taxonID	type	description	language	source
0394879FFFF9A1571B45FD39FDF2FB59.taxon	diagnosis	Diagnosis — This species resembles C. schoenleinianum (Klotzsch) Farron because of its auriculate to deeply cordate leaf base, but differs by the leaf blade being broader in the basal part, and by having caducous stipules, a serrate leaf margin, a flattened peduncle and shorter and more compact racemes of 3 – 11 cm long.	en	Bissiengou, P., Chatrou, L. W., Wieringa, J. J., Sosef, M. S. M. (2013): Taxonomic novelties in the genus Campylospermum (Ochnaceae). Blumea 58 (1): 1-7, DOI: 10.3767/000651913X667817, URL: https://doi.org/10.3767/000651913x667817
0394879FFFF9A1571B45FD39FDF2FB59.taxon	materials_examined	Type. Strijk 73 (holo WAG; iso LBV), Gabon, Woleu-Ntem, Bordamur concession area, some 40 km from WWF-station, N 1 ° 14 ', E 11 ° 53 ', 10 Oct. 2002.	en	Bissiengou, P., Chatrou, L. W., Wieringa, J. J., Sosef, M. S. M. (2013): Taxonomic novelties in the genus Campylospermum (Ochnaceae). Blumea 58 (1): 1-7, DOI: 10.3767/000651913X667817, URL: https://doi.org/10.3767/000651913x667817
0394879FFFF9A1571B45FD39FDF2FB59.taxon	description	Understorey treelet up to 6 m tall. Stipules caducous, 3 – 5 mm long. Leaves: petiole 0 – 2 mm long; leaf blade elliptic-obovate to oblanceolate, (9 –) 14 – 24 (– 31) by (4 –) 6 – 8 (– 11) cm, base auriculate to deeply cordate, often clasping the twig, apex acute, coriaceous to thick leathery, margin serrate, upper surface glossy green, lower surface dull paler green; midrib flattened above, prominent below, main lateral veins 10 – 20 mm apart, 16 – 19 on either side, curved upward to run parallel to the margin, intermediate lateral veins prominent on both sides, 0 – 2 in between each pair of main laterals, tertiary venation scalariform, very distinct on both sides. Inflorescences terminal or rarely axillary, its main axis 12 – 15 (– 18) cm long, flattened; racemes 1 – 6, 3 – 9 (– 11) cm long; pairwise scales absent; bracts persistent at the base of the raceme and pedicel, triangular, c. 3 – 5 mm long; cymules 5 – 9 mm apart, 2 – 4 (– 6) - flowered. Flowers: pedicel 5 – 7 mm, articulated at 2 – 3 mm from the base; sepals 5, ovate, in flower 7 – 8 by 2 – 3 mm, yellowish green, in fruit persistent and accrescent, 9 – 10 by 3 – 4 mm, bright red; petals 5, obovate, 7 – 12 by 3 – 4 mm, cuneate at base, rounded at apex, bright yellow; stamens 10, caducous, orange-yellow, filaments less than 1 mm long, anthers 3 – 4 mm long, transversely wrinkled, poricidal; ovary c. 2 mm wide; style c. 4 mm long, curved, yellow. Fruits: receptacle a ± flattened-globose shape, c. 5 mm wide, orange-red; drupelets 1 – 3 well developed per receptacle, reniform, c. 8 by 5 mm, black at maturity; cotyledons incumbent, dissimilar in size with a small outer cotyledon.	en	Bissiengou, P., Chatrou, L. W., Wieringa, J. J., Sosef, M. S. M. (2013): Taxonomic novelties in the genus Campylospermum (Ochnaceae). Blumea 58 (1): 1-7, DOI: 10.3767/000651913X667817, URL: https://doi.org/10.3767/000651913x667817
0394879FFFF9A1571B45FD39FDF2FB59.taxon	distribution	Distribution — Equatorial Guinea (Rio Muni) and Gabon (Woleu-Ntem province).	en	Bissiengou, P., Chatrou, L. W., Wieringa, J. J., Sosef, M. S. M. (2013): Taxonomic novelties in the genus Campylospermum (Ochnaceae). Blumea 58 (1): 1-7, DOI: 10.3767/000651913X667817, URL: https://doi.org/10.3767/000651913x667817
0394879FFFF9A1571B45FD39FDF2FB59.taxon	biology_ecology	Ecology — In primary or secondary forest; on brown clay soil; at 475 – 600 m altitude. Phenology — Flowers and mature fruits collected from October to November, flowers also observed in April, which coincides with the two rainy seasons.	en	Bissiengou, P., Chatrou, L. W., Wieringa, J. J., Sosef, M. S. M. (2013): Taxonomic novelties in the genus Campylospermum (Ochnaceae). Blumea 58 (1): 1-7, DOI: 10.3767/000651913X667817, URL: https://doi.org/10.3767/000651913x667817
0394879FFFF9A1571B45FD39FDF2FB59.taxon	conservation	Preliminary IUCN conservation status — VU B 1 / B 2 ab (ii, iii, iv). EOO = 6098 km 2, AOO = 1653 km 2, locations = 7 (cell width = 15.37 km). All collections have been made recently (the oldest one is from 1983). The only collection from Equatorial Guinea is from within a protected area (Inselberg at Akuom). The other ten collections from Gabon are from a fairly restricted area where logging companies operate which may lead to a decline in the AOO and / or extent of suitable habitat and hence we propose the ‘ Vulnerable’ category.	en	Bissiengou, P., Chatrou, L. W., Wieringa, J. J., Sosef, M. S. M. (2013): Taxonomic novelties in the genus Campylospermum (Ochnaceae). Blumea 58 (1): 1-7, DOI: 10.3767/000651913X667817, URL: https://doi.org/10.3767/000651913x667817
0394879FFFF9A1571B45FD39FDF2FB59.taxon	materials_examined	Additional material. EQUATORIAL GUINEA, Rio Muni, Wele Nzas: Parmentier 358 (BRLU), Inselberg d’Akuom, à 1 h 30 de marche (6 km) du village de Nzuameyong, à 25 km d’Anisoc, N 1 ° 50 ' E 10 ° 55 ', 600 m, 13 June 1999 (st). – GABON, Woleu-Ntem: Bissiengou 667 (LBV, WAG), c. 8 km SSW of Mitzic, FOREENEX forest exploitation, N 0 ° 42.9 ' E 11 ° 32.0 ', 547 m, 6 Nov. 2009 (fr); Bissiengou 707 (LBV, WAG), 8.5 km SSW of Mitzic, FOREENEX forest exploitation, N 0 ° 42.5 ' E 11 ° 31.9 ', 501 m, 6 Nov. 2009 (fl); Bissiengou 714 (LBV, WAG), 13 km SE of Mitzic, FOREENEX forest exploitation, road from forestry camp to Madouaka village, N 0 ° 42.4 ' E 11 ° 38.2 ', 480 m, 7 Nov. 2009 (fl); Bissiengou 730 (LBV, WAG), 13 km SE of Mitzic, FOREENEX forest exploitation, road from forestry camp to Madouaka village, N 0 ° 42.1 ' E 11 ° 38.8 ', 490 m, 7 Nov. 2009 (fr); Bissiengou 769 (LBV, WAG), 35 km NNE of Mitzic, forestry road in Bordamur forest exploitation, 2 km from forestry camp, N 1 ° 02.8 ' E 11 ° 43.3 ', 545 m, 8 Nov. 2009 (st); Breteler 8797 (LBV, WAG), 30 – 40 km NE of Saint Germain, E of Okano River, c. N 0 ° 40 ' E 11 ° 40 ', 475 m, 18 Apr. 1988 (fl, fr); A. M. Louis 377 (K, LBV, WAG), Oveng, 65 km N of road Benguié-Lalara, forest exploitation camp, N 0 ° 42 ' E 11 ° 23 ', 500 m, 5 Nov. 1983 (fl, fr); Maas 10052 (LBV, WAG), RN 2, S of Mitzic, FOREENEX- concession, N 0 ° 42.8 ' E 11 ° 38.9 ', 503 m, 30 Oct. 2011 (fl); Reitsma 2493 (WAG), Inventory Oveng, c. 25 km WSW of Mitzic, N 0 ° 44 ' E 11 ° 22 ', 6 Nov. 1986 (fl, fr).	en	Bissiengou, P., Chatrou, L. W., Wieringa, J. J., Sosef, M. S. M. (2013): Taxonomic novelties in the genus Campylospermum (Ochnaceae). Blumea 58 (1): 1-7, DOI: 10.3767/000651913X667817, URL: https://doi.org/10.3767/000651913x667817
0394879FFFF9A1571B45FD39FDF2FB59.taxon	discussion	Note — Especially the auriculate to deeply cordate leaf base is highly reminiscent of C. schoenleinianum, a species confined to western tropical Africa (from Guinea to Ghana).	en	Bissiengou, P., Chatrou, L. W., Wieringa, J. J., Sosef, M. S. M. (2013): Taxonomic novelties in the genus Campylospermum (Ochnaceae). Blumea 58 (1): 1-7, DOI: 10.3767/000651913X667817, URL: https://doi.org/10.3767/000651913x667817
0394879FFFFBA1571B4AFAA8FFD0F9A0.taxon	diagnosis	Diagnosis — A species resembling C. claessensii (De Wild.) Farron, but with persistent, 7 – 15 mm long, stipules and an unbranched and compact inflorescence.	en	Bissiengou, P., Chatrou, L. W., Wieringa, J. J., Sosef, M. S. M. (2013): Taxonomic novelties in the genus Campylospermum (Ochnaceae). Blumea 58 (1): 1-7, DOI: 10.3767/000651913X667817, URL: https://doi.org/10.3767/000651913x667817
0394879FFFFBA1571B4AFAA8FFD0F9A0.taxon	materials_examined	Type. Breteler & Breteler­Klein Breteler 13124 (holo WAG; iso LBV, WAG), Gabon, Moyen-Ogooué, c. 20 – 30 km NNW of Ndjolé, N 0 ° 03 ' E 10 ° 45 ', 2 Oct. 1994.	en	Bissiengou, P., Chatrou, L. W., Wieringa, J. J., Sosef, M. S. M. (2013): Taxonomic novelties in the genus Campylospermum (Ochnaceae). Blumea 58 (1): 1-7, DOI: 10.3767/000651913X667817, URL: https://doi.org/10.3767/000651913x667817
0394879FFFFBA1571B4AF9B2FAAAF797.taxon	description	Tree or treelet, up to 7 m tall. Stipules persistent, narrowly triangular, 7 – 15 cm long. Leaves: petiole canaliculated above, 3 – 7 mm long; leaf blade narrowly elliptic to narrowly elliptic-obovate, 16 – 30 (– 35) by 6 – 10 (– 11) cm, base cuneate to tapering, apex acuminate or sometimes acute, thick leathery to coriaceous, margin serrulate or sometimes entire toward the base, rarely entire, upper surface flat or rarely bullate, dark green, lower surface paler green, both sides glossy, young leaves purplered; midrib flattened above, prominent below, main lateral veins 6 – 23 mm apart, 14 – 27 on either side, prominent on both sides, curved upward to run parallel to the margin, intermediate lateral veins not to slightly prominent on both sides, 0 – 2 in between each pair of main laterals, tertiary venation scalariform, running perpendicular to the midrib, indistinct on the upper side, slightly distinct on the lower. Inflorescences terminal, unbranched, erect, compact, 3 – 13 cm long; peduncle robust; pairwise scales persistent at the base of the peduncle, narrowly triangular; bracts persistent at the base of the cymule, triangular, 2 – 3 mm long; cymules 3 – 5 (– 10) mm apart, 4 – 8 - flowered. Flowers: pedicel 5 – 15 mm, articulated at 3 – 9 mm from the base, pale green; sepals 5, ovate, in flower 7 – 9 by 2 – 3 mm, greenish, in fruit 9 – 10 by 3 – 4 mm, persistent and accrescent, red; petals 5, obovate, yellow, 10 – 15 by 5 – 9 mm, cuneate at base, rounded at apex; stamens 10, pale yellow, filaments less than 1 mm long, anthers 5 – 7 mm long, transversely wrinkled, poricidal; ovary c. 2 mm wide; style 6 – 7 mm long, curved. Fruits: receptacle a ± flattened-globose shape, 3 – 4 mm wide, red; drupelets 1 – 2 well developed per receptacle, ellipsoid, c. 7 by 5 mm, black at maturity; cotyledons incumbent, dissimilar in size with a smaller outer cotyledon.	en	Bissiengou, P., Chatrou, L. W., Wieringa, J. J., Sosef, M. S. M. (2013): Taxonomic novelties in the genus Campylospermum (Ochnaceae). Blumea 58 (1): 1-7, DOI: 10.3767/000651913X667817, URL: https://doi.org/10.3767/000651913x667817
0394879FFFFBA1571B4AF9B2FAAAF797.taxon	distribution	Distribution — Endemic to Gabon (Moyen-Ogooué and Ngounié provinces).	en	Bissiengou, P., Chatrou, L. W., Wieringa, J. J., Sosef, M. S. M. (2013): Taxonomic novelties in the genus Campylospermum (Ochnaceae). Blumea 58 (1): 1-7, DOI: 10.3767/000651913X667817, URL: https://doi.org/10.3767/000651913x667817
0394879FFFFBA1571B4AF9B2FAAAF797.taxon	biology_ecology	Ecology — In primary and secondary forest, sometimes along rivers or streams; at 70 – 400 m altitude. Phenology — Flowers observed from August to November and in April; fruits collected from October to January.	en	Bissiengou, P., Chatrou, L. W., Wieringa, J. J., Sosef, M. S. M. (2013): Taxonomic novelties in the genus Campylospermum (Ochnaceae). Blumea 58 (1): 1-7, DOI: 10.3767/000651913X667817, URL: https://doi.org/10.3767/000651913x667817
0394879FFFFBA1571B4AF9B2FAAAF797.taxon	conservation	Preliminary IUCN conservation status — VU B 1 ab (ii, iii, iv). EOO = 4476 km 2, AOO = 2348 km 2, locations = 10 (cell width = 15.32 km). All known records of this taxon are from areas exposed to mining and / or logging activities rendering a projected decline in its AOO and / or quality of the habitat most likely and hence we propose the ‘ Vulnerable’ category.	en	Bissiengou, P., Chatrou, L. W., Wieringa, J. J., Sosef, M. S. M. (2013): Taxonomic novelties in the genus Campylospermum (Ochnaceae). Blumea 58 (1): 1-7, DOI: 10.3767/000651913X667817, URL: https://doi.org/10.3767/000651913x667817
0394879FFFFBA1571B4AF9B2FAAAF797.taxon	materials_examined	Additional material. GABON, Moyen-Ogooué: Azizet Issembé 176 (LBV, WAG), Camp Mboumi, Base, c. S 0 ° 25 ' E 10 ° 50 ', 16 Aug. 1999 (fl); Breteler 10375 (LBV, WAG), Missanga, 5 – 15 km NNW of Ndjolé, c. E 0 ° 05 ' E 10 ° 45 ', 11 Nov. 1991 (fl); Breteler 10983 (WAG), 5 – 30 km NNW of Ndjolé, c. S 0 ° 05 ' E 10 ° 45 ', 21 Apr. 1992 (fl); Breteler 14655 (WAG), M’Boumi, chantier SHM, c. 30 km S of Ndjolé, on border of the Ogooué river near Ndjolé, S 0 ° 25 ' E 10 ° 50 ', 17 Nov. 1998 (st); Breteler 14658 (LBV, WAG), idem; Dibata 66 (LBV, MO, WAG), ENE de Belle Vue, Layon ×, S 0 ° 35 ' E 10 ° 39 ', 23 Jan. 1987 (fl). Ngounié: Bissiengou 610 (LBV, WAG), along a forestry road of chantier EFT (Exploitation Forestière de Tsanba) starting at Ndjemba village on Fougamou-Lambaréné road, S 1 ° 03.7 ' E 10 ° 28.6 ', 129 m, 29 Oct. 2009 (fl); Bissiengou 627 (LBV, WAG), idem, S 1 ° 10.9 ' E 10 ° 28.4 ', 165 m, 29 Oct. 2009 (fl); Bissiengou 628 (LBV, WAG), idem; Bissiengou 1432 (LBV, WAG), Sindara, après village Matadi 7 route exploitation forestière EGG (ancien IFL), S 1 ° 02.76 ' E 10 ° 42.50 ', 72 m, 22 June 2011 (st); Bissiengou 1451 (LBV, WAG), Fougamou, village Nzemba route du chantier forestier EGBD, entrée école, S 1 ° 03.06 ' E 10 ° 30.37 ', 97 m, 24 June 2011 (st); Bissiengou 1452 (LBV, WAG), idem; Breteler 5725 (WAG), 50 km SE of Lambaréné, c. S 1 ° 04 ' E 10 ° 30 ', 28 Sept. 1968 (fl); Breteler 14052 (WAG), between Yombi and Fougamou, E slope of Koumounabouali ridge, c. S 1 ° 20 ' E 10 ° 40 ', 22 Sept. 1997 (st); Leeuwenberg 13617 (WAG), right bank Ngounié R., SE of Sindara, km 17 SW of Chantier Waka, S 1 ° 13 ' E 10 ° 49 ', 400 m, 22 Sept. 1985 (fl); A. M. Louis 1324 (WAG), 2 – 3 km SE of Forestry Camp Waka situated ± 32 km SE of Sindara, Waka R. Basin, c. S 1 ° 14 ' E 10 ° 53 ', 350 m, 12 Dec. 1983 (fr); Wieringa 4402 (WAG), 2 km on the road branching off near Ikobey to Magonga, S 1 ° 01.67 ' E 10 ° 57.46 ', 200 m, 25 Nov. 2001 (fr).	en	Bissiengou, P., Chatrou, L. W., Wieringa, J. J., Sosef, M. S. M. (2013): Taxonomic novelties in the genus Campylospermum (Ochnaceae). Blumea 58 (1): 1-7, DOI: 10.3767/000651913X667817, URL: https://doi.org/10.3767/000651913x667817
0394879FFFFCA1501B4AFFFBFDAFF924.taxon	diagnosis	Diagnosis — Like the typical subspecies, but the leaf blade bullate between the lateral veins or rarely flat and the scalariform tertiary veins running perpendicular to the main lateral veins. Type. Wieringa 2852 (holo WAG; iso BR, LBV, MO), Gabon, Ogooué-Maritime, Rabi, 0.6 km on the road to platform Rabi 78, S 1 ° 55.1 ', E 9 ° 50.8 ', 4 Oct. 1994. Distribution — Endemic to Gabon (Ogooué-Maritime, western Ngounié and Nyanga provinces). Ecology — In primary, moist and wet forest, in valleys with small streams; on sandy soil; at 3 – 350 m altitude. Phenology — Flowers observed from October to December; fruits collected from October to January. IUCN conservation status — VU B 1 / B 2 ab (ii, iii, iv). EOO = 6862 km 2, AOO = 1545 km 2, locations = 10 (cell width = 12.43 km). This taxon is known only from 15 recent collections, five of which fall within the Loango National Park and the Moukalaba-Doudou National Park. The remaining ten collections originate from logging and oil concessions while even the Loango park is under development threat from Chinese mining companies. Therefore this subspecies is best placed in the category ‘ Vulnerable’. Additional material. GABON, Ngounié: Le Testu 5834 (BR, P), Agouma, c. S 1 ° 36 ' E 10 ° 10 ', Dec. 1925 (st). Nyanga: Jongkind 5734 (LBV, WAG), Doudou Mountains, Chantier SFN-Bakker, S 2 ° 39.2 ' E 10 ° 27.0 ', 180 m, 22 Nov. 2003 (st). Ogooué-Maritime: Arends 635 (WAG), Monts Doudou, W of Doussala and Réserve de Faune de Moukalaba, c. S 2 ° 15 ' E 10 ° 20 ', 350 m, 5 Dec. 1984 (fr); van Bergen 126 (WAG), savannah road to Vera, 32 km E from junction to Mayonami, S 2 ° 43.4 ' E 10 ° 12.2 ', 70 m, 21 Nov. 1995 (fr); Breteler 10152 (WAG), Rabi-Kounga, Echira road, c. S 2 ° 00 ' E 9 ° 50 ', 27 Oct. 1991 (fl); Farron 7416 (P), Koumouloundou, route d’accès au Rembo-Rabi, S 2 ° 00 ' E 9 ° 36 ', 10 June 1970 (fl); Haegens 89 (WAG), Rabi, W of Shell platform 76, S 1 ° 57 ' E 9 ° 51 ', 30 m, 2 Dec. 1993 (fl); van Nek 117 (WAG), Rabi-E, N of Pechoud Camp, S 1 ° 56.5 ' E 9 ° 52.9 ', 26 Oct. 1990 (st); van Nek 289 (LBV, MO, WAG), Rabi-NW, near Rembo Rabi, NW of Rabi site, S 1 ° 53.7 ' E 9 ° 50.7 ', 13 Nov. 1990 (fl); van Nek 560 (LBV, WAG), near Rabi, S 1 ° 57.6 ' E 9 ° 52.8 ', 11 Jan. 1991 (fr); Schoenmaker 34 (WAG), Rabi-Kounga, opposite Buzzichelli, S 1 ° 56 ' E 9 ° 53 ', 15 Oct. 1991 (fl); Schoenmaker 134 (WAG), Rabi-Kounga, direction Echira, c. S 1 ° 59 ' E 9 ° 51 ', 11 Nov. 1991 (fl); J. J. F. E. de Wilde 9843 (LBV, WAG), 30 km S of Rabi, high bank of an affluent of the Echira River, S 2 ° 08 ' E 9 ° 43 ', 15 m, 28 Nov. 1989 (fr); J. J. F. E. de Wilde 11363 (LBV, WAG), Gamba, N’Dogo Lagoon, near Sounga, S 2 ° 25.9 ' E 9 ° 42.7 ', 3 m, 10 Dec. 1994 (fl). Notes — This subspecies can be added to the list of taxa endemic to the greater Loango-Rabi area (Wieringa & Mackinder 2012). It is also an addition to the recently published checklist of Loango National Park (Harris et al. 2012), and provides further evidence for the unique vegetation in this part of Gabon (Wieringa & Sosef 2011). Apart from the fact that the material shows a scalariform venation different from that of the typical subspecies, and a tendency to have more obviously bullate leaves, there seem to be no other differentiating characters. This, together with the parapatric distribution of the two entities, made us decide to distinguish the taxon at the subspecies level.	en	Bissiengou, P., Chatrou, L. W., Wieringa, J. J., Sosef, M. S. M. (2013): Taxonomic novelties in the genus Campylospermum (Ochnaceae). Blumea 58 (1): 1-7, DOI: 10.3767/000651913X667817, URL: https://doi.org/10.3767/000651913x667817
0394879FFFFCA1511B4AF933FE04FDF8.taxon	diagnosis	Diagnosis — A species resembling C. calanthum (Gilg) Farron, but with a glaucous upper leaf surface (at least when dry), longer petiole, and a pendulous inflorescence that carries 0 – 2 (– 3) racemes.	en	Bissiengou, P., Chatrou, L. W., Wieringa, J. J., Sosef, M. S. M. (2013): Taxonomic novelties in the genus Campylospermum (Ochnaceae). Blumea 58 (1): 1-7, DOI: 10.3767/000651913X667817, URL: https://doi.org/10.3767/000651913x667817
0394879FFFFCA1511B4AF933FE04FDF8.taxon	materials_examined	Type. J. J. de Wilde et al. 10165 (holo WAG; iso LBV), Gabon, Estuaire, 1 km W of Nkan, along the road from Assok to Méla, N 0 ° 40 ', E 10 ° 19 ', 23 Jan. 1991.	en	Bissiengou, P., Chatrou, L. W., Wieringa, J. J., Sosef, M. S. M. (2013): Taxonomic novelties in the genus Campylospermum (Ochnaceae). Blumea 58 (1): 1-7, DOI: 10.3767/000651913X667817, URL: https://doi.org/10.3767/000651913x667817
0394879FFFFCA1511B4AF933FE04FDF8.taxon	description	Treelet up to 2 m tall. Stipules caducous, 2 – 3 mm long. Leaves: petiole 7 – 15 (– 20) mm long, canaliculate above; leaf blade narrowly elliptic to narrowly elliptic-obovate, 13 – 25 (– 30) by 4 – 7 (– 8) cm, base cuneate, apex acuminate, leathery to coriaceous, margin serrulate, upper surface glossy, dark green but turning glaucous when dry, lower surface pale green, young leaves reddish; midrib generally prominent on both sides, main lateral veins 10 – 15 mm apart, 14 – 16 on either side, slightly prominent above, prominent below, curved upward to run parallel to the margin, tertiary venation scalariform, running perpendicular to the midrib thus causing the formation of a decreasing series of intermediate lateral veins, indistinct above, distinct below. Inflorescences terminal, pendulous, (9 –) 15 – 33 cm long; peduncle slender; racemes 0 – 2 (– 3), 3 – 6 (– 10) cm long; pairwise scales persistent at the base of peduncle; bracts caducous, triangular, c. 2 – 3 mm long; cymules 0.5 – 1 (– 1.5) cm apart, 1 – 4 - flowered. Flowers: pedicel 3 – 12 mm, articulated at 1 – 3 mm from the base; sepals, ovate, in flower 6 – 7 by 2 – 3 mm, green-yellowish, in fruit 8 – 9 by 3 – 4 mm, persistent and accrescent, bright red; petals obovate, 7 – 9 by 4 – 5 mm, shortly clawed at base, rounded at apex, bright yellow; stamens 10, filaments less than 1 mm long, anthers 5 – 6 mm long, transversely wrinkled, poricidal; ovary c. 2 mm wide; style slender, 6 – 7 mm long. Fruits: receptacle a ± flattened-globose shape, c. 3 mm wide, red; drupelets 1 – 4 well developed per receptacle, reniform, black at maturity; cotyledons accumbent, similar in size.	en	Bissiengou, P., Chatrou, L. W., Wieringa, J. J., Sosef, M. S. M. (2013): Taxonomic novelties in the genus Campylospermum (Ochnaceae). Blumea 58 (1): 1-7, DOI: 10.3767/000651913X667817, URL: https://doi.org/10.3767/000651913x667817
0394879FFFFCA1511B4AF933FE04FDF8.taxon	distribution	Distribution — Continental Equatorial Guinea (Rio Muni), Gabon and south-western Republic of the Congo (Niari).	en	Bissiengou, P., Chatrou, L. W., Wieringa, J. J., Sosef, M. S. M. (2013): Taxonomic novelties in the genus Campylospermum (Ochnaceae). Blumea 58 (1): 1-7, DOI: 10.3767/000651913X667817, URL: https://doi.org/10.3767/000651913x667817
0394879FFFFCA1511B4AF933FE04FDF8.taxon	biology_ecology	Ecology — In primary and secondary forest, on creek banks and adjacent to swampy areas; at 210 – 1000 m altitude. Phenology — Flowers collected in January, April, July and August; fruits collected from March to May and in August.	en	Bissiengou, P., Chatrou, L. W., Wieringa, J. J., Sosef, M. S. M. (2013): Taxonomic novelties in the genus Campylospermum (Ochnaceae). Blumea 58 (1): 1-7, DOI: 10.3767/000651913X667817, URL: https://doi.org/10.3767/000651913x667817
0394879FFFFCA1511B4AF933FE04FDF8.taxon	conservation	Preliminary IUCN conservation status — LC. EOO = 72,683 km 2, AOO = 23,157 km 2, locations = 9 (cell width = 50.79 km). This species, although being sub-endemic to Gabon, has a fairly wide distribution and hence its EOO and AOO are comparatively large, and above the IUCN thresholds. Potential future threats could come from habitat destruction (logging or land transformation) but a fair number of populations are located within protected areas (PN de Monte Alen, Monts de Cristal NP, Waka NP, Lopé Reserve). Therefore, the category of ‘ Least Concern’ is here proposed.	en	Bissiengou, P., Chatrou, L. W., Wieringa, J. J., Sosef, M. S. M. (2013): Taxonomic novelties in the genus Campylospermum (Ochnaceae). Blumea 58 (1): 1-7, DOI: 10.3767/000651913X667817, URL: https://doi.org/10.3767/000651913x667817
0394879FFFFCA1511B4AF933FE04FDF8.taxon	materials_examined	Additional material. CONGO (BRAZZAVILLE), Niari: Bissiengou 1326 (LBV, WAG), route Malinga-Divénié, 4 km du village Mollo, S 2 ° 29.64 ' E 12 ° 09.61 ', 480 m, 13 June 2011 (st). – EQUATORIAL GUINEA, Rio Muni, Centro Sur: Desmet, G. 242 (BRLU), Mundung ‘ Les 4 montagnes’, N 1 ° 38 ' E 10 ° 11 ', 1000 m, 8 Jan. 2003 (fr); Ngomo 418 (BRLU), Parc National de Monte Alen, c. N 1 ° 40 ' E 10 ° 17 ', 30 July 1998 (fl); Pérez Viso 1139 (MA), Parque Nacional de Monte Alén, Esamalang, camino hacia Mondung, c. N 1 ° 33 ' E 10 ° 12 ', 13 May 1999 (fr); Senterre 4122 (BRLU), Parc National de Monte Alen, 11 km à l’Est de la cabaña de Mosumo, N 1 ° 36.4 ' E 10 ° 08.5 ', 12 July 2003 (st); Wilks 1796 (LBV, WAG), Monts de Cristal, 10 km ENE d’Okuamkos, N 1 ° 10 ' E 10 ° 16 ', 12 Aug. 1988 (fl). – GABON, Estuaire: J. J. F. E. de Wilde 10257 (LBV, WAG), Crystal Mountains, 25 km on the road Tchimbélé-Kinguélé, N 0 ° 31 ' E 10 ° 18 ', 360 m, 26 Jan. 1991 (fl). Ngounié: A. M. Louis 3058 (LBV, WAG), route chantier Leroy Massika entre Mouila et Yeno, S 1 ° 40 ' E 11 ° 15 ', 600 m, 27 Apr. 1989 (fr); Bissiengou 1343 (LBV, WAG), route Malinga-Rebé, S 2 ° 23.07 ' E 12 ° 03.35 ', 489 m, 14 June 2011 (st); Bissiengou 1436 (LBV, WAG), Sindara, après village Matadi 7 route exploitation forestière EGG (ancien IFL), S 1 ° 02.26 ' E 10 ° 42.47 ', 49 m, 22 June 2011 (st); Wieringa 5192 (LBV, WAG), upper Waka area, 13 km on IFL forestry road B 2, S 1 ° 20.5 ' E 10 ° 52.2 ', 180 m, 31 Mar. 2004 (fr). Ogooué-Ivindo: Bissiengou 1056 (LBV, WAG), Nord-Est du parc de la Lopé, 25 km du carrefour qui mène à l’ancien Booué Bac, S 0 ° 11.42 ' E 11 ° 50.05 ', 251 m, 6 Mar. 2010 (fr); Dibata 117 (MO, WAG), Chantier Koumameyong, côté rivière-Marécage à raphia, c. N 0 ° 31 ' E 11 ° 55 ', Mar. 1987 (fr). Ogooué-Maritime: van Valkenburg 3162 (BR, LBV, MO, WAG), old logging road leading southward from chantier CBG Peni, S 2 ° 07.76 ' E 10 ° 24.95 ', 210 m, 22 Apr. 2005 (fl). Woleu-Ntem: Bissiengou 961 (LBV, WAG), Parc des Monts de Cristal, le long de la rivière Mbé, piste après la case picnic sur la droite, c. N 0 ° 37 ' E 10 ° 24 ', 13 Feb. 2010 (st); Wieringa 500 (BR, C, LBV, MO, PRE, WAG), Crystal mountains, 5.5 km NNE of Tchimbélé, N 0 ° 40 ' E 10 ° 25 ', 31 Jan. 1990 (fl).	en	Bissiengou, P., Chatrou, L. W., Wieringa, J. J., Sosef, M. S. M. (2013): Taxonomic novelties in the genus Campylospermum (Ochnaceae). Blumea 58 (1): 1-7, DOI: 10.3767/000651913X667817, URL: https://doi.org/10.3767/000651913x667817
0394879FFFFDA151198EFAD2FABFFA7E.taxon	materials_examined	Type: Welwitsch 4604 (holo LISU; iso BM), Angola, Pungo Andongo, 1857.	en	Bissiengou, P., Chatrou, L. W., Wieringa, J. J., Sosef, M. S. M. (2013): Taxonomic novelties in the genus Campylospermum (Ochnaceae). Blumea 58 (1): 1-7, DOI: 10.3767/000651913X667817, URL: https://doi.org/10.3767/000651913x667817
0394879FFFFDA151198EFA7FFC49F92A.taxon	materials_examined	Type: Zenker 1762 (holo P; iso B, COI, G, K, L, LY, P, S, WAG), Cameroon, Bipinde, 1898.	en	Bissiengou, P., Chatrou, L. W., Wieringa, J. J., Sosef, M. S. M. (2013): Taxonomic novelties in the genus Campylospermum (Ochnaceae). Blumea 58 (1): 1-7, DOI: 10.3767/000651913X667817, URL: https://doi.org/10.3767/000651913x667817
0394879FFFFDA151198EF92BFB95F794.taxon	discussion	Note — The name Ouratea dusenii Engl. & Gilg, published in 1903, has been used most often in literature as the correct name for this species. Recently, Bissiengou & Sosef (2008) transferred that name to the genus Campylospermum. However, it now turns out that the name Monelasmum glomeratum Tiegh. (with type specimen Thollon 140) refers to the same taxon and is thus also available. Since it was published in 1902, hence one year before O. dusenii, Monelasmum glomeratum Tiegh. has priority which necessitates the new combination C. glomeratum (Tiegh.) Biss.	en	Bissiengou, P., Chatrou, L. W., Wieringa, J. J., Sosef, M. S. M. (2013): Taxonomic novelties in the genus Campylospermum (Ochnaceae). Blumea 58 (1): 1-7, DOI: 10.3767/000651913X667817, URL: https://doi.org/10.3767/000651913x667817
0394879FFFFDA1511B45FD0AFA3AFB48.taxon	diagnosis	Diagnosis — A species similar to C. paucinervatum Sosef, but leaf blade papery, with an acuminate apex, entire to serrulate margin and 7 – 11 main lateral veins on either side of the midrib; inflorescence lax, 4 – 8 (– 12) cm long, with 1 – 4 (– 6) racemes of 1 – 7 (– 11) cm long; pedicels 10 – 15 mm, articulating at 2 – 5 mm from the base.	en	Bissiengou, P., Chatrou, L. W., Wieringa, J. J., Sosef, M. S. M. (2013): Taxonomic novelties in the genus Campylospermum (Ochnaceae). Blumea 58 (1): 1-7, DOI: 10.3767/000651913X667817, URL: https://doi.org/10.3767/000651913x667817
0394879FFFFDA1511B45FD0AFA3AFB48.taxon	materials_examined	Type. Haegens & v. d. Burgt 106 (holo WAG; iso LBV), Gabon, Ogooué-Maritime, Rabi-Kounga, road to Divangui, S 1 ° 54 ', E 9 ° 55 ', 6 Dec. 1993.	en	Bissiengou, P., Chatrou, L. W., Wieringa, J. J., Sosef, M. S. M. (2013): Taxonomic novelties in the genus Campylospermum (Ochnaceae). Blumea 58 (1): 1-7, DOI: 10.3767/000651913X667817, URL: https://doi.org/10.3767/000651913x667817
0394879FFFFDA1511B45FD0AFA3AFB48.taxon	description	Treelet up to 4 m tall. Stipules caducous, 2 – 3 mm long. Leaves: petiole 2 – 5 mm long; leaf blade narrowly elliptic to narrowly elliptic-obovate, 6 – 15 by 1.5 – 4.5 cm, base cuneate, apex acuminate, papery, margin entire to serrulate, glossy dark green above, paler green below; midrib slightly prominent above, prominent below, main lateral veins 9 – 10 mm apart, 7 – 11 on either side, curved upward to run parallel to the margin, tertiary venation distinct on both sides, scalariform, running perpendicular to the midrib thus causing the formation of a series of slightly prominent intermediate lateral veins. Inflorescences terminal, lax, 4 – 8 (– 12) cm long; peduncle slender; racemes 1 – 4 (– 6), 1 – 7 (– 11) cm long, held ± horizontally, not seldom with secondary branches; pairwise scales at the base of peduncle absent; bracts early caducous; cymules (5 –) 10 – 15 mm apart, 1 – 2 (– 3) - flowered. Flowers: pedicel 4 – 10 (– 15) mm, articulated at 1 – 6 mm from the base; sepals ovate, in flower 5 – 7 by 2 – 3 mm, greenish yellow, in fruit 6 – 7 by 3 – 3.5 mm, persistent and accrescent, red; petals obovate, 5 – 11 by 4 – 7 mm, cuneate at base, rounded to emarginate at apex; stamens 10, yelloworange, filaments less than 1 mm long, anthers 3 – 4 mm long; ovary c. 2 mm wide; style slender, curved, c. 4 mm long. Fruits: receptacle a ± flattened-globose shape, c. 3 mm wide, red; drupelets 1 – 3 well developed per receptacle, ellipsoid, c. 8 by 5 mm, orange to orange-red at maturity; cotyledons incumbent, similar in size.	en	Bissiengou, P., Chatrou, L. W., Wieringa, J. J., Sosef, M. S. M. (2013): Taxonomic novelties in the genus Campylospermum (Ochnaceae). Blumea 58 (1): 1-7, DOI: 10.3767/000651913X667817, URL: https://doi.org/10.3767/000651913x667817
0394879FFFFDA1511B45FD0AFA3AFB48.taxon	distribution	Distribution — Endemic to Gabon, only found in the coastal plain, in the Ogooué-Maritime (Rabi-Kounga) and Estuaire (Mondah forest and Bikele) provinces.	en	Bissiengou, P., Chatrou, L. W., Wieringa, J. J., Sosef, M. S. M. (2013): Taxonomic novelties in the genus Campylospermum (Ochnaceae). Blumea 58 (1): 1-7, DOI: 10.3767/000651913X667817, URL: https://doi.org/10.3767/000651913x667817
0394879FFFFDA1511B45FD0AFA3AFB48.taxon	biology_ecology	Ecology — Primary and secondary, moist forest, near swampy areas; on sandy soil; at 5 – 30 m altitude. Phenology — Flowers collected from August to November, coinciding with the end of the long dry season and start of the main rainy season; mature fruits from November to December.	en	Bissiengou, P., Chatrou, L. W., Wieringa, J. J., Sosef, M. S. M. (2013): Taxonomic novelties in the genus Campylospermum (Ochnaceae). Blumea 58 (1): 1-7, DOI: 10.3767/000651913X667817, URL: https://doi.org/10.3767/000651913x667817
0394879FFFFDA1511B45FD0AFA3AFB48.taxon	conservation	IUCN conservation status — VU. B 1 ab (ii, iii, iv). EOO = 5959 km 2, AOO = 3388 km 2, locations = 4 (cell width = 29.10 km). This species seems to have a disjunct distribution, but this is uncertain because the area between the two occupied areas is not well explored. In a situation involving a disjunct distribution the sliding scale grid size method leads to an overly large estimation of especially the AOO. Some of the occurrences in the vicinity of the capital Libreville are under threat of habitat destruction due to urbanization and overexploitation. This is likely to lead to a decline in the area of occupancy and / or number of populations or even extent of suitable habitat and hence we propose the category ‘ Vulnerable’.	en	Bissiengou, P., Chatrou, L. W., Wieringa, J. J., Sosef, M. S. M. (2013): Taxonomic novelties in the genus Campylospermum (Ochnaceae). Blumea 58 (1): 1-7, DOI: 10.3767/000651913X667817, URL: https://doi.org/10.3767/000651913x667817
0394879FFFFDA1511B45FD0AFA3AFB48.taxon	materials_examined	Additional material. GABON, Estuaire: A. M. Louis 1201 (WAG), secondary forest ± 17 km E of Libreville, S of Bikelé village, N 0 ° 23 ' E 9 ° 35 ', 7 Dec. 1983 (fr); Bissiengou 815 (LBV, WAG), Mondah forest, parcelle des conservateurs, N 0 ° 35 ' E 9 ° 20 ', 10 Nov. 2009 (fl); Breteler & J. J. F. E. de Wilde 386 (BR, C, K, LBV, MO, P, PRE, SRGH, WAG), Mondah forest, 25 km along the road Libreville-Cap Esterias, N 0 ° 32 ' E 9 ° 23 ', 5 m, 2 Sept. 1978 (fl); de Saint Aubin 2076 (P), 18 km E de Libreville, c. N 0 ° 24 ' E 9 ° 34 ', Oct. 1961 (fl); Reitsma 1328 (MA, WAG), between Cap Santa Clara and Cap Esterias, N 0 ° 34 ' E 9 ° 22 ', 15 Aug. 1985 (fl); Wilks 1632 (MO, WAG), Forêt de Mondah, N 0 ° 35 ' E 9 ° 20 ', 16 Sept. 1987 (fl, fr); Wilks 2090 (LBV, MO, P, WAG), Nyonyie survey, around 1600 m on transect S, S 0 ° 02.9 ' E 9 ° 23.0 ', 3 July 1990 (fl, fr); Wilks 2097 (MO, WAG), Nyonyie survey, 2710 m on transect S, S 0 ° 03.5 ' E 9 ° 23.0 ', 4 July 1990 (fl). Ogooué-Maritime: Breteler 10161 (LBV, WAG), Rabi-Kounga, Echira road, c. S 2 ° 00 ' E 9 ° 50 ', 27 Oct. 1991 (fl); J. J. F. E. de Wilde 9725 (WAG), 1 km on the road Rabi-Divangui, S 1 ° 54 ' E 9 ° 53 ', 25 Nov. 1989 (fr); Schoenmaker 135 (WAG), Rabi-Kounga, direction Echira, c. S 1 ° 59 ' E 9 ° 51 ', 11 Nov. 1991 (fl).	en	Bissiengou, P., Chatrou, L. W., Wieringa, J. J., Sosef, M. S. M. (2013): Taxonomic novelties in the genus Campylospermum (Ochnaceae). Blumea 58 (1): 1-7, DOI: 10.3767/000651913X667817, URL: https://doi.org/10.3767/000651913x667817
0394879FFFFEA1521B45FFFBFDF7FF51.taxon	materials_examined	Type: Zenker 3577 (holo BR; iso E, G, S, US), Cameroon, Bipinde, 1908.	en	Bissiengou, P., Chatrou, L. W., Wieringa, J. J., Sosef, M. S. M. (2013): Taxonomic novelties in the genus Campylospermum (Ochnaceae). Blumea 58 (1): 1-7, DOI: 10.3767/000651913X667817, URL: https://doi.org/10.3767/000651913x667817
0394879FFFFEA1521B45FE93FD79FE25.taxon	materials_examined	Type: Bullock 3877 (holo K; iso BR, K), Zambia, Lunzua River, 19 miles W of Abercorn, 1951.	en	Bissiengou, P., Chatrou, L. W., Wieringa, J. J., Sosef, M. S. M. (2013): Taxonomic novelties in the genus Campylospermum (Ochnaceae). Blumea 58 (1): 1-7, DOI: 10.3767/000651913X667817, URL: https://doi.org/10.3767/000651913x667817
0394879FFFFEA1521B45FE28FE89FD96.taxon	materials_examined	Type: Schlieben 6110 (holo B †; iso BR, G, Z), Tanzania, 50 km W. Lindi, Lake Lutamba, Noto Plateau, 9 Mar. 1935.	en	Bissiengou, P., Chatrou, L. W., Wieringa, J. J., Sosef, M. S. M. (2013): Taxonomic novelties in the genus Campylospermum (Ochnaceae). Blumea 58 (1): 1-7, DOI: 10.3767/000651913X667817, URL: https://doi.org/10.3767/000651913x667817
0394879FFFFEA1521B45FDD6FD91FD41.taxon	materials_examined	Type: Welwitsch 4606 (holo BM; iso K), São Tomé and Príncipe, Pico de Papagaio, Sept. 1853.	en	Bissiengou, P., Chatrou, L. W., Wieringa, J. J., Sosef, M. S. M. (2013): Taxonomic novelties in the genus Campylospermum (Ochnaceae). Blumea 58 (1): 1-7, DOI: 10.3767/000651913X667817, URL: https://doi.org/10.3767/000651913x667817
0394879FFFFEA1521B45FC84FD8BFCFE.taxon	materials_examined	Type: Griffon du Bellay 296 (holo P), Gabon, Pyrat, Oct. 1863.	en	Bissiengou, P., Chatrou, L. W., Wieringa, J. J., Sosef, M. S. M. (2013): Taxonomic novelties in the genus Campylospermum (Ochnaceae). Blumea 58 (1): 1-7, DOI: 10.3767/000651913X667817, URL: https://doi.org/10.3767/000651913x667817
0394879FFFFEA1521B45FCFFFF07FC51.taxon	materials_examined	Type: Warnecke 468 (holo Z; iso BM, E, EA, K), Tanzania, Amani, 1907.	en	Bissiengou, P., Chatrou, L. W., Wieringa, J. J., Sosef, M. S. M. (2013): Taxonomic novelties in the genus Campylospermum (Ochnaceae). Blumea 58 (1): 1-7, DOI: 10.3767/000651913X667817, URL: https://doi.org/10.3767/000651913x667817
