taxonID	type	description	language	source
03AD285FFF8ADA1C9C10A8CFFF6130C0.taxon	description	Zoobank LSID: urn: lsid: zoobank. org: act: 8 C 8 D 5 ABA- 3 C 0 B- 4 B 9 E- 8 EBF- 1 D 08 F 56 C 5362.	en	Dai, Rui, Chen, Ziyi, Du, Sile, Ren, Dong, Yao, Yunzhi (2025): New Early Cretaceous pentatomorph bugs from China and the systematic position of Kobdocoridae. Acta Palaeontologica Polonica 70 (1): 17-24, DOI: 10.4202/app.01186.2024, URL: https://doi.org/10.4202/app.01186.2024
03AD285FFF8ADA1C9C10A8CFFF6130C0.taxon	etymology	Etymology: A combination of Latin macula, a spot and Classic Greek coris, true bug, in reference to the presence of the characteristic spots on the hemelytra. Masculine.	en	Dai, Rui, Chen, Ziyi, Du, Sile, Ren, Dong, Yao, Yunzhi (2025): New Early Cretaceous pentatomorph bugs from China and the systematic position of Kobdocoridae. Acta Palaeontologica Polonica 70 (1): 17-24, DOI: 10.4202/app.01186.2024, URL: https://doi.org/10.4202/app.01186.2024
03AD285FFF8ADA1C9C10A8CFFF6130C0.taxon	type_taxon	Type species: Maculocris yixiani Dai, Yao, & Ren sp. nov.; by monotypy; see below.	en	Dai, Rui, Chen, Ziyi, Du, Sile, Ren, Dong, Yao, Yunzhi (2025): New Early Cretaceous pentatomorph bugs from China and the systematic position of Kobdocoridae. Acta Palaeontologica Polonica 70 (1): 17-24, DOI: 10.4202/app.01186.2024, URL: https://doi.org/10.4202/app.01186.2024
03AD285FFF8ADA1C9C10A8CFFF6130C0.taxon	diagnosis	Diagnosis. — As for the type species.	en	Dai, Rui, Chen, Ziyi, Du, Sile, Ren, Dong, Yao, Yunzhi (2025): New Early Cretaceous pentatomorph bugs from China and the systematic position of Kobdocoridae. Acta Palaeontologica Polonica 70 (1): 17-24, DOI: 10.4202/app.01186.2024, URL: https://doi.org/10.4202/app.01186.2024
03AD285FFF8ADA1C9C10A8CFFF6130C0.taxon	discussion	Remarks. — The new genus and Clavaticoris Yao et al., 2012, from the same locality, the two significantly differ in morphological characteristics: head pentagonal, with similar length and width (vs. the latter with a rectangular head, length greater than the width); second segment shorter than the last two segment combined (vs. second segment much greater than the last two segment combined); pronotum callosities present (vs. callosities absent); labium only extends to the metacoxae (vs. labium extending to the third abdominal segment); and the ovipositor long, bisecting the last four abdominal segments (vs. the ovipositor only bisecting the last three abdominal segments). The new genus and Halonatusivena Du et al., 2016, are from the same locality, but are also the most morphologically similar. Both of them have pentagonal head with similar length and width; the length of the second segment is no longer than the sum of the last two segments; both have distinct callosities on the pronotum. These shared characteristics distinguish them from other genera within the family. Therefore, we believe that the new genus is more closely related to Halonatusivena. However, they can be distinctly differentiated based on the following characteristics: the new genus has a larger body size of about 20 mm (vs. medium-sized, less than 10 mm of Halonatusivena); eyes relatively small, anterior almost adjacent to margin of pronotum. (vs. compound eyes are larger and anterior distant from the margin of pronotum of Halonatusivena); the distance between the ocelli is greater than the distance from an ocellus to a compound eye (vs. ocelli are close together, with the distance between them smaller than the distance from an ocellus to a compound eye of Halonatusivena); scutellum punctate, triangulate without median trifurcate carina, covering party the clavus (vs. scutellum without puncture, with a approximately T-shaped carina, narrowing gradually towards the apex and not covering the clavus of Halonatusivena). Stratigraphic and geographic range. — As for the type species.	en	Dai, Rui, Chen, Ziyi, Du, Sile, Ren, Dong, Yao, Yunzhi (2025): New Early Cretaceous pentatomorph bugs from China and the systematic position of Kobdocoridae. Acta Palaeontologica Polonica 70 (1): 17-24, DOI: 10.4202/app.01186.2024, URL: https://doi.org/10.4202/app.01186.2024
03AD285FFF89DA1C9FAFA9E0FA7137BF.taxon	description	Figs. 1 – 3. Zoobank LSID: urn: lsid: zoobank. org: act: C 2 EC 4776 - 4122 - 46 EA-BE 22 - A 3 DFFF 4 B 7 DB 4.	en	Dai, Rui, Chen, Ziyi, Du, Sile, Ren, Dong, Yao, Yunzhi (2025): New Early Cretaceous pentatomorph bugs from China and the systematic position of Kobdocoridae. Acta Palaeontologica Polonica 70 (1): 17-24, DOI: 10.4202/app.01186.2024, URL: https://doi.org/10.4202/app.01186.2024
03AD285FFF89DA1C9FAFA9E0FA7137BF.taxon	etymology	Etymology: Derived from the Yixian Formation, where it was discovered.	en	Dai, Rui, Chen, Ziyi, Du, Sile, Ren, Dong, Yao, Yunzhi (2025): New Early Cretaceous pentatomorph bugs from China and the systematic position of Kobdocoridae. Acta Palaeontologica Polonica 70 (1): 17-24, DOI: 10.4202/app.01186.2024, URL: https://doi.org/10.4202/app.01186.2024
03AD285FFF89DA1C9FAFA9E0FA7137BF.taxon	materials_examined	Type material: Holotype CNU-HET-LB 2024010, male, well preserved bodies, antennae and forewings. Paratypes: CNU-HET-LB 2024011, male; CNU-HET-LB 2024012 – 15 female; CNU-HET-LB 2024016 – 19, male; well preserved bodies, labium, leges and ovipositor from the type locality and horizon. Type locality: Huangbanjigou, Chaomidian Village, Beipiao City, Liaoning Province, China. Type horizon: Yixian Formation, lower Aptian, Lower Cretaceous.	en	Dai, Rui, Chen, Ziyi, Du, Sile, Ren, Dong, Yao, Yunzhi (2025): New Early Cretaceous pentatomorph bugs from China and the systematic position of Kobdocoridae. Acta Palaeontologica Polonica 70 (1): 17-24, DOI: 10.4202/app.01186.2024, URL: https://doi.org/10.4202/app.01186.2024
03AD285FFF89DA1C9FAFA9E0FA7137BF.taxon	diagnosis	Diagnosis. — Body large (Fig. 1 A). Head nearly rounded pentagonal, as long as wide (Fig. 2 B). Eyes relatively small, anterior almost adjacent to margin of pronotum (Fig. 1 A). Length of antennomere II less than the combined length of antennomere III and IV (Fig. 3 B 4). Bucculae reaching at the midline of compound eyes, half the length of head (Fig. 2 B). Labium reaching metacoxae (Fig. 1 C). Ocelli situated closer to eyes than to each other (Fig. 2 A). Pronotum impunctate; callosities well-developed, oval; posterior angles rounded (Fig. 2 A). Scutellum punctate, triangular without sharp vertices, covered part of abdominal (Fig. 3 C). Forewings corium with two rounded markings (Fig. 1 A). Ovipositor long, bisecting the last four abdominal segments (Fig. 1 C, D).	en	Dai, Rui, Chen, Ziyi, Du, Sile, Ren, Dong, Yao, Yunzhi (2025): New Early Cretaceous pentatomorph bugs from China and the systematic position of Kobdocoridae. Acta Palaeontologica Polonica 70 (1): 17-24, DOI: 10.4202/app.01186.2024, URL: https://doi.org/10.4202/app.01186.2024
03AD285FFF89DA1C9FAFA9E0FA7137BF.taxon	description	Description. — Body elongated, about 2.76 times as long as wide. Head pentagonal, slightly shorter than pronotum, length about equal to width. Mandibular plates not projecting past midpoint of clypeus. Antenna four segments, slender, slightly shorter than half of body, antennomere I shortest, slightly exceeding the apex of head, antennomere II longest, antennomere III and IV subequal in length; antennomeres I – III subcylindrical, antennomere IV fusiform. Labium four-segmented, straight, appressed to venter, reaching metacoxae, labiomere I longest, 2.05 times as long as labiomere II, labiomere II – III subequal in length, labiomere IV shortest, tapering apically. Compund eyes relatively small, anterior almost adjacent to margin of pronotum. Ocelli present, closer to eyes than to each other, interocular gap narrower than the diameter of the ocelli. Thorax smooth. Pronotum 1.56 times as wide as long, lateral margin convex, posterior angles feebly rounded. Pronotal collar and carinae absent. Pronotum with two large callosities on anterior area, callus broader than intercallus space. Scutellum subequilateral triangular; about 0.93 times as long as pronotum at midline, 0.85 times as wide as long. Hemelytron poorly preserved, membrane venation not visible. Corium relatively small, almost 1 / 3 length of hemelytron. Corium with two obvious rounded markings, one of oval shapes in 1 / 3 length middle of corium, the other one nearly circular and relatively small on posterior margin of corium. R, M and Cu fuse in the basal of corium. Clavus well developed, clavate, tapers from the base to the apex, claval commissure absent. Legs slender. Femora stout, about 2 times as thick as corresponding tibiae. Hind legs longer than fore and mid legs, metafemur longest, about 1.28 times as long as mesofemur, about 1.09 times as long as profemur, metatibia longest, about 1.27 times as long as mesotibia, about 1.65 times as long as protibia. Tarsi three-segmented, second shortest, third tarsomeres 3.91 times the length of the second. Pulvilli present. Abdomen oval, abdominal connexivum sternites II – VII visible, Ovipositor long, bisecting the last four abdominal segments, about 1 / 3 of body length (Table 1).	en	Dai, Rui, Chen, Ziyi, Du, Sile, Ren, Dong, Yao, Yunzhi (2025): New Early Cretaceous pentatomorph bugs from China and the systematic position of Kobdocoridae. Acta Palaeontologica Polonica 70 (1): 17-24, DOI: 10.4202/app.01186.2024, URL: https://doi.org/10.4202/app.01186.2024
03AD285FFF89DA1C9FAFA9E0FA7137BF.taxon	distribution	Stratigraphic and geographic range. — Yixian Formation, lower Aptian, Lower Cretaceous of Huangbanjigou, Chaomidian Village, Beipiao City, Liaoning Province, China.	en	Dai, Rui, Chen, Ziyi, Du, Sile, Ren, Dong, Yao, Yunzhi (2025): New Early Cretaceous pentatomorph bugs from China and the systematic position of Kobdocoridae. Acta Palaeontologica Polonica 70 (1): 17-24, DOI: 10.4202/app.01186.2024, URL: https://doi.org/10.4202/app.01186.2024
