swelling medio-basally (Fig. 18 E); posterior margin narrow, roundly triangular somewhat projecting over posterior margin of tergum IX (Figs. 18 G – H). Legs: Moderately slender and of average shape with all carinae rather minutely dentate. Profemora roughly equal in length to head, pro- and mesothorax combined, mesofemora slightly shorter than mesothorax, metafemora almost reaching posterior margin of abdominal segment V and metatibiae reaching beyond tip of abdomen by about combined length of terminal abdominal segments. Anterodorsal carina of profemora fairly distinct serrate teeth, those on the posteroventral carina somewhat less in number, comparatively smaller and often black; posterodorsal carina supplied with about 20 minute denticles. Dentations on two outer ventral carinae of meso- and metafemora more numerous and comparatively larger than those of dorsal carinae. Meso- and metabasitarsi roughly as long as remaining joints except claw taken together and with a few minute denticles in basal half of two outer ventral carinae; probasitarsus slightly longer and unarmed. Variability. Both sexes, but ♀♀ in particular, show a considerably size range. The specimen from Nabunturan in the collection of RBINS and one of the examples from Sibagat in the author’s collection (coll. FH No. 0630 - 54) are notably larger than all other specimens at hand at a total length of 183.0 mm (see table 5 below). Interestingly, also the smallest examined specimens with a total length of only 141.0 mm is from Sibagat (coll. FH, No. 0630 - 53). The ♀ from San Agustin is just slightly smaller than the two aforementioned large specimens. These large specimens are generally slenderer in overall shape and also have comparatively slenderer limbs, but show no other significant morphological differences from all other examined specimens. All three localities are lowland habitats in the eastern region of Mindanao. The majority of ♀♀ from Mount Apo and Mount Matutum range from 146.0 mm to 167.0 mm in total length. Variability is also seen in the length and shape of the subgenital plate, which can have the apex broad and just slightly projecting beyond the apex of the abdomen or is notably elongated, somewhat tapering towards a more narrowly rounded apex that extends beyond the abdomen by as much as the combined length of terga IX and X. While ♂♂ are fairly consistent in colour, remarkably variability is seen in ♀♀. The great majority of specimens shows pretty mottling that consist of various tones of ochre, brown, grey, cream and white with the white areas strongly varying in density, size and shape. Considerably less specimens are olive or green and mostly these have a variably shaped either continuous or interrupted white medio-longitudinal streak along the dorsal body surface and sometimes even the head (e. g. the example from Nabunturan in RBINS; Fig. 23 B). Even more rarely are plain buff to ochre specimens (e. g. examples from San Agustin and the large ♀ from Sibagat). Captive reared ♀♀ originating from Tampakan are more or less uniformly green to olive (Figs. 16 A – B) with only single ones showing a white medio-longitudinal streak along the dorsal body surface. The lower portion of the genae is more or less whitish in all examples at hand and the postocular streak may be represented either as a washed dark green or brown streak or a variable cluster of dark brown to blackish speckles (Figs. 17 F – H). The only noteworthy chromatic variability seen in ♂♂ concerns to the postocular streak which mostly is black (Fig. 18 K) but dark green in the captive reared specimens from Tampakan (Fig. 18 L) and colour of the apex of the meso- and metafemora, which ranges from washed mid brown (Fig. 18 D) to distinctly black. Egg (Figs. 21 A – B). Medium-sized, chorion slightly longer than high, flattened laterally and distinctly oval in cross-section; polar-area with a shallow indention anteriorly and the anterior margin strongly arched if seen in lateral aspect. Surface wholly covered by an irregular wide-meshed network of obtuse granular bulges that leave impressed areas inbetween them; surface of the indented areas minutely and densely granular. Micropylar plate small, spearhead-shaped with the broaded anterior portion oval and acutely triangular at anterior tip and the lower portion strongly narrowed with the strongly inflated and bulgy outer margin fused with another. Internal surface lowered, flat and with a small bowl-shaped micropylar cup at lower end of widened anterior area. Operculum elliptical and strongly arched with the dorsal and ventral portions downward-directed. Outer margin somewhat raised and labiate. Dorsal portion with a small but distinct pit that is surrounded by a low rim. Capitulum an irregular, mushroom-like protrusion that consists of several irregular raised and radially directed ridges that fuse in centre. Colour greyish ochre with the indented portions of the chorion drab. Operculum dark brown and the raised ridges of the capitulum with a slight reddish hue. Micropylar cup black. Measurements [mm]: Length incl. operculum 3.4, length 3.1, width 2.1, height 2.4, length of micropylar plate 1.7.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF8D5D15FF52FE0BA773FD02.taxon	discussion	Remarks. A culture stock of S. mindanaense sp. nov. was reared for several generations based on eggs imported from Tampakan, Bukidnon Province and laid by specimens collected by Benjié Mabanta (Philippines) during 2005 and 2006 (Harman, 2013: 13). The culture was misidentified as “ Eucarcharus feruloides ” and included on the Phasmid Study Group culture list as culture No. PSG 287. Meanwhile the culture has vanished. In captivity in the Philippines guava (Psidium guajava, Myrtaceae) has frequently been accepted as food. Alternative food plants accepted in captivity in Europe included eucalyptus (Eucalyptus spp., Myrtaceae), hypericum (Hypericum patulum, Hypericaceae), oak (Quercus spp., Fagaceae), beech (Fagus sylvatica, Fagaceae), bramble and raspberry (Rubus spp., Rosaceae), wild roses (Rosa spp., Rosaceae) and also strawberry (Fragaria vesca, Rosaceae). Females are prolific egg-layers and produce an average of 4 – 5 eggs per day and ♀, which are simply flicked away by an abrupt movement of the abdomen. The development is fairly rapid, with eggs hatching after only 12 – 16 weeks and nymphs reaching maturity after no more than 4 – 5 at average temperatures of 22 – 25 ° C. Nymphs and adult insects are difficult to handle and behave hectically if disturbed, with immatures in particular easily shedding limbs if a leg is grasped by a predator or the uncareful breeder.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF8D5D15FF52FE0BA773FD02.taxon	distribution	Distribution. Philippines. Mindanao (endemic): " Mindanao " [RBINS]; Province South Cotabato (Mount Apo [FH, OC]; Mount Matutum [FH, OC]; Mount Matutum, Tupi Municipality [FH]); Province North Cotabato (Mount Apo, Marvel River 1100 – 1400 [OZ]; Mount Apo, Lake Venado 2280 m [OZ]); Province Cotabato (Magpet Municipality [FH]); Province Bukidnon (Tampakan 1300 m [FH]; Mount Imbayao, Baungon, Tanalaong River [OZ]); Province Llanao del Norte (Kapatagan [FH]); Province Agusan del Sur (Sibagat [RBINS, FH]; San Agustin [FH]); Province Davao de Oro (Compostela Valley, near Mount Diwata [FH]; Nabunturan [RBINS]); Province Sultan Kudarat (Palimbang Municipality, Milbuk [OZ]); Province Surigao del Norte (Surigao [ANSP]).	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF865D15FF52FD47A011F85B.taxon	description	Seow-Choen, 2019: 268. Brock & Büscher, 2022: 559. [Not: Caudell, 1927: 19 Misidentification that most certainly relates to S. borrii Stål, 1875] Differentiation. A differential diagnosis of the ♂ of this species (the only sex described) is only possible upon literary sources, since the holotype is lost and no new material has yet become available. From annulatus, the only other Bornean member of the genus whose ♂ is known, this ♂ can be readily be separated by the much smaller size (body length 104.0 mm vs.> 131.0 mm in annulatus), lack of a dark medio-longitudinal streak on the tegmina and costal region of the alae and white posteromedian marking of the abdominal sterna. The brief original description given by Redtenbacher (1908. 451) does not provide any additional characters that would serve for distinction.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF865D15FF52FD47A011F85B.taxon	discussion	Remarks. Unfortunately, the unique ♂ holotype of Redtenbacher’s species was destroyed by a fire during the Hungarian Revolution in 1956 at the museum in Budapest (HNHM). So far, no other ♂ could be traced that matches the brief original description and could serve for a neotype in order to provide stability for the taxon. The ♂ from Tjibodas, Mount Gede, W-Java by Caudell (1927: 19) and in the collection of the USNM could not be examined but most certainly is a specimen of the Javanese S. borrii Stål, 1875. Below, a translation of Redtenbacher’s original description is provided. It is worth mentioning that at a length of 48.8 mm (Redtenbacher, 1908: 451) this species has fairly long alae for the genus, which is shared with annulatus the only other known Bornean ♂ of the genus. As stated above, it is not unlikely that the unknown ♀ of nigricornis is represented by S. decoris (Seow-Choen, 2016), a moderately sized Bornean species that is here transferred from the genus Phobaeticus Brunner v. Wattenwyl, 1907 (comb. nov.). Body length of holotype ♂ 104.0 mm (Redtenbacher, 1908: 451). Description according to Redtenbacher (1908: 451): “ ♂. Antennae blackish-brown. Head unarmed, not gibbose posteriorly. Tegmina yellowish brown, anterior margin yellowish-white. Alae transparent brown, anterior margin white. (Front legs missing in specimen at hand). Four posterior femora yellowish, dorsally sparsely, ventrally densely armed with black spines on both outer carinae. Four posterior tibiae dorsally and ventrally on both outer carinae armed with small black spines, without enlarged teeth. Metatarsus not cristate. Anal segment truncated apically, with two very short, rounded lobes. Abdominal sterna brownish and with a white marking just before apex. Poculum rounded, convex. ”	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF865D15FF52FD47A011F85B.taxon	distribution	Distribution. “ Borneo ” [HNHM]	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF855D10FF52FF2AA125FD66.taxon	description	(Fig. 19) HT, ♀: Philippines, Panay Island, 09.06.1998, Purchased from dealer, IV. 2000, Accession Number: PEB- 3035 [NHMUK, ex coll. PEB No. 3035]. Differentiation. The ♀ of this new species (the only sex known) is morphologically very close to that of S. maganda sp. nov. from the island of Mindoro with which it shares the overall shape and long, lanceolate subgenital plate (Figs. 19 G-I). It may however be separated by the slightly less globose head and lack of the black postocular speckles seen in maganda (Figs. 19 D – E), less incrassate basal portion of the meso- and metafemora, rather spinose than serrate ventral dentations of the meso- and metafemora, which are comparatively less developed but more numerous and not as distinctly black as in maganda, less enlarged apical tooth of the two outer ventral carinae of the meso- and metafemora (Fig. 19 F), as well as the more narrowed and bilobed posterior margin of the anal segment (Fig. 19 H).	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF855D10FF52FF2AA125FD66.taxon	etymology	Etymology. Named after its type-locality, the island of Panay. Neuter.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF855D10FF52FF2AA125FD66.taxon	description	Description. The unique holotype (Figs. 19 A – C) lacks the right mid leg and claw of the left mesotarsus. The left side of abdominal terga VII and VIII shows damage due to an injury that most certainly happened when the insect was still alive. ♀ (Fig. 19). Of average size (body length incl. subgenital plate 166.0 mm) and form for the genus with a long and lanceolate subgenital plate. General colour of the holotype ochraceous buff to mid brown, the abdomen somewhat darker brown due to preservation. Bases of profemora pale green (Figs. 19 D – E), all limbs with a slight orange hue and the carinae of mid and hind legs light green (Fig. 19 F). Dentations of the legs dark brown and tipped with black (ventral dentations in particular). Eyes russet and the antennae very dark reddish brown. Head (Figs. 19 D – E): Oval, about 1.4 x longer than wide, broadest at the eyes and with the genae just slightly narrowed in posterior portion; vertex moderately convex with the coronal line somewhat indented posteriorly and the lateral furrow notably indented throughout almost its entire length. Eyes moderately convex and their diameter corresponding to about 0.43 x length of gena. Antennae almost reaching to posterior margin of mesothorax and roughly equal in length to profemora; the joints slightly unequal in length and irregularly decreasing in length towards tip of antennae. Scapus rectangular, somewhat compressed dorsoventrally and about 1.8 x longer than wide. Thorax: Pronotum roughly rectangular and slightly shorter but notably narrower than head (Fig. 19 E); the anterior margin gently concave and slightly inflated with a distinct transverse furrow behind; the median transverse sulcus weakly arched and just not reaching lateral margins of segment. Meso- and metanotum both with a fairly distinct medio-longitudinal carina. Mesothorax notably constricted anteriorly, somewhat inflated pre-medially and 4.5 x longer than prothorax; the anterior margin of mesonotum with a shallow transverse bulge. Metanotum parallel-sided, about 0.43 x length of mesonotum and 2 x longer than wide. Pleurae simple, the sterna with a shallow medio-longitudinal ridge. Abdomen: Median segment 1.4 x longer than wide and about 0.7 x length of metanotum, almost rectangular with lateral margins very weakly concave. Segments II – VII slightly sub-uniform in diameter, II – IV slightly increasing in length, V as long as IV and VI – VII decreasing in length; II somewhat longer than median segment and IV – V about 2.5 x longer than wide. Terga smooth except for a shallow longitudinal bulge close to lateral margins and V with posterior margin just weakly inflated medially; sterna simple and only with a short median longitudinal carina posteriorly. Praeopercular organ formed by a fairly broad indention of the posterior margin of sternum VII, which has a narrowing pre-posteriorly, as well as a black verrucose swelling in front (Fig. 19 I). Terga VIII and IX uniform in width, VIII less than half as long as VII and IX roughly quadrate in outline and half as long as VIII. Anal segment with basal portion as wide as IX, tectate longitudinally and with a distinct concave medio-lateral excavation (Fig. 19 G); the posterior portion narrowed and unevenly angularly bi-lobate with a small and narrow posteromedian indention (Fig. 19 H). Epiproct very small and fully concealed under anal segment (Fig. 19 H). Cerci conical and tapering towards a fairly pointed tip, almost reaching to posterior margin of anal segment. Gonapophyses VIII moderately elongated and not reaching tip of anal segment, mostly concealed under it. Subgenital plate lanceolate with apical portion slightly but gradually narrowing towards a pointed apex; projecting beyond tip of abdomen by almost the length of terga VIII-X taken together (Figs. 19 G – I). Legs: Moderately stocky with all carinae dentate; the ventral dentations generally more pronounced than the dentations of dorsal carinae. Profemora almost as long as pro- and mesothorax combined, mesofemora slightly shorter than mesothorax, metafemora reaching to posterior margin of abdominal segment IV and metatibiae reaching about one-third along segment VII. Posteroventral carina of profemora with about 22 distinct, triangular and acute teeth, the anteroventral carina with a similar number of somewhat more serrate teeth; posterodorsal carina supplied with about 18 – 20 small denticles. Dentations of mid and hind legs somewhat erratic dorsally; the two outer ventral carinae of meso- and metafemora with the two apical teeth somewhat enlarged and stronger than preceding. Medioventral carina of meso- and metafemora with nine (mesofemora) or 10 – 12 fairly distinct and strong spines, the three or four sub-basal ones of which are most prominent; medioventral carina of meso- and metatibiae spinose as well. Basitarsi with all carinae minutely denticulate (teeth least distinct on probasitarsus) and the dorsal carinae slightly raised and rounded apically; almost as long as remaining joints except claw taken together. Measurements [mm]: Body (incl. subgenital plate) 166.0, body 155.0, pronotum 5.4, mesonotum 29.5, metanotum 13.9, median segment 7.8, profemora 35.7, mesofemora 27.6, metafemora 33.2, protibiae 36.0, mesotibiae 22.7, metatibiae 33.8, antennae 26.0.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF855D10FF52FF2AA125FD66.taxon	discussion	Comments. So far only known from the unique ♀ holotype. Male and egg unknown.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF855D10FF52FF2AA125FD66.taxon	distribution	Distribution. Philippines (Panay [NHMUK – type locality]).	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF825D6FFF52FF2AA67CF9DE.taxon	description	(Fig. 20) HT, ♂: Coll. I. R. Sc. N. B., Philippines, Luzon, Benguet, Tuba, Leg. T. Heitzmann [RBINS]. Differentiation. The ♂ of this new species (the only sex known) is easily distinguished from its Philippine congeneric by the inconspicuous dark green overall colouration and is smaller than the other two Luzonian S. fallax (Brunner v. Wattenwyl, 1907) and S. feruloides (Westwood, 1859). From the first it is easily separated by the more elongate and ovoid head, which is not as distinctly yellow as in fallax and lacks the characteristic black postocular streak seen in that species (only a faint and washed green postocular stripe in tubaense; Figs. 20 C – D), lack of the distinct black posterior transverse band of abdominal segments II – VII seen in fallax (posterior portion only dark brown in tubaense), just very weakly indicated annulations of the meso- and metatibiae (brown with three distinct yellow to orange annulae in fallax), shorter poculum which does not reach to the posterior margin of abdominal tergum IX (Fig. 20 E) and larger, more elongate vomer (Fig. 20 G). From the much slenderer feruloides this ♂ is readily distinguished by lacking the black abdominal terga II-VI, black longitudinal lateral lines of the mesonotum and mesopleurae as well as the having the costal region of the alae not as distinctly striped longitudinally as in feruloides.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF825D6FFF52FF2AA67CF9DE.taxon	etymology	Etymology. named after the Tuba Municipality in southern Benguet Province, Luzon, the type locality of this new species. Neuter.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF825D6FFF52FF2AA67CF9DE.taxon	description	Description. The unique holotype lacks the left front leg and left mesotarsus, but is otherwise in good condition and seems to show no notable discolouration caused by the dehydrating process. ♂ (Fig. 20). Rather small (body length 95.0 mm) and of rather typical form for the genus, but with colouration less complex and more uniform than other Philippine congenerics. Overall colour fairly plain mid to dark green, the abdomen greyish buff with the two terminal terga dark brown; tergum VIII with a light grey lateral marking in basal half. Head greyish ochre, the genae straw-coloured with a bold but faint and washed green postocular streak (Figs. 20 C – D). Pronotum olive with a brownish wash. Tegmina and costal region of alae cream-coloured with the anterior margin broadly white; all main longitudinal veins marked with dark brown; anal fan hyaline with greyish brown veins. Abdominal terga II – V each with a washed blackish posteromedian marking. Profemora ochre, meso- and metafemora green, all with basal portion slightly bluish and apex washed dark brown. Tibiae ochre with a slight greenish hue and three weakly indicated darker annulae. Antennae greyish brown with ventral surface black in basal one-third. Head (Figs. 20 C – D): Oval, somewhat depressed dorsoventrally, only 1.2 x longer than wide, broadest at eyes and slightly narrowing towards posterior. Vertex weakly convex with coronal line and lateral furrow slightly impressed and with a notable median impression just behind frons, that laterally extends into a shallow furrow that reaches to the compound eye. Frons shallowly bi-gibbose and with a small C-shaped impression in front. Eyes very large, projecting more than hemispherical and their diameter corresponding to 0.66 x length of gena. Antennae slightly projecting over posterior margin of abdominal segment III; scapus somewhat narrowed towards base, the pedicellus round in cross-section and barrel-shaped; following joints slightly and irregularly unequal and essentially first increasing, then decreasing in length. Thorax: Pronotum shorter and much narrower than head, basically rounded rectangular in outline and 1.8 x longer than wide; anterior portion somewhat narrowed and lateral margins gently concave; the anterior margin inflated and bulgy medially and with a small pit some distance off each outer angle; transverse median sulcus distinctly impressed, almost straight and almost reaching to lateral margins of segment (Figs. 20 C – D). Mesothorax slender, elongate and uniform in diameter with only a slight widening in posterior portion and at anterior margin; almost 8.5 x longer than prothorax. Mesonotum with a shallow transverse swelling at anterior margin and a few indicated granules in anterodorsal area. Meso- and metasternum with a fine and rather low medio-longitudinal carina. Tegmina spatulate with basal half gradually narrowing and the central hump fairly well developed and obtusely rounded. Alae reaching about three-quarters the way along abdominal segment III. Abdomen: Abdominal segments II – VI slightly sub-equal in length but uniform in diameter and about 5.2 x longer than wide. Tergum V with an obtusely rounded posteromedian swelling. VII only about 0.75 x the length of preceding segments. Sterna II – VII all with a fairly distinct and acute medio-longitudinal carina. Tergum VIII trapezoidal in outline with anterior margin only two-thirds the width of posterior margin; length a little more than half of VII. IX notably longer than VIII, tube-shaped and slightly widened basally; the lateral margins somewhat inward folded. Anal segment only two-thirds the length of IX (Fig. 20 E), similar in width but somewhat narrowing towards the posterior; obtusely tectate longitudinally with the medio-longitudinal keel most pronounced posteriorly; the posterior margin labiate, slightly inflated, glossy and with a widely triangular emargination. Ventral surfaces of posterior margin supplied with small denticles and facing each other at an angle of ca. 70 °. Vomer gently arched towards the left, fairly slender and gradually narrowing towards a short but acute and up-curved terminal hook; the lateral margins notably inflated and the ventral surface with a deep medio-longitudinal furrow (Fig. 20 G). Phallus strongly enlarged and projecting over posterior margin of poculum. Cerci elongate with apex club-like and noticeably projecting beyond tip of abdomen. The poculum rather small, bowl-shaped with the base moderately bulgy and rounded (Fig. 20 E) and the posterior margin narrowly rounded; not reaching to posterior of tergum IX (Fig. 20 G). Legs: Of average length and shape for the genus; dentations weakly developed and minute. Profemora somewhat longer than head, pro- and mesothorax taken together, mesofemora scarcely longer than pro- and mesothorax combined, metafemora slightly projecting over posterior margin of abdominal segment V and metatibiae projecting considerably beyond tip of abdomen. Teeth of profemora strongly reduced. Mid and hind legs with all carinae fairly regularly denticulate; the medioventral carina supplied with about ten (mesofemora) to about twelve small spines. Tarsi all elongate with basitarsus longer than remaining tarsomeres taken together (just scarcely longer in mesotarsi although); ventral carinae of meso- and metabasitarsi supplied with a few minute denticles. Measurements [mm]: Body 95.0, pronotum 2.4, mesonotum 24.0, metanotum 3.9, median segment 7.8, tegmina 5.4, alae 24.0, profemora 32.8, mesofemora 26.8, metafemora 31.5, protibiae 34.8, mesotibiae 25.5, metatibiae 33.8, antennae 62.0.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF825D6FFF52FF2AA67CF9DE.taxon	discussion	Remarks. Female and egg unknown.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
038187C1FF825D6FFF52FF2AA67CF9DE.taxon	distribution	Distribution. Philippines (Luzon, Province Benguet, Tuba Municipality [RBINS – type locality]). Summary The genus Sadyattes Stål, 1875 was taxonomically revised at the species level and illustrations of almost all fifteen known species are provided along with identification keys. Six new species have been described from the Philippines and four species are transferred to Sadyattes from other genera. The previously unrecognised ♀ of the type-species S. borrii is re-described and found to have been described as a distinct species. The previously unknown ♂♂ of two species as well as the eggs of two species are described and illustrated for the first time. Sadyattes was previously only known from a unique ♂ from an unknown locality and thus the taxonomic position and relationships were obscure. The genus was however recognised as a member of the tribe Stephanacridini Günther, 1953 (subfamily Platycraninae; see Hennemann, 2020) and removed from Pharnaciini (Clitumninae) by Hennemann & Conle (2008), where it was placed by Günther (1953). It was until now that additional material has become available which uncovered the identity and type-locality of Sadyattes borrii Stål, 1875, the type-species of the genus, and represented the key to consequently also allowing a new recognition and delimitation of Sadyattes. As a result, it was found that Eucarcharus Brunner v. Wattenwyl, 1907 is a synonym of Sadyattes (syn. nov.) and that the Javanese paralectotype (the only other type specimen besides the lectotype from the Philippines) of its type species E. feruloides (Westwood, 1859), which actually is an endemic of the Philippines, is the opposite sex of Stål’s Sadyattes borrii. Comprehensive new material, especially from the Philippine Islands, has generated as many as six new species from the Philippines that are described herein. This quadruples the number of known species of Sadyattes from throughout the Philippines, since only to species were known previously, namely S. fallax (Brunner v. Wattenwyl, 1907) and S. feruloides (Westwood, 1859) both of which are here transferred from Eucarcharus (comb. nov.). No new species have been revealed by the present study from the other regions of the distributional range of the genus, but the previously unknown ♂ of the Malayan S. incertus (Redtenbacher, 1908; comb. nov.) could be accounted for and described for the first time. The three Bornean species of the genus, all of which are only known from one sex, are discussed and it seems likely that at least one of them represents the previously unknown ♀ of a species only known from the ♂, namely S. nigricornis (Redtenbacher, 1908) and S. decoris Seow-Choen, 2016; comb. nov.). In total the number of known species know counts to fifteen, but there are still noteworthy gaps in our knowledge of Sadyattes, because some of species are currently only known from one sex and the eggs of only seven species are known. Thus, two species are only known from the ♂ and three only from the ♀. The distributional range of Sadyattes is biogeographically interesting in that it comprises great parts of Southeast Asia i. e. islands on the Sunda Shelf (Peninsular Malaysia, Sumatra and Nicobar Islands, Java and Borneo) as well as the Philippine Islands, the latter of which are currently regarded as part of Wallacea (e. g. Vallejo, 2011) as defined by Huxley (1868). The genus has no species in the remainder of Wallacea, e. g. the Maluku Islands or Sulawesi. However, the distribution of Sadyattes fits very well to the definition of Wallace’s Line by Wallace (1876), who placed it southeast of the Philippines. To the current state of knowledge Sadyattes has one species on Java, two on Sumatra (including nearby smaller islands), two in Peninsular Malaysia (one of which also occurs in the Andaman Islands), three species in Borneo and nine species throughout the Philippine Islands. Within the Philippines the distribution of Sadyattes comprises all of the major biogeographical regions with the exceptions of Palawan and the Sulu Islands. However, since there are also representatives in Borneo and the two mentioned Philippine biogeographic regions represent transitions zones between the Philippines and Borneo, it is likely that Sadyattes can also be found there, but there have been no records yet. The small number of known specimens of certain species suggest these to be fairly rare in the corresponding habitats. Only two Philippine species, namely S. maganda sp. nov. from the island of Mindoro and S. mindanaense sp. nov. from Mindanao, appear to be fairly common and abundant in certain localities of their distributional range, which is seen by the large number of specimens that are available for examination. Very little is known about the natural habitats, life-cycles, behaviours or host plants of Sadyattes - species and only two species, S. leytensis (Zompro, 1997; comb. nov.) and S. mindanaense sp. nov., have been reared in captivity in Europe for some generations. Seow-Choen (2021: 773) stated to have reared specimens of S. enganensis (Redtenbacher, 1908) from eggs in captivity and that these accepted guava (Psidium guayava, Myrtaceae), Syzygium aqueum (Myrtaceae) and mango (Mangifera indica, Anarcadiaceae) as alternative food plants. These insects are believed to be rather polyphagous than specialized feeders since in captivity in Europe S. mindanaense sp. nov. frequently accepted various plants as alternative nutrition, including guava (Psidium guayava, Myrtaceae), eucalyptus (Eucalyptus spp., Myrtaceae), hypericum (Hypericum patulum, Hypericaceae), oak (Quercus spp., Fagaceae), beech (Fagus sylvatica, Fagaceae), bramble and raspberry (Rubus spp., Rosaceae), wild roses (Rosa spp., Rosaceae) and also strawberry (Fragaria vesca, Rosaceae). It is hoped that future collections of species of Sadyattes will also contribute to our knowledge of biological aspects. The descriptions of as many as six new species of the rarely known genus Sadyattes from the Philippines, which as stated above quadruple the number of species known from the archipelago, are yet again good evidence for the still poor degree of exploration of the phasmid fauna of the Philippine Islands. Some of these newly described species are stunning in the aspect that the ♀♀ are large and the ♂♂ often colourful and therefore eye-catching insects, which makes it even more remarkable that they have not been discovered earlier. But the proportionally high number of newly discovered species lines up very well with other current studies describing a good number of new species from an individual clade (e. g. Hennemann et al., 2016; Cumming et al., 2023; Hennemann, 2023) and the certitude that still a plethora of new species of Phasmatodea from throughout the Philippines await formal description. Many of these species are endemics of individual islands or certain isolated habitats on islands, which may also be the case for some of the newly described species of Sadyattes, for instance S. maganda sp. nov. on the island of Mindoro, S. panayense sp. nov. on the island of Panay and S. tubaense sp. nov. from Luzon. However, still too little is known about these species, two of which are so far only known from a unique holotype specimen. Unfortunately, the impressive biodiversity of Philippine Phasmatodea is badly threatened by a progressive destruction and loss of forest, which are the natural habitat for most phasmids. This becomes obvious by some alarming figures. From 2001 to 2021 the Philippines have lost 158 kha of humid primary forest, making up 12 % of its total tree cover loss and the total area of humid primary forest in the Philippines has decreased by as much as 3.4 % in this period of less than twenty years. Roughly, 92 % of this loss occurred in areas where the main reason is deforestation by human (Source: https: // globalforestwarch. org, accessed 28.07.2024). Since small and isolated island faunas, of which there are a plethora throughout the Philippine archipelago, are particularly vulnerable, such degrees of deforestations have severe impact on the biodiversity of the Philippines. It is well known that protected forests experience the lowest deforestation rates and thus it must be hoped there will rapidly be considerably more efforts to protect the remaining forest areas and highly tender isolated island faunas within the Philippines to help retain as much as possible of the highly threatened biodiversity of the archipelago, much of which is still so poorly studied. Thus, it is hoped that the descriptions of several large and colourful stick insects such as the species of Sadyattes can support considerations that ensure the survival and protection of endangered areas within the Philippines.	en	Hennemann, Frank H. (2025): A review of Sadyattes Stål, 1875, with the descriptions of six new species from the Philippines (Phasmatodea: Platycraninae: Stephanacridini). Zootaxa 5610 (1): 1-72, DOI: 10.11646/zootaxa.5610.1.1, URL: https://doi.org/10.11646/zootaxa.5610.1.1
