taxonID	type	description	language	source
03A2878C2D72FFA4FF2FC67DFB91F800.taxon	description	(Fig. 1 & 2)	en	Vijayakrishnan, Balaji, Shinde, Gaurav Anil (2025): Gangetic Glyptothorax (Siluriformes: Sisoridae): Two new species and revalidation of Glyptothorax dakpathari Tilak & Husain 1976. Zootaxa 5613 (3): 500-518, DOI: 10.11646/zootaxa.5613.3.4, URL: https://doi.org/10.11646/zootaxa.5613.3.4
03A2878C2D72FFA4FF2FC67DFB91F800.taxon	materials_examined	Material examined. Holotype: ZSI / NRC F- 10669, 63.7 mm SL; India: Uttarakhand: Ganges River at Haridwar. Paratypes: 61.5 – 69.9 mm SL; ZSI / NRC F- 10670 (6); India: Uttarakhand: Ganges River at Haridwar.	en	Vijayakrishnan, Balaji, Shinde, Gaurav Anil (2025): Gangetic Glyptothorax (Siluriformes: Sisoridae): Two new species and revalidation of Glyptothorax dakpathari Tilak & Husain 1976. Zootaxa 5613 (3): 500-518, DOI: 10.11646/zootaxa.5613.3.4, URL: https://doi.org/10.11646/zootaxa.5613.3.4
03A2878C2D72FFA4FF2FC67DFB91F800.taxon	diagnosis	Diagnosis. Glyptothorax bhurainu can be distinguished from all Gangetic congeners except G. pectinopterus, G. conirostris, G. brevipinnis, G. dakpathari and G. himalaicus in the presence (vs. absence) of plicate ventral surface on pectoral-fin spine and first pelvic-fin ray (Fig. 3 a & b). It differs from both G. brevipinnis and G. pectinopterus in possessing serrated (vs. smooth) posterior margin of dorsal-fin spine, longer thoracic adhesive apparatus reaching (vs. not reaching) posterior limit of pectoral-fin base, and shorter adipose-fin base (10.3 – 12.5 % SL vs. 13.9 – 18.0). It differs from G. dakpathari in having an elliptical- (vs. ovoid-) shaped thoracic adhesive apparatus with spindle- (vs. ovoid-) shaped central depression (Fig. 4 a), longer anterior nuchal plate element with rounded anterior margin and pale spots spread throughout (vs. shorter anterior nuchal plate element with pointed anterior margin and pale spots only at lateral tips) (Fig. 5 a), and a shorter dorsal-fin spine (12.9 – 14.8 % SL vs. 15.6 – 15.8); from G. conirostris in having an elliptical- (vs. ovoid-) shaped thoracic adhesive apparatus, absence (vs. presence) of medial pit and in the absence (vs. presence) of anteromedial striae of the thoracic adhesive apparatus; and from G. himalaicus in the absence (vs. presence) of irregular blotches on dorso-lateral surface of body, absence (vs. presence) of medial pit, and a shorter maxillary barbel (82.4 – 116.9 % HL vs. 124.0). Glyptothorax bhurainu further differs from G. telchitta and G. botius in having a deeper body (depth at anus 13.6 – 14.6 % SL vs. 9.9 – 13.3) and deeper caudal peduncle (8.4 – 9.4 % SL vs. 3.1 – 5.9); from G. cavia in having elliptical- (vs. rhomboid-) shaped thoracic adhesive apparatus, absence (vs. presence) of anteromedial striae of the thoracic adhesive apparatus, and shorter head (23.6 – 24.1 % SL vs. 29.5); from G. indicus in having unculiferous ridges of thoracic adhesive apparatus not extending (vs. extending) anteriorly onto gular region, and shorter adipose-fin base (10.3 – 12.5 % SL vs. 13.6 – 15.3); from G. garhwali in the absence (vs. presence) of anteromedial striae in thoracic adhesive apparatus, and serrated (vs. smooth) posterior margin of dorsal-fin spine; and from G. gracilis in having a narrow head (head depth 9.3 – 10.9 % SL vs. 11.9 – 13.9), and a larger eye (11.3 – 13.3 % HL vs. 9.4 – 10.8).	en	Vijayakrishnan, Balaji, Shinde, Gaurav Anil (2025): Gangetic Glyptothorax (Siluriformes: Sisoridae): Two new species and revalidation of Glyptothorax dakpathari Tilak & Husain 1976. Zootaxa 5613 (3): 500-518, DOI: 10.11646/zootaxa.5613.3.4, URL: https://doi.org/10.11646/zootaxa.5613.3.4
03A2878C2D72FFA4FF2FC67DFB91F800.taxon	description	Description. Morphometric data in Table 1. Head depressed; body subcylindrical. Dorsal profile rising evenly from tip of snout to origin of dorsal fin, then sloping gently ventral from origin of dorsal fin to end of caudal peduncle. Ventral profile straight till anal-fin base, sloping gently from anal-fin base to end of caudal peduncle. Anus and urogenital openings located almost vertical through middle of adpressed pelvic fin. Skin tuberculate. Lateral line midlateral, complete. Head depressed and narrow, triangular when viewed laterally. Snout prominent. Anterior and posterior nares large and separated only by base of nasal barbel. Gill opening broad, extending from immediately ventral to post-temporal to isthmus. Bony elements of dorsal surface of head covered with thick, tuberculate skin. Eyes small and round, located on dorsal surface of head. Barbels in four pairs. Maxillary barbel long and slender, extending beyond pectoral-fin base. Nasal barbel slender, extending beyond anterior orbital margin. Inner mandibular barbel extending to pectoral-fin base and outer mandibular barbel extending to middle of pectoral-fin base. Mouth inferior, lips papillate, premaxillary tooth band slightly exposed when mouth closed. Oral teeth small and villiform, in irregular rows on all tooth-bearing surfaces. Premaxillary teeth in single broad semi-lunate band. Dentary teeth in single crescent shaped band, consisting of two separate halves tightly bound at midline. Thoracic adhesive apparatus consisting of striae in elliptical field (Figure 4 a) extending from isthmus to posterior limit of pectoral-fin base; anterolateral edges of adhesive apparatus gently convex. Striae uninterrupted, anteromedial striae absent. No medial pit devoid of striae. Dorsal fin located above anterior third of body, with I, 6 * (7) rays; fin margin gently concave; spine short and gently curved, anterior margin smooth, posterior with 4 – 5 serrations. Adipose fin with anterior and posterior margin slightly concave. Caudal fin strongly forked, with equal upper and lower lobes and i, 7,8, i * (7) principal rays. Procurrent rays symmetrical and extending slightly anterior to fin base. Anal-fin base ventral to adipose-fin origin. Anal fin with straight anterior margin and straight or slightly concave posterior margin; with ii, 9 (2) or ii, 10 * (5) rays. Pelvic-fin origin immediately posterior to vertical through posterior end of dorsal-fin base. Pelvic fin with slightly convex margin and i, 5 * (7) rays; tip of adpressed fin almost reaching anal-fin origin. Pectoral fin with I, 9 (2) or I, 10 * (5) rays; pectoral spine broad and covered with thick skin. Anterior spine margin smooth, posterior margin with 8 – 9 serrations. Ventral surface of pectoral-fin spine and first pelvic-fin ray with plicae (Figure 3 a & b). Colouration. In 70 % alcohol (Fig 1.): dorsal and lateral surfaces of head bluish grey, fading to yellowish cream on ventral surface. Posterior half of supraoccipital process, and both central portion and lateral tips of nuchal plate elements yellowish-cream. A thin, pale mid-dorsal stripe extending from base of last dorsal-fin ray to caudal peduncle, stripe interrupted in some individuals. Dorsal, pectoral, pelvic and anal fins black at base and anterior margins, yellow on rays, and hyaline distally. Pectoral and pelvic fins with prominent yellow on ventral base. Adipose fin yellow with black at base, black subdistal patch on centre of fin, and hyaline distal margin. Caudal fin with dark black vertical band at its base, outer rays yellow to brown, and inner rays cream or hyaline, tip of lobes hyaline. Maxillary and nasal barbels with grey at base, yellow distally and ventrally. Mandibular barbels yellow. Colouration in life: dorsal and lateral surfaces of body and head brown to dark brown, with darker fins (Fig 2.).	en	Vijayakrishnan, Balaji, Shinde, Gaurav Anil (2025): Gangetic Glyptothorax (Siluriformes: Sisoridae): Two new species and revalidation of Glyptothorax dakpathari Tilak & Husain 1976. Zootaxa 5613 (3): 500-518, DOI: 10.11646/zootaxa.5613.3.4, URL: https://doi.org/10.11646/zootaxa.5613.3.4
03A2878C2D72FFA4FF2FC67DFB91F800.taxon	etymology	Etymology. The specific name comes from the Garhwali word ‘ bhurainu’ meaning the one who is brown, in allusion to the live colouration of this species. Garhwali is the primary language in the Garhwal region of Uttarakhand State, India, where the type locality is located.	en	Vijayakrishnan, Balaji, Shinde, Gaurav Anil (2025): Gangetic Glyptothorax (Siluriformes: Sisoridae): Two new species and revalidation of Glyptothorax dakpathari Tilak & Husain 1976. Zootaxa 5613 (3): 500-518, DOI: 10.11646/zootaxa.5613.3.4, URL: https://doi.org/10.11646/zootaxa.5613.3.4
03A2878C2D72FFA4FF2FC67DFB91F800.taxon	biology_ecology	Habitat. Glyptothorax bhurainu was collected from a shallow, fast-flowing stream with a predominantly sandy bottom with cobblestone and pebbles as major substrates (Fig. 6). Co-occurring fishes include Glyptothorax himalaicus, Garra sp., Botia almorhae, Labeo sp., and Tariqilabeo latius.	en	Vijayakrishnan, Balaji, Shinde, Gaurav Anil (2025): Gangetic Glyptothorax (Siluriformes: Sisoridae): Two new species and revalidation of Glyptothorax dakpathari Tilak & Husain 1976. Zootaxa 5613 (3): 500-518, DOI: 10.11646/zootaxa.5613.3.4, URL: https://doi.org/10.11646/zootaxa.5613.3.4
03A2878C2D72FFA4FF2FC67DFB91F800.taxon	distribution	Distribution. Glyptothorax bhurainu is currently known only from the upper reaches of the Ganges River basin in Haridwar District, Uttarakhand, northern India (Fig. 7). The Ganges at the type locality is formed by the confluence of the Alaknanda and Bhagirathi Rivers, approximately 100 kms upstream.	en	Vijayakrishnan, Balaji, Shinde, Gaurav Anil (2025): Gangetic Glyptothorax (Siluriformes: Sisoridae): Two new species and revalidation of Glyptothorax dakpathari Tilak & Husain 1976. Zootaxa 5613 (3): 500-518, DOI: 10.11646/zootaxa.5613.3.4, URL: https://doi.org/10.11646/zootaxa.5613.3.4
03A2878C2D7BFFACFF2FC321FC2FFE07.taxon	description	urn: lsid: zoobank. org: act: 9 C 1 C 7 EAC-A 086 - 4884 - 9 CE 5 - 2391 C 5 D 6784 B (Fig. 8 & 9)	en	Vijayakrishnan, Balaji, Shinde, Gaurav Anil (2025): Gangetic Glyptothorax (Siluriformes: Sisoridae): Two new species and revalidation of Glyptothorax dakpathari Tilak & Husain 1976. Zootaxa 5613 (3): 500-518, DOI: 10.11646/zootaxa.5613.3.4, URL: https://doi.org/10.11646/zootaxa.5613.3.4
03A2878C2D7BFFACFF2FC321FC2FFE07.taxon	materials_examined	Type material. Holotype: ZSI / NRC F- 10671, 92.2 mm SL; India: Uttarakhand: Ganges River at Haridwar.	en	Vijayakrishnan, Balaji, Shinde, Gaurav Anil (2025): Gangetic Glyptothorax (Siluriformes: Sisoridae): Two new species and revalidation of Glyptothorax dakpathari Tilak & Husain 1976. Zootaxa 5613 (3): 500-518, DOI: 10.11646/zootaxa.5613.3.4, URL: https://doi.org/10.11646/zootaxa.5613.3.4
03A2878C2D7BFFACFF2FC321FC2FFE07.taxon	diagnosis	Diagnosis. Glyptothorax himalaicus can be distinguished from all Gangetic congeners except G. pectinopterus, G. brevipinnis, G. conirostris, G. bhurainu and G. dakpathari in the presence (vs. absence) of plicate ventral surface on pectoral-fin spine and first pelvic-fin ray (Fig. 3 c & d). It can be readily distinguished from all of the abovementioned species in having the dorso-lateral surface of body covered with irregular blotches (vs. plain body). It further differs from G. brevipinnis and G. pectinopterus in possessing serrated (vs. smooth) posterior margin of dorsal-fin spine and longer thoracic adhesive apparatus reaching (vs. not reaching) posterior limit of pectoral-fin base; from G. dakpathari in having thoracic adhesive apparatus open caudally (vs. completely encinctured by striae) (Fig. 4 b), and longer anterior nuchal plate element with rounded anterior margin and pale spots spread throughout (vs. shorter anterior nuchal plate element with pointed anterior margin and pale spots only at lateral tips) (Fig. 5 b); and from G. conirostris in having elliptical- (vs. ovoid-) shaped thoracic adhesive apparatus and in the absence (vs. presence) of anteromedial striae on the thoracic adhesive apparatus. Glyptothorax himalaicus further differs from G. telchitta and G. botius in having longer pectoral-fin spine (18.3 % SL vs. 12.3 – 17.0) and deeper caudal peduncle (7.9 % SL vs. 3.1 – 5.9); from G. cavia in having elliptical- (vs. rhomboid-) shaped thoracic adhesive apparatus, absence (vs. presence) of anteromedial striae on the thoracic adhesive apparatus, and shorter head (23.1 % SL vs. 29.5); from G. indicus in having unculiferous ridges of thoracic adhesive apparatus not extending (vs. extending) anteriorly onto gular region, and shorter adipose-fin base (11.2 % SL vs. 13.6 – 15.3); from G. garhwali in the absence (vs. presence) of anteromedial striae on thoracic adhesive apparatus and serrated (vs. smooth) posterior margin of dorsal-fin spine; and from G. gracilis in having more slender head (head depth 10.6 % SL vs. 11.9 – 13.9) and larger eye (14.5 % HL vs. 9.4 – 10.8).	en	Vijayakrishnan, Balaji, Shinde, Gaurav Anil (2025): Gangetic Glyptothorax (Siluriformes: Sisoridae): Two new species and revalidation of Glyptothorax dakpathari Tilak & Husain 1976. Zootaxa 5613 (3): 500-518, DOI: 10.11646/zootaxa.5613.3.4, URL: https://doi.org/10.11646/zootaxa.5613.3.4
03A2878C2D7BFFACFF2FC321FC2FFE07.taxon	description	Description. Morphometric data in Table 2. Head depressed; body subcylindrical. Dorsal profile rising evenly from tip of snout to origin of dorsal fin, then sloping gently ventral from origin of dorsal fin to end of caudal peduncle. Ventral profile straight till anal-fin base, sloping gently from anal-fin base to end of caudal peduncle. Anus and urogenital openings located almost vertical through middle of adpressed pelvic fin. Skin tuberculate. Lateral line complete and midlateral. Head depressed and narrow, triangular when viewed laterally. Snout prominent. Anterior and posterior nares large and separated only by base of nasal barbel. Gill opening broad, extending from immediately ventral to post-temporal to isthmus. Bony elements of dorsal surface of head covered with thick, tuberculate skin. Eyes small and round, located on dorsal surface of head. Barbels in four pairs. Maxillary barbel long and slender, extending beyond pectoral-fin base. Nasal barbel slender, extending beyond anterior orbital margin. Inner mandibular barbel extending to pectoral-fin base and outer mandibular barbel extending to middle of pectoral-fin base. Mouth inferior, lips papillate, premaxillary tooth band partially almost wholly exposed when mouth is closed. Oral teeth small and villiform. Premaxillary teeth in a single broad semi-lunate band. Dentary teeth in two narrow crescent shaped bands separated at midline. Thoracic adhesive apparatus consisting of striae in elliptical field (Figure 4 b) extending from isthmus to posterior limit of pectoral-fin base; anterolateral edges of adhesive apparatus gently convex. Striae uninterrupted, anteromedial striae absent. Medial pit present and posteromedial, extending for about half length of thoracic adhesive apparatus. Dorsal fin located above anterior third of body, with I, 6 * (1) rays; fin margin gently concave; spine short and gently curved, anterior margin smooth, posterior with 6 serrations. Adipose fin with anterior margin and posterior margins straight. Caudal fin strongly forked, with i, 7,8, i * (1) principal rays. Procurrent rays symmetrical and extending slightly anterior to fin base. Anal-fin base ventral to adipose-fin origin. Anal fin with straight anterior margin and straight or slightly concave posterior margin; with ii, 10 * (1) rays. Pelvic-fin origin immediately posterior to vertical through posterior end of dorsal-fin base. Pelvic fin with slightly convex margin and i, 6 * (1) rays; tip of adpressed fin almost reaching anal-fin origin. Pectoral fin with I, 9 * (1); pectoral spine broad and covered with thick skin. Anterior spine margin smooth, posterior margin with 13 serrations. Ventral surface of pectoral-fin spine and first pelvic-fin ray with plicae (Figure 3 c & d). Colouration. In 70 % alcohol (Fig 8): dorsal and lateral surfaces of head grey, with irregular dark blotches, fading to yellowish-cream on ventral surfaces. Posterior half of supraoccipital process and both central portion and lateral tips of nuchal plate elements yellow. A thin, pale mid-dorsal stripe extending from base of last dorsal- fin ray to origin of adipose fin. Dorsal, pectoral, pelvic and anal fins yellow at base, black on anterior margins and rays, and hyaline distally. Adipose fin yellow with hyaline distal margin. Caudal fin with dark black patch at its base, rays yellow with irregular black spots, hyaline distally. Barbels grey dorsally and cream ventrally. Colouration in life similar, with darker body and fins (Fig 9).	en	Vijayakrishnan, Balaji, Shinde, Gaurav Anil (2025): Gangetic Glyptothorax (Siluriformes: Sisoridae): Two new species and revalidation of Glyptothorax dakpathari Tilak & Husain 1976. Zootaxa 5613 (3): 500-518, DOI: 10.11646/zootaxa.5613.3.4, URL: https://doi.org/10.11646/zootaxa.5613.3.4
03A2878C2D7BFFACFF2FC321FC2FFE07.taxon	etymology	Etymology. The specific name comes from the Latin ‘ himalaicus’ meaning of, or pertaining to the Himalayas, in reference to the location of the type locality in Haridwar, at the foothills of the Garhwal Himalayas in Uttarakhand State, India.	en	Vijayakrishnan, Balaji, Shinde, Gaurav Anil (2025): Gangetic Glyptothorax (Siluriformes: Sisoridae): Two new species and revalidation of Glyptothorax dakpathari Tilak & Husain 1976. Zootaxa 5613 (3): 500-518, DOI: 10.11646/zootaxa.5613.3.4, URL: https://doi.org/10.11646/zootaxa.5613.3.4
03A2878C2D7BFFACFF2FC321FC2FFE07.taxon	biology_ecology	Habitat. Glyptothorax himalaicus was collected from a shallow, fast-flowing stream with a predominantly sandy bottom with cobblestone and pebbles as substrates (Fig. 6). Co-occurring fishes include Glyptothorax bhurainu, Garra sp., Botia almorhae, Labeo sp., and Tariqilabeo latius.	en	Vijayakrishnan, Balaji, Shinde, Gaurav Anil (2025): Gangetic Glyptothorax (Siluriformes: Sisoridae): Two new species and revalidation of Glyptothorax dakpathari Tilak & Husain 1976. Zootaxa 5613 (3): 500-518, DOI: 10.11646/zootaxa.5613.3.4, URL: https://doi.org/10.11646/zootaxa.5613.3.4
03A2878C2D7BFFACFF2FC321FC2FFE07.taxon	distribution	Distribution. Glyptothorax himalaicus is currently known only from the upper reaches of the Ganges basin in Haridwar district, Uttarakhand, northern India (Fig. 7). The Ganges at the type locality is formed by the confluence of the Alaknanda and Bhagirathi Rivers, approximately 100 kms upstream.	en	Vijayakrishnan, Balaji, Shinde, Gaurav Anil (2025): Gangetic Glyptothorax (Siluriformes: Sisoridae): Two new species and revalidation of Glyptothorax dakpathari Tilak & Husain 1976. Zootaxa 5613 (3): 500-518, DOI: 10.11646/zootaxa.5613.3.4, URL: https://doi.org/10.11646/zootaxa.5613.3.4
03A2878C2D7FFFAEFF2FC658FE50F8C1.taxon	description	(Fig. 10)	en	Vijayakrishnan, Balaji, Shinde, Gaurav Anil (2025): Gangetic Glyptothorax (Siluriformes: Sisoridae): Two new species and revalidation of Glyptothorax dakpathari Tilak & Husain 1976. Zootaxa 5613 (3): 500-518, DOI: 10.11646/zootaxa.5613.3.4, URL: https://doi.org/10.11646/zootaxa.5613.3.4
03A2878C2D7FFFAEFF2FC658FE50F8C1.taxon	materials_examined	Material examined. ZSI / NRC V- 988 (1), 116.8 mm SL, holotype; ZSI / NRC V- 989 (2), 103.2 – 110.0 mm SL, paratypes; India: Uttarakhand, Yamuna River below Dakpathar barrage, Dehradun District, India.	en	Vijayakrishnan, Balaji, Shinde, Gaurav Anil (2025): Gangetic Glyptothorax (Siluriformes: Sisoridae): Two new species and revalidation of Glyptothorax dakpathari Tilak & Husain 1976. Zootaxa 5613 (3): 500-518, DOI: 10.11646/zootaxa.5613.3.4, URL: https://doi.org/10.11646/zootaxa.5613.3.4
03A2878C2D7FFFAEFF2FC658FE50F8C1.taxon	diagnosis	Diagnosis. Glyptothorax dakpathari can be distinguished from all Gangetic congeners except G. pectinopterus, G. brevipinnis, G. conirostris, G. bhurainu and G. himalaicus in the presence (vs. absence) of plicate ventral surface of pectoral-fin spine and first pelvic-fin ray (Fig. 3 e & f). It differs from both G. brevipinnis and G. pectinopterus in possessing serrated (vs. smooth) posterior margin of dorsal-fin spine, longer thoracic adhesive apparatus reaching (vs. not reaching) posterior limit of pectoral-fin base, and shorter adipose-fin base (9.5 – 11.9 % SL vs. 13.9 – 18.0). It can be differentiated from G. conirostris in having thoracic adhesive apparatus completely encinctured by striae (vs. open caudally), absence (vs. presence) of medial pit and absence (vs. presence) of anteromedial striae on the thoracic adhesive apparatus (Fig. 4 c). Glyptothorax dakpathari further differs from G. telchitta and G. botius in having a deeper body (depth at anus 15.1 – 18.1 % SL vs. 9.9 – 13.3) and deeper caudal peduncle (8.4 – 8.8 % SL vs. 3.1 – 5.9); from G. cavia and G. indicus in having a shorter head (20.7 – 23.1 % SL vs. 27.6 – 29.9), and longer caudal peduncle (21.0 – 22.5 % SL vs. 16.3 – 19.1); from G. garhwali in possessing serrated (vs. smooth) posterior margin of dorsal-fin spine and thoracic adhesive apparatus completely encinctured by striae (vs. open caudally); and from G. gracilis in having a thoracic adhesive apparatus with ovoid (vs. elliptic) central depression, and larger eye (12.2 – 14.4 % HL vs. 9.4 – 10.8).	en	Vijayakrishnan, Balaji, Shinde, Gaurav Anil (2025): Gangetic Glyptothorax (Siluriformes: Sisoridae): Two new species and revalidation of Glyptothorax dakpathari Tilak & Husain 1976. Zootaxa 5613 (3): 500-518, DOI: 10.11646/zootaxa.5613.3.4, URL: https://doi.org/10.11646/zootaxa.5613.3.4
03A2878C2D7FFFAEFF2FC658FE50F8C1.taxon	description	Description. Morphometric data in Table 3. Head depressed; body subcylindrical. Dorsal profile rising evenly from tip of snout to origin of dorsal fin, then sloping gently ventrally from origin of dorsal fin to end of caudal peduncle. Ventral profile straight to anal-fin base, then sloping gently dorsally from anal-fin base to end of caudal peduncle. Anus and urogenital openings located at vertical through middle of adpressed pelvic fin. Skin tuberculate. Lateral line complete and midlateral. Head depressed and narrow, rounded when viewed laterally. Snout prominent and rounded when viewed dorsally. Anterior and posterior nares large and separated only by base of nasal barbel. Gill opening broad, extending from immediately ventral to post-temporal to isthmus. Bony elements of dorsal surface of head covered with thick, tuberculate skin. Eye rounded, located entirely on dorsal half of head. Barbels in four pairs. Maxillary barbel long and slender, extending to middle of pectoral-fin base. Nasal barbel slender, extending to anterior orbital margin. Inner mandibular barbel extending to pectoral-fin base and outer mandibular barbel extending to middle of pectoral-fin base. Mouth inferior, lips papillate, premaxillary tooth band partially almost wholly exposed when mouth closed. Oral teeth small and villiform. Premaxillary teeth in single broad semi-lunate band. Dentary teeth in two narrow crescent-shaped bands separated at midline. Thoracic adhesive apparatus consisting of striae in ovate field (Fig. 4 c) extending from isthmus to beyond posterior limit of pectoral-fin base. Striae uninterrupted, anteromedial striae absent. Medial pit absent. Dorsal fin located above anterior third of body, with I, 5 * (2) or I, 6 (2) rays; fin margin gently concave; spine short and gently curved, anterior margin smooth, posterior with 6 – 9 serrations. Adipose fin with anterior margin and posterior margins straight. Caudal fin strongly forked. Procurrent rays symmetrical and extending slightly anterior to finbase. Anal-fin base ventral to adipose-fin origin. Anal fin with straight anterior margin and straight or slightly concave posterior margin; with ii, 7 (1) or ii, 8 * (2) rays. Pelvic-fin origin immediately posterior to vertical through posterior end of dorsal-fin base. Pelvic fin with slightly convex margin and i, 5 (1) or i, 6 * (2) rays; tip of adpressed fin reaching anal-fin origin. Pectoral fin with I, 10 * (3); pectoral spine broad and covered with thick skin. Anterior spine margin smooth, posterior margin with 11 – 14 serrations. Ventral surface of pectoral-fin spine and first pelvic-fin ray with plicae (Figure 3 e & f). Colouration. In 70 % ethanol: dorsal and lateral surface of head and body brown, fading to beige on ventral surface. All fins brown, with brown rays; outer rays darker than inner rays. Base of dorsal, adipose and caudal fins with pale black marking. Maxillary and nasal barbels brown dorsally, beige ventrally. Mandibular barbels beige.	en	Vijayakrishnan, Balaji, Shinde, Gaurav Anil (2025): Gangetic Glyptothorax (Siluriformes: Sisoridae): Two new species and revalidation of Glyptothorax dakpathari Tilak & Husain 1976. Zootaxa 5613 (3): 500-518, DOI: 10.11646/zootaxa.5613.3.4, URL: https://doi.org/10.11646/zootaxa.5613.3.4
03A2878C2D7FFFAEFF2FC658FE50F8C1.taxon	distribution	Distribution. Glyptothorax dakpathari is currently known only from the type locality, the Yamuna River below Dakpathar Barrage in Dehradun District, Uttarakhand, India (Fig. 7). The Yamuna is a major tributary of the Ganges River in northern India.	en	Vijayakrishnan, Balaji, Shinde, Gaurav Anil (2025): Gangetic Glyptothorax (Siluriformes: Sisoridae): Two new species and revalidation of Glyptothorax dakpathari Tilak & Husain 1976. Zootaxa 5613 (3): 500-518, DOI: 10.11646/zootaxa.5613.3.4, URL: https://doi.org/10.11646/zootaxa.5613.3.4
