identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03A387E5FFAEFFCD4CC9FA6FCB5142BC.text	03A387E5FFAEFFCD4CC9FA6FCB5142BC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudococcus Westwood 1840	<div><p>Genus Pseudococcus Westwood</p><p>Pseudococcus Westwood, 1840: 118, 447 . Type species: Dactylopius longispinus Targioni Tozzetti, 1867: 75, designated by Melville 1983: 77.</p><p>Trechocorys Curtis, 1843: 444 . Type species: Coccus adonidum Linnaeus 1767: 740 .</p><p>Boisduvalia Signoret, 1875: 338 . Type species: Coccus laurinus Boisduval 1867: 353 .</p><p>Oudablis Signoret, 1882: clvii.</p><p>Generic diagnosis (slightly modified from Williams 2004). Body of adult female normally broadly oval; anal lobe bars absent. Antennae each with 7 or 8 segments. Legs well developed, claw without denticle, translucent pores frequently present on hind legs. Circulus present or absent.Anterior and posterior ostioles present. Cerarii numbering 12‒17 pairs, with preocular pair (C 2) always absent. Most cerarii each bearing 2 conical setae, except for those on head and sometimes on thorax, where some with 3 conical setae each. Anal lobe bars absent. Anal lobe cerarii often sclerotized; usually all cerarii containing auxiliary setae, but these sometimes absent anterior to penultimate cerarii (C 17). Oral rim tubular ducts usually present on dorsum, if absent, then some present on venter. Oral collar tubular ducts present on venter, often of several different sizes. Discoidal pores usually present, sometimes 1 or 2 situated adjacent to rim of each oral rim tubular duct, and sometimes next to eyes. Quinquelocular pores always absent.</p></div>	https://treatment.plazi.org/id/03A387E5FFAEFFCD4CC9FA6FCB5142BC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zarkani, Agustin;Watson, Gillian W.;Kaydan, Mehmet Bora	Zarkani, Agustin, Watson, Gillian W., Kaydan, Mehmet Bora (2024): A new species in the mealybug genus Pseudococcus Westwood (Hemiptera: Coccomorpha: Pseudococcidae) from Indonesia. Zootaxa 5555 (4): 590-598, DOI: 10.11646/zootaxa.5555.4.6, URL: https://doi.org/10.11646/zootaxa.5555.4.6
03A387E5FFADFFC94CC9FEACCDEC41A1.text	03A387E5FFADFFC94CC9FEACCDEC41A1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudococcus iffahae Zarkani & Watson & Kaydan 2024	<div><p>Pseudococcus iffahae Zarkani &amp; Kaydan, sp. nov.</p><p>(Figs 1, 2 and 3)</p><p>Material examined. Holotype, adult female, left label: AZ 1220 / 11.xi. 2022 / Indonesia, Sumatra, Bengkulu / Piper aduncum / 3°27’59.1”S / 102°42’59.1”E / 900 m a.s.l; right label: Pseudococcus iffahae Zarkani &amp; Kaydan, 1 ♀ / coll. A. Zarkani / det. M.B. Kaydan. In addition to the holotype specimen (ringed with red ink on the coverslip), the slide mount also contains 2 specimens of Paracoccus sp. (MMUB) .</p><p>Paratypes. INDONESIA: same data as for holotype; 6 ♀♀ on 3 slides, each slide with 2 specimens (AZ1221–1223) (2 slides at MMUB, 1 slide at MZB) .</p><p>Description of adult female (n = 7)</p><p>Appearance in life (Fig. 1). Living on underside of leaves, frequently attended by unidentified ants (not visible in Fig. 1). Body of adult female yellowish-orange in 70% ethanol; in life, body covered by thin layer of white to yellowish powdery wax on all surfaces, with pairs of narrow white wax projections arranged segmentally on body margins, those on posterior end longest (up to half as long as body); at maturity, an ovisac of white wax filaments is produced posteriorly.</p><p>Slide-mounted adult female (Figs 2 and 3). Body elongated oval to broadly oval, 4.24 (3.70–4.25) mm long, 1.83 (1.83–2.05) mm wide. Eyespots present on margin, each about 35 μm wide, without associated discoidal pores. Antenna 8 segmented, 1.25 (1.00–1.25) mm long; each antennal segmental lengths (in μm): segment I, 100 (100–110); II, 100 (100–105); III, 113 (113–118); IV, 75 (75–80); V, 75 (75–80); VI, 63 (63–68); VII, 75 (75–80); VIII (apical segment), 150 (150–155), with 4 fleshy setae each about 35–40 μm long, and an apical seta 40 (38–50) μm long. Clypeolabral shield 210 (200–270) μm long and 180 (150–190) μm wide. Labium 3 segmented, 210 (200–270) μm long, with basal segment 100 (100–130) μm wide. Anterior spiracles each 60 (60–65) μm long and 10 (10–11) μm wide across atrium, posterior spiracles each 70 (70–75) μm long and 15 (15–17) μm wide across atrium. Circulus situated between ventral abdominal segments III and IV, 200 (140–200) μm wide, divided by intersegmental line. Legs well developed; segment lengths for hind leg (in μm): coxa, 230 (200–250); trochanter + femur, 520 (500–540); tibia + tarsus, 580 (560–680); claw 40 (40–50), without a denticle. Ratio of lengths of hind tibia + tarsus to trochanter + femur, 1.12 (1.08–1.12): 1; ratio of lengths of tibia to tarsus, 3.8 (3.6–4): 1; ratio of lengths of trochanter + femur to greatest width of femur, 4.3 (4.2–4.5): 1. Hind legs without translucent pores. Tarsal digitules capitate, each 50–55 μm long. Claw digitules capitate, each about 40.0–42.5 μm long. Anterior and posterior ostioles well developed, each with a total for both lips of 8 (8–13) trilocular pores and 6 or 7 setae. Cerarii numbering 17 pairs, preocular pair (C 2) absent. Anal ring about 100 (100–120) μm long and 140 (140–150) μm wide, with 2 rows of pores and bearing 6 setae, each seta 230–250 μm long.</p><p>Dorsum. Derm membranous. Setae slender and flagellate, mostly each about 100–170 μm long, intermixed with shorter setae of various lengths and some minute setae, each about 10–15 μm long. Multilocular disc-pores absent. Trilocular pores scattered, widely spaced and evenly distributed, each 2.5–3.0 μm in diameter. Oral rim tubular ducts few (numbering 4 or 5), present in sub-median and median areas of thorax and abdomen, each duct 11–13 μm long, with rim about 10 μm in diameter, often each with 1 or 2 discoidal pores adjacent to rim. Oral collar tubular ducts present near margins (Fig. 2D), occurring singly by most abdominal cerarii but becoming more numerous anteriorly (particularly on head), each duct about as wide as a trilocular pore, orifices often with narrow indistinct rims and each associated with 1 or 2 discoidal pores. Anal lobe cerarii well developed (Fig. 3H), each about 20–32 μm long and 12.5 μm wide, set on an almost circular sclerotization with 2 enlarged conical setae, numerous trilocular pores, and 3–5 hair-like auxiliary setae, each 125–138 μm long. Cerarii on more anterior segments each situated on sclerotized and mostly containing 2 (anteriormost 4 pairs of cerarii often each containing 3) conical enlarged setae, 2 or 3 slender straight auxiliary setae and a concentration of trilocular pores.</p><p>Venter. Derm membranous. Setae slender and flagellate, each 15–88 μm long, longest setae located medially on head. Apical setae on anal lobes each 230 (230–320) μm long. Multilocular disc-pores, each 6.3–7.5 μm in diameter, very few, numbering 14 (14–20), present mainly on abdominal segments VII and VIII. Vulva leading to heavily sclerotized internal mushroom-shaped structure (Fig. 2A), about 100–160 μm long and 120–170 μm in diameter. Orifices of oral rim tubular ducts and largest oral collar ducts often each associated with 1 or 2 discoidal pores. Trilocular pores as on dorsum, each 2.5–3.0 μm across, evenly distributed throughout. Oral rim tubular ducts of 2 sizes (Fig. 2C): longer ducts each 11–13 μm long, with rim about 10 μm in diameter (Fig. 3B), and shorter ducts each 8–10 μm long and with rim about 8 μm in diameter (Fig. 3C). Two sizes of oral rim tubular ducts often intermixed, forming a submarginal to marginal cluster of 5–7 anteriorly on prothorax by C 7 and single ducts present in submarginal areas of abdominal segments II–V. Oral collar tubular ducts (Fig 2B) of 3 sizes (Fig. 3 bottom right): (i) largest ducts each 12–13 μm long and about 6 μm wide (Fig. 3D), similar to those on dorsum but often each with an indistinct rim, present on abdominal margins of segments IV‒VII, and on head between antennal bases; (ii) slightly smaller ducts (Fig. 3E), each 10–12 μm long and about 5 μm wide, varying in number, present posterior to vulva and across medial posterior edges of abdominal segments V‒VII; and (iii) minute ducts (Fig. 3F), each 6–7 μm long and about 2.5 μm wide, few, with 1 or 2 situated medially on abdominal segments V and VI.</p><p>Etymology. The species is named iffahae after the collector of the new species, Iffah Izzatun Niswah, daughter of Agustin Zarkani, in recognition of her great assistance in the field during mealybug-collecting activities.</p><p>Host plant. Piper aduncum L. ( Piperaceae) (Fig. 1).</p><p>Distribution. Indonesia (Sumatra, Bengkulu Province).</p><p>Comments. Pseudococcus iffahae sp. nov. is closest to P. philippinicus in having: (i) dorsal oral collar tubular ducts with faint narrow rims; and (ii) vulva leading to a sclerotized internal structure. However, P. iffahae can be distinguished from P. philippinicus by having (character states for P. philippinicus are given in parenthesis): (i) vulva leading to sclerotized internal mushroom-shaped structure (vulva saccate, leading to 4 sclerotized internal pouches); (ii) translucent pores absent from hind legs (translucent pores present on hind femur and tibia), and (iii) anal lobe cerarius with 3–5 hair-like auxiliary setae (with about 8 auxiliary setae).</p><p>The new species is also morphologically close to P. longispinus in having: (i) ventral multilocular disc-pores absent from anterior to abdominal segment VI; (ii) tibia + tarsus of hind legs either mostly same length as trochanter + femur or longer; and (iii) penultimate cerarii (C 17) heavily sclerotized and larger in diameter than anal ring. However, P. iffahae differs by having (character states for P. longispinus given in parenthesis): (i) dorsal oral rim tubular ducts of only one size, these present in submedian to median areas, often each with 1 or 2 discoidal pores adjacent to rim (dorsal oral rim ducts of two sizes present near margin, in groups of 2 or 3, each without discoidal pores adjacent to rim); (ii) dorsal oral collar ducts present around margins (usually absent, occasionally with very few (1 or 2) present near margins); and (iii) hind legs without translucent pores (hind coxa and tibia with translucent pores).</p><p>Pseudococcus iffahae also resembles P. odermatti Miller &amp; Williams in possessing oral rim and oral collar tubular ducts each with discoidal pores adjacent to the rim or orifice on both dorsum and venter. However, P. iffahae can be readily distinguished by having (character states for P. odermatti given in parenthesis): (i) multilocular disc-pores few, on venter of abdominal segments VII and VIII only (multilocular disc-pores numerous, on venter of abdominal segments IV‒IX); (ii) dorsal oral rim tubular ducts few, present on submedian and median areas, absent from margins (dorsal oral rim tubular ducts quite numerous, present on margins and submedian areas only); and (iii) translucent pores absent from hind legs (present on hind femur and tibia).</p></div>	https://treatment.plazi.org/id/03A387E5FFADFFC94CC9FEACCDEC41A1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zarkani, Agustin;Watson, Gillian W.;Kaydan, Mehmet Bora	Zarkani, Agustin, Watson, Gillian W., Kaydan, Mehmet Bora (2024): A new species in the mealybug genus Pseudococcus Westwood (Hemiptera: Coccomorpha: Pseudococcidae) from Indonesia. Zootaxa 5555 (4): 590-598, DOI: 10.11646/zootaxa.5555.4.6, URL: https://doi.org/10.11646/zootaxa.5555.4.6
03A387E5FFA9FFC84CC9FB90CBBE4091.text	03A387E5FFA9FFC84CC9FB90CBBE4091.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudococcus Westwood 1840	<div><p>Key to adult female Pseudococcus in Southeast and southern Asia, adapted from Williams (2004).</p><p>Note: Although it seems likely that P. maritimus is not present in the area covered by the key, the species is included below in case it is found there in the future.</p><p>1(0) Ventral multilocular disc pores present anterior to abdominal segment VI........................................ 6</p><p>- Ventral multilocular disc pores absent from anterior to abdominal segment VI..................................... 2</p><p>2(1) Tibia + tarsus of hind legs noticeably shorter than trochanter + femur. Oral collar tubular ducts absent.................................................................................................. P. leptotrichotus Williams</p><p>- Tibia + tarsus of hind legs either same length as trochanter + femur or longer. Oral collar tubular ducts present, at least on venter.............................................................................................. 3</p><p>3(2) Penultimate cerarii (C 17) at most only lightly sclerotized, smaller in diameter than anal ring...... P. dendrobiorum Williams</p><p>- Penultimate cerarii (C 17) heavily sclerotized, larger in diameter than anal ring...................................... 4</p><p>4(3) Dorsal oral rim tubular ducts absent................................................... P. philippinicus Williams</p><p>- Dorsal oral rim tubular ducts present..................................................................... 5</p><p>5(4) Dorsal oral rim tubular ducts present by margin in groups of 2 or 3, usually with 1 duct larger than other 1 or 2, and without discoidal pores adjacent to rim. Hind coxa and tibia with translucent pores............. P. longispinus (Targioni Tozzetti)</p><p>- Dorsal oral rim tubular ducts present in submarginal and medial areas of abdomen and thorax only, all same size, often each with 1 or 2 discoidal pores adjacent to rim. Hind legs without translucent pores....... P. iffahae Zarkani &amp; Kaydan sp. nov.</p><p>6(1) Dorsal oral collar tubular ducts absent. Vulva normal, without sclerotised internal structures..... P. pseudocitriculus Betrem</p><p>- Dorsal oral collar tubular ducts present, sometimes with faint narrow rims present singly on most segments. Vulva noticeably well developed, with sclerotized internal structures........................................................... 7</p><p>7(6) Eyes each with 1 or more associated discoidal pores.......................................................... 8</p><p>- Eyes each without associated discoidal pores.............................................................. 10</p><p>8(7) Eyes each with a sclerotized rim containing 4–9 discoidal pores...................... P. jackbeardsleyi Gimpel &amp; Miller</p><p>- Eyes each without a sclerotized rim but closely associated with 1–3 discoidal pores................................ 9</p><p>9(8) Hind tibia parallel-sided. Legs extending well beyond margins of body. Normally with an oral rim tubular duct present above each anterior ostiole, next to postocular cerarius. Ventral marginal oral collar tubular ducts opposite each mid-coxa usually numbering more than 5.............................................................. P. maritimus (Ehrhorn)</p><p>- Hind tibia often with slightly convex margins. Legs rarely extending well beyond margins of body. Without an oral rim tubular duct present above each anterior ostiole, next to postocular cerarius. Ventral marginal oral collar tubular ducts opposite each mid-coxa usually numbering fewer than 5.................................................. P. viburni (Signoret)</p><p>10(7) Dorsal setae short, shorter than or same length as ventral setae................................................ 11</p><p>- Dorsal setae longer than ventral setae.................................................................... 13</p><p>11(10) Dorsum without any oral collar tubular ducts (check margins of posteriormost abdominal segments). Hind leg without translucent pores on coxa but pores present on femur and tibia. Ventral submargin of prosoma without any tubular ducts.................................................................................... P. calceolariae (Maskell)</p><p>- Dorsum with at least a few oral collar tubular ducts. Hind leg with translucent pores on coxa and sometimes on femur and/or tibia. Ventral submargin of prosoma with at least a few tubular ducts........................................... 12</p><p>12(11) Dorsal setae very short, many only half as long as medium-sized ventral setae or less. Translucent pores present on hind coxa and sometimes tibia but not on femur. Cerarii usually numbering 16 pairs, ocular pair (C 3) often absent; anal lobe cerarii situated on membranous cuticle. Circulus sometimes absent. Only on Poaceae ................. P. saccharicola Takahashi</p><p>- Dorsal setae only slightly shorter than ventral setae. Translucent pores present on hind coxa, tibia and femur. Cerarii numbering 17 pairs; anal lobe cerarii situated on sclerotised cuticle. Circulus always present. On various hosts, occasionally including Poaceae ........................................................................... P. comstocki (Kuwana)</p><p>13(11) Abdominal segment VII submargin (posterior to posterior ostiole) with tubular duct(s) present. Ventral submargin between prothorax and abdominal segment II with more than 5 oral rim tubular ducts on each side.......................... 14</p><p>- Abdominal segment VII submargin (posterior to posterior ostiole) without tubular duct(s). Ventral submargin between prothorax and abdominal segment II with only 2 or 3 oral rim tubular ducts on each side.................................... 15</p><p>14(13) Dorsal setae about same length as ventral setae. Abdominal segments without dorsal oral rim tubular ducts on midline........................................................................................... P. cryptus Hempel</p><p>- Many dorsal setae obviously longer than ventral setae. A few abdominal segments with dorsal oral rim tubular ducts on midline.................................................. P. gilbertensis Beardsley (= apodemus Williams, 2004)</p><p>15(13) Dorsal oral rim tubular ducts present on abdomen................................................ P. baliteus Lit</p><p>- Dorsal oral rim tubular ducts absent from abdomen....................................... P. aurantiacus Williams</p></div>	https://treatment.plazi.org/id/03A387E5FFA9FFC84CC9FB90CBBE4091	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zarkani, Agustin;Watson, Gillian W.;Kaydan, Mehmet Bora	Zarkani, Agustin, Watson, Gillian W., Kaydan, Mehmet Bora (2024): A new species in the mealybug genus Pseudococcus Westwood (Hemiptera: Coccomorpha: Pseudococcidae) from Indonesia. Zootaxa 5555 (4): 590-598, DOI: 10.11646/zootaxa.5555.4.6, URL: https://doi.org/10.11646/zootaxa.5555.4.6
