identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03B0B164090BBD09B8CAFDCAB18A2DEE.text	03B0B164090BBD09B8CAFDCAB18A2DEE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Laemophloeus claudioruyi Bento & Rafael 2025	<div><p>Laemophloeus claudioruyi sp. nov.</p><p>(Figs. 1A–C; 2A, D; 3A, B; 4A)</p><p>Type material. Holotype male deposited at INPA, labeled (Fig. 1C): “ Brasil, Amazonas / Reserva Ducke / 26 Km NE Manaus / Barbosa, M. G. V. ” (white, printed) // “Plot B / Malaise 2 / Abr/95” (white, printed) // “HOLOTYPE / Laemophloeus / claudioruyi sp. nov. / det. M. Bento, 2024” (red, printed and handwritten).</p><p>Diagnosis. Frontoclypeal suture deeply impressed laterally and effaced at middle. Scape entirely cylindrical. Antennomere 3 short, 1.3 times longer than pedicel. Anterior margin of clypeus with mandibular and labral emarginations U-shaped. Lateral margins of pronotum with small, distinct antebasal denticle. Claspers with two transverse, somewhat rhomboidal lobes, with apical inner angles broadly rounded, not produced. Procoxal cavities closed. Parameres fused to basal piece and fused to each other at midline. Endophallus with an extremely elongate, twisted apical sclerite.</p><p>Description of holotype male. Length 1.6 mm, width at mid-elytra 0.54 mm. Body elongate, parallel sided, strongly flat dorsoventrally. Coloration. General color uniformly testaceous, with head slightly darker. Head (Fig. 2A, D) large, 1.9 times wider than long, distinctly wider and slightly shorter than pronotum; surface moderately punctate, with punctures much smaller than eye facets, each bearing a minute seta; eyes laterally protruding, not excavated anteriad. Frontoclypeal suture deeply impressed laterally and completely effaced at middle. Frons quite broad, with interocular width 8.2 times wider than transverse eye diameter; median longitudinal line absent. Clypeus with five distinct anterior emarginations: labral emargination broad, as wide as labrum; mandibular emarginations slightly narrower than labral emargination, moderately deep, forming narrowly rounded outer tooth; antennal emarginations narrow and extremely deep, represented by U-shaped, oblique incisions. Labrum large, as wide as median emargination of clypeus, with anterior margin rounded, not emarginated; surface moderately setose, with long apical setae. Mandibles robust, moderately large, 0.7 times as long as head, conjunctly rounded in closed position. Pregular suture (dividing submentum and gula) extending laterally to form subgenal sutures, reaching inner margin of eyes. Genal processes weakly projected anteriorly, with narrowly rounded apex. Antennae long, distinctly longer than half the body length; scape oblong, cylindrical, short, about 1.8 times longer than wide and shorter than antennomeres 2–3 combined; pedicel globose, about 0.5 times length of scape; flagellum gradually widened toward apex, with antennomeres weakly elongate and narrower basally; antennomere 3 subequal to antennomeres 4–8, not distinctly longer; club 3-segmented, comprised of antennomeres 9–11; antennomeres 9 and 10 subglobose, with lateral sensillar fields; antennomere 11 large and elongate, almost 2 times longer than scape, with a broad sensillar field. Pronotum (Fig. 2A) subtrapezoidal, 1.5 times wider than long, with anterior margin 1.1 times wider than posterior margin. Surface moderately punctate, not microreticulate; punctures small, each bearing a minute seta. Sublateral lines as a wide, deeply impressed groove, with minute, transverse setae. Lateral margins slightly curved, with a weak, shallow posterior notch forming a small antebasal denticle near posterior angle. Anterior angles not produced, narrowly rounded; posterior angles acute, slightly produced laterally. Elytra (Fig. 1A) not maculate, 1.5 times longer than conjunctly wide. First and second cells basally open; third cell closed, barely defined at base. Striae of cells 1–3 complete and well defined. Humeral carina well marked. Lateral side external to humeral carina strongly declivous and not explanate, with surface moderately covered with minute pubescence. Epipleuron gradually narrowed and incomplete, narrower than lateral side of elytron. Thoracic venter (Fig. 1B). Prosternal process broad, with apex expanded posteriad of procoxae, 3.5 times wider than procoxal diameter, with apical margin moderately setose. Procoxal cavities closed. Legs slender, with tibiae distinctly shorter than femurs. Femora robust; metafemur enlarged, 2.8 times longer than wide, with medial width less than half the length. Pro-, meso-, and metatarsus shorter than respective tibiae, with inner surface moderately setose. Tarsal formula 5-5-4. Genitalia (Figs. 3A, B; 4A). Claspers with two transverse, somewhat rhomboidal lobes, with apical inner angles broadly rounded, not produced; apex with surface densely covered with large setal punctures; apical margin straight with a few short setae. Parameres narrowly triangular, fused to basal piece (no suture) and medially fused to each other at basal two-thirds, with apex free and gradually acute; each paramere with two inner setal punctures and two large setal punctures at outer margin. Median lobe with anterior strut 1.4 times longer than tegmen; outer borders wide; apex sharply acute. Endophallus with two rounded, grooved basal sclerites, and an extremely elongate, twisted apical sclerite.</p><p>Female. Unknown.</p><p>Etymology. This species is dedicated to the entomologist Claudio Ruy Vasconcelos da Fonseca, who has contributed enormously to research and education in Amazonas state (Brazil) for the last 40 years.</p><p>Type-locality. Brazil. Amazonas: Manaus.</p><p>Remarks. This species is very similar to Laemophloeus incisus Sharp, 1899 (Figs. 2B, E; 3E, F; 4C) and L. assignatus Bremer, 2025 because of the elytra uniformly light testaceous, not infuscate, antennal emarginations of clypeus incised, club antennomeres simply pubescent, and antennomere 11 elongate (only in major males of L. incisus). However, Laemophloeus claudioruyi sp. nov. is easily distinguished from L. incisus in the following respects (characters of L. incisus parenthesized): antennal scape entirely cylindrical (scape with posterior face flat or slightly concave); antennomere 3 short, 1.3 times longer than pedicel (elongate, almost twice longer than pedicel); anterior margin of clypeus with mandibular and labral emarginations U-shaped (anterior margin of clypeus somewhat Mshaped, with mandibular and labral emarginations almost straight); frontoclypeal suture deeply impressed laterally and effaced at middle (frontoclypeal suture indistinct); lateral margins of pronotum with small, distinct antebasal denticle (lateral margins of pronotum without antebasal denticle); procoxal cavities closed (procoxal cavities widely open); aedeagus with parameres fused to basal piece and fused to each other at midline (parameres not fused to basal piece and completely separated from each other). In addition, Laemophloeus claudioruyi sp. nov. shares with L. assignatus the anterior margin of clypeus with mandibular and labral emarginations U-shaped, but it is distinguished by (characters of L. assignatus parenthesized): frontoclypeal suture deeply impressed laterally and effaced at middle (frontoclypeal suture complete, V-shaped); lateral margins of pronotum with small, distinct antebasal denticle (lateral margins of pronotum without antebasal denticle); procoxal cavities closed (procoxal cavities widely open); and parameres short and barely acute at apex (parameres elongate, slender, sharply acute at apex).</p></div>	https://treatment.plazi.org/id/03B0B164090BBD09B8CAFDCAB18A2DEE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bento, Matheus;Rafael, José Albertino	Bento, Matheus, Rafael, José Albertino (2025): Two new species of the lined flat bark beetle genus Laemophloeus Dejean (Coleoptera: Cucujoidea: Laemophloeidae) from Amazonia. Zootaxa 5679 (3): 431-439, DOI: 10.11646/zootaxa.5679.3.7, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03B0B1640909BD0DB8CAFAFAB1232B32.text	03B0B1640909BD0DB8CAFAFAB1232B32.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Laemophloeus obeliscus Bento & Rafael 2025	<div><p>Laemophloeus obeliscus sp. nov.</p><p>(Figs. 1D–F; 2C, F; 3G, H; 4D)</p><p>Type material. Holotype male deposited at INPA, labeled (Fig. 1F): “ Brasil, AM, Manaus, ZF-2, BR / 174, Km 14, 24.x.2021 / lençol, luz mista, M. Bento, / F. Xavier et al. (legs.)” (white, printed) // “HOLOTYPE / Laemophloeus / obeliscus sp. nov. / det. M. Bento, 2024” (red, handwritten and printed).</p><p>Diagnosis. Frontoclypeal suture deeply impressed, straight. Antennal emarginations of clypeus present. Male antennomeres 4–10 moniliform. Second elytral cell closed. Procoxal cavities closed. Male tarsal formula 5-5-4. Claspers with apex slightly deflected internally, weakly sickle-shaped. Parameres abruptly acute. Endophallus with a long, filament-shaped apical sclerite.</p><p>Description of holotype male. Length 1.5 mm, width at mid-elytra 0.59 mm. Body strongly flat dorsoventrally, robust, parallel sided with lateral margins slightly curved. Coloration. General color uniformly testaceous, with anterior region of head, antennae, and tarsi slightly darker. Head (Fig. 2C, F) large, twice wider than long, slightly wider and slightly shorter than pronotum; surface moderately punctate, with punctures much smaller than eye facets, each bearing a minute seta; eyes laterally protruding, not excavated anteriad. Frontoclypeal suture complete, deeply impressed, laterally connected with lateral lines of head. Frons broad, with interocular width 5.3 times wider than transverse eye diameter; median longitudinal line barely defined. Clypeus with five distinct anterior emarginations: labral emargination broad, as wide as labrum; mandibular emarginations distinctly narrower than labral emargination, moderately deep, forming broadly rounded, obtuse outer tooth; antennal emarginations wide and shallow, not incised. Labrum large, as wide as median emargination of clypeus, with anterior margin rounded, not emargined; surface moderately setose, with long apical setae. Mandibles robust, moderately large, 0.6 times as long as head, conjunctly rounded in closed position. Pregular suture (dividing submentum and gula) extending laterally to genal processes, subgenal sutures absent. Genal processes weakly projected anteriorly, with narrowly rounded apex. Antennae long, as long as half the body length, moniliform; scape oblong, with posterior face flat, short, about 1.6 times longer than wide and shorter than antennomeres 2–3 combined; pedicel globose, about 0.4 times length of scape; flagellum gradually widened toward apex, with antennomeres weakly subglobose and narrower basally; antennomere 3 elongate, distinctly longer than antennomeres 4–8; club 3-segmented comprised of antennomeres 9–11; antennomeres 9 and 10 subglobose, with lateral sensillar fields; antennomere 11 large and elongate, 1.4 times longer than scape, with a broad sensillar field. Pronotum (Fig. 2C) subtrapezoidal, 1.4 times wider than long, with anterior margin 1.1 times wider than posterior margin. Surface moderately punctate, not microreticulate; punctures small, each bearing a minute seta. Sublateral lines as a wide, deeply impressed groove, with minute, transverse setae. Lateral margins slightly curved, with a weak, shallow posterior notch forming a small antebasal denticle near posterior angle. Anterior angles not produced, narrowly rounded; posterior angles acute, slightly produced laterally. Elytra (Fig. 1D) not maculate, 1.5 times longer than conjunctly wide. First cell basally open; second and third cells closed, well defined at base. Striae of cells 1–3 complete, well defined; stria of cell 1 (sutural stria) sparsely punctuated at mid-elytra. Humeral carina well marked. Lateral side external to humeral carina strongly declivous and weakly explanate, with surface moderately covered with minute pubescence. Epipleuron gradually narrowed and incomplete, narrower than lateral side of elytron. Thoracic venter (Fig. 1E). Prosternal process broad, with apex expanded posteriad of procoxae, 3.2 times wider than procoxal diameter, with apical margin moderately setose. Procoxal cavities closed. Legs robust, with tibiae distinctly shorter than femurs. Femurs robust; metafemur enlarged, twice longer than wide. Pro-, meso-, and metatarsus shorter than respective tibiae, with inner surface moderately setose. Tarsal formula 5-5-4. Genitalia (Figs. 3G, H; 4D). Claspers with two triangular, somewhat sickle-shaped lobes, with apex narrowly rounded and slightly deflected internally, bearing 4 large, thick setae; inner margin curved at apical third, with 2 large basal setae. Parameres narrowly triangular, not fused to basal piece and medially free from each other, with apex abruptly acute; each paramere with 7 large setal punctures at basal half. Median lobe with anterior strut extremely elongate, 3.4 times longer than tegmen; apex blunt. Endophallus bearing a long, filament-shaped sclerite enclosed by a large membranous pouch.</p><p>Female. Unknown.</p><p>Etymology. The specific epithet derives from the Greek word “ obeliskos ”, the diminutive form of “ obelos ”, which means “spit, spear”. The term is applied in reference to the sharply pointed parameres of this species.</p><p>Type-locality. Brazil. Amazonas: Manaus.</p><p>Remarks. Among all the species within Laemophloeus (s. str.) possessing 3-segmented antennal club, Laemophloeus obeliscus sp. nov. is superficially similar to Laemophloeus capitesculptus Thomas, 2014 and Laemophloeus corporeflavus Thomas, 2014 (Figs. 3C, D; 4B) because of their general appearance, dorsal color mostly testaceous, and clypeal emarginations shallow. However, the new species can be separated from both by: frontoclypeal suture complete, deeply impressed (laterally effaced, not complete in L. capitesculptus; absent in L. corporeflavus); antennal emarginations of clypeus present (absent in L. capitesculptus; present in L. corporeflavus); male antennomeres 4–10 moniliform (elongate); second elytral cell closed (open in L. capitesculptus and L. corporeflavus); procoxal cavities closed (narrowly open L. capitesculptus and L. corporeflavus); male tarsal formula 5-5-4 (5-5- 5 in L. capitesculptus; 5-5- 4 in L. corporeflavus); and endophallus with a long, filament-shaped apical sclerite (flat, hairy apical sclerite in L. capitesculptus; shared with L. corporeflavus).</p></div>	https://treatment.plazi.org/id/03B0B1640909BD0DB8CAFAFAB1232B32	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bento, Matheus;Rafael, José Albertino	Bento, Matheus, Rafael, José Albertino (2025): Two new species of the lined flat bark beetle genus Laemophloeus Dejean (Coleoptera: Cucujoidea: Laemophloeidae) from Amazonia. Zootaxa 5679 (3): 431-439, DOI: 10.11646/zootaxa.5679.3.7, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03B0B164090DBD02B8CAFD9BB0F62A64.text	03B0B164090DBD02B8CAFD9BB0F62A64.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Laemophloeus Dejean 1835	<div><p>Identification key to Neotropical species of Laemophloeus s. str.</p><p>[modified from Thomas (2013; 2014) and Bento (2021)].</p><p>1. Antennal club composed of 6–8 antennomeres.............................................................. 2</p><p>— Antennal club composed of 3 antennomeres................................................................ 7</p><p>2(1). Elytra maculate........................................................................................5</p><p>— Elytra not maculate, may be infuscate...................................................................... 3</p><p>3(2). Male clypeus with deep antennal emarginations (e. g., Fig. 2A, B). Body color more or less uniformly testaceous or dark testaceous............................................................................................ 4</p><p>— Male clypeus with shallow antennal emarginations (e. g., Fig. 2C). Head and pronotum dark testaceous, elytra testaceous, infuscate medially............................................................... .. L. concinnus Thomas, 2013</p><p>4(3). Frons without V-shaped excavation, with longitudinal line usually deeply impressed. Lateral sides of head not excavated, with eyes not laterally protruding. Antennal emarginations of clypeus rounded.................. L. sexarticulatus Kessel, 1926</p><p>— Frons with a broad, V-shaped excavation, with longitudinal line effaced to barely defined. Lateral sides of head strongly excavated, with laterally protruding, somewhat pedunculated eyes. Antennal emarginations of clypeus somewhat V-shaped................................................................................ .. L. souzalimai Bento, 2021</p><p>5(2). Frons unmodified, not excavate. Ground color of body testaceous. Elytra dark basally and along suture, forming ill-defined, pale, elongate, maculae.......................................................... L. macrognathus Reitter, 1876</p><p>— Frons medially excavate. Ground color of body castaneous or piceous. Elytra with well-defined pale maculae............6</p><p>6(5). Medial excavation of frons broadly V-shaped. Surface of head between punctures smooth, not microreticulate. Genal processes anteriorly projected............................................................. .. L. germaini Grouvelle, 1896</p><p>— Medial excavation of frons as a deep groove. Surface of head between punctures strongly microreticulate. Genal processes unprojected..................................................................... L. buenavista Thomas, 2013</p><p>7(1). Antennal club with complex pubescence. Male anterior margin of clypeus with deep antennal emarginations and paired median horns............................................................................................... 8</p><p>— Antennal club simply pubescent. Male anterior margin of clypeus with or without deep antennal emarginations, without horns................................................................................................9</p><p>8(7). Clypeal horns of major male narrow. Clypeus sinuate laterally between tip of horn and antennal insertion. Female with a clypeal emargination over each mandible.................................................... L. lecontei Grouvelle, 1876</p><p>— Clypeal horns of major male broad. Clypeus straight laterally between tip of horn and antennal insertion. Female clypeus without an emargination over each mandible.............................................. L. taurus Thomas, 2014</p><p>9(7). Male anterior margin of clypeus with deep antennal emarginations (e. g., Fig. 2A, B)................................10</p><p>— Male anterior margin of clypeus without deep antennal emarginations (e. g., Fig. 2C).............................. 13</p><p>10(9). Antennal scape of males with a dorsal setal brush. Elytra with color pattern (Neotropical specimens with circum-scutellar area infuscate)................................................................ .. L. megacephalus Grouvelle, 1876</p><p>— Antennal scape without setal brush. Elytra uniformly light testaceous............................................11</p><p>11(10). Frontoclypeal suture deeply impressed laterally and effaced at middle (Fig. 2A). Antennal scape entirely cylindrical (Fig. 2A, D). Lateral margin of pronotum with antebasal denticle (Fig. 2A). Procoxal cavities closed posteriorly (Fig. 2D)............................................................................................. L. claudioruyi sp. nov.</p><p>— Frontoclypeal suture indistinct (Fig. 2B) or complete. Antennal scape with posterior face flat or slightly concave (Fig. 2B, E). Lateral margin of pronotum without antebasal denticle (Fig. 2B). Procoxal cavities widely open posteriorly (Fig. 2E)..... 12</p><p>12(11). Frontoclypeal suture indistinct (Fig. 2B).Antennomere 11 elongate in major males, as long as antennomeres 9 and 10 combined. Lateral margins of pronotum narrowing posteriorly (Fig. 2B). Male claspers subquadrate (Fig. 3F). Parameres fused to basal piece and fused to each other at middle (Fig. 3E)............................................ L. incisus Sharp, 1899</p><p>— Frontoclypeal suture complete, V-shaped. Antennomere 11 short, distinctly shorter than antennomeres 9 and 10 combined. Lateral margins of pronotum straight to weakly convex. Male claspers with inner angles narrowly rounded, weakly curved. Parameres separated from basal piece and fused to each other only at base................... .. L. assignatus Bremer, 2025</p><p>13(9). Frontoclypeal suture depressed medially to form a distinct triangular fovea in males (females unknown). Elytra without a color pattern.................................................................... ... L. capitesculptus Thomas, 2014</p><p>— Frontoclypeal suture not forming triangular fovea. Elytra with or without color pattern..............................14</p><p>14(13). Elytra uniformly testaceous, without color pattern............................................................15</p><p>— Elytra often with color pattern.......................................................................... 16</p><p>15(14). Body elongate. Frontoclypeal suture barely defined to almost indistinct. Lateral margins of pronotum without distinct antebasal denticle. Elytra not explanate laterally. Procoxal cavities narrowly open posteriorly............ L. mathani Grouvelle, 1889</p><p>— Body oblong-ovate (Fig. 1D). Frontoclypeal suture complete and deeply impressed (Fig. 2C). Lateral margins of pronotum with antebasal denticle (Fig. 2C). Elytra explanate laterally (Fig. 1D, E). Procoxal cavities closed posteriorly (Fig. 2F).............................................................................................. L. obeliscus sp. nov.</p><p>16(14). Frons on each side with a fine diagonal line extending from frontoclypeal suture posteriorly to near eye.................17</p><p>— Frons without diagonal lines............................................................................18</p><p>17(16). Body color testaceous with elytral base and suture darkly infuscate. Antennal club short and flattened. Base of each paramere with four setae.................................................................................. L. planaclavatus Thomas, 2014</p><p>— Body color dark brown, with pale elytral maculae. Antennal club not short and flattened. Base of each paramere with one seta (female unknown)................................................................... L. dozieri Thomas, 2014</p><p>18(16). Head and pronotum darkly infuscate. Elytra infuscate along suture and both basally and apically to form well-defined pale discal maculae........................................................... L. insulatestudinorum Thomas, 2014</p><p>— Head and pronotum usually bright testaceous. Elytra darkly infuscate along suture and/or basally, not forming pale maculae19</p><p>19(18).Frontoclypeal suture present, barely defined at middle. Lateral margins of pronotum irregularly denticulate. Elytra infuscate at base and along sutural margin. Parameres with large basal setae extending well beyond apex of parameres................................................................................................ L. suturalis Reitter, 1876</p><p>— Frontoclypeal suture indistinct. Lateral margins of pronotum mostly inerm. Elytra barely infuscate at base. Parameres without large basal setae.............................................................. L. corporeflavus Thomas, 2014</p></div>	https://treatment.plazi.org/id/03B0B164090DBD02B8CAFD9BB0F62A64	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bento, Matheus;Rafael, José Albertino	Bento, Matheus, Rafael, José Albertino (2025): Two new species of the lined flat bark beetle genus Laemophloeus Dejean (Coleoptera: Cucujoidea: Laemophloeidae) from Amazonia. Zootaxa 5679 (3): 431-439, DOI: 10.11646/zootaxa.5679.3.7, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
