taxonID	type	description	language	source
0D49832FFF8F8209FF68FB82FD57FCF6.taxon	type_taxon	Type species: Siphonaria sipho Sowerby I 1823, by subsequent designation of Gray (1847: 181). — Numerous subsequent references.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF8F8209FF68FB82FD57FCF6.taxon	description	Hubendiculata Iredale & McMichael 1962: 82. Invalid; incorrect subsequent spelling of Hubendickula. Siphonacmea Habe 1958 b: 35. Type species: Acmaea oblongata Yokoyama, 1926, by original designation. — Habe 1962: 96, pl. 44, fig. 15; 1964: 144, pl. 44, fig. 14; Toyohara et al. 2001: 27 – 35.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF8F8209FF68FB82FD57FCF6.taxon	discussion	Taxonomic remarks. The genus Siphonaria was originally described to contain four species, S. sipho, S. concinna, S. tristensis and S. exigua, without an original type species designation. Sowerby referred to two species as ‘ typical species of the genus’ (“ Adanson’s Mouret ” (Le. Lepas mouret Adanson, 1757; unfigured) and “ Patella Sipho ”). However, none of these were available names. Gray (1847: 181) subsequently designated S. sipho as the type species. Anton (1838: 26) designated Siphonaria sowerbyi Michelin, 1832 and Hubendick (1946: 19) designated Siphonaria laciniosa (Linnè, 1758) as type species, but none of these taxa were originally included species rendering these subsequent type designations invalid (Art. 74.1 of the Code). Several subsequent authors incorrectly attributed the genus name Siphonaria to Blainville, 1824, but Blainville (1824 a: 268) explicitly attributed Siphonaria to Sowerby. Consequently, Blainville did not establish a junior homonym of Sowerby’s name. Hubendick (1946: 18) treated Iredale’s taxa Ellsiphon, Legosiphon, Hebesiphon, Mallorisiphon, Mestosiphon, Planesiphon (although not valid), Parellsiphon, Torquisiphon, Triellsiphon, and Pugillaria as junior synonyms of Siphonaria as these were essentially based on conchological attributes. Several authors treated Liriola as a subgenus of Siphonaria (e. g., Suter 1913: 601; Hubendick 1945: 60, 1946: 8 – 9, 18 – 19; Morrison 1963: 7; Abbott 1974: 335). By contrast, Fischer and Crosse (1900: 108) treated Liriola as a distinct genus. Contrary to Hubendick (1945: 15, fig. 1, 1946: 8, fig. 3), two determinant anatomical characters specified for the subgenus Liriola appear not to be diagnostic: The ‘ epiphallus and spermoviduct’ (i. e. ED and CD) do not open separately (i. e., the genital pore is monaulic) in several species and a ‘ spermatheca’ (i. e., BC) is present in the type species S. thersites as well as other species occasionally associated with this taxon (S. obliquata, S. tasmanica, S. funiculata, S. virgulata and S. obvirgulata, S. stowae, S. lateralis, S. acmaeoides, S. capensis, S. belcheri, and S. carbo). Indeed, all species studied herein possess a monoaulic genital opening and a BC as part of the RS. Hubendick (1946: 30) assigned Patellopsis Nobre, 1886 as a section to Liriola; however, incorrectly stated the type species to be S. pectinata (Linnè, 1758). However, the type species is S. algesirae Quoy & Gaimard, 1833 by original designation (Nobre, 1886: 32) although subsequently treated as a synonym of S. pectinata by Hubendick (1946: 43), Morrison (1972: 53), and Giribet & Kawauchi (2015: 5). The subsequent designation of S. lateralis Gould, 1848 as the type species for Kerguelenia Mabille & Rochebrune, 1889 by Hubendick (1945: 60) is invalid because S. lateralis was not originally included in Kerguelenia (Art. 67.2, 69.1 of the Code). Siphonaria redimiculum Reeve, 1856 and S. macgillivrayi Reeve, 1856 are the only originally included species. Powell (1946: 91) subsequently designated S. redimiculum Reeve, 1856 as the type species of Kerguelenia. However, Kerguelenia Mabille & Rochebrune, 1889 is preoccupied by Kerguelenia Stebbing, 1888 (Crustacea: Amphipoda). Finlay (1927: 442) noted the similarity of S. obliquata with Kerguenella, but for being conchologically “ quite aberrant ” placed it in a new genus, Benhamina. Torquisiphon has been treated as a junior synonym of Hebesiphon by Morrison (1972: 60). Hubendick (1946: 22) mentioned Talisiphon tasmanicus as a synonym of Siphonaria tasmanica implying that he likely considered Talisiphon as a junior synonym of Siphonaria. By contrast, Morrison (1963: 7) treated Talisiphon as a section of the subgenus Liriola. Hubendick (1946: 18) treated Mouretus as a synonym of Siphonaria. The type species of Mouretus is S. adansonii Blainville, 1824 by original designation of Blainville (1824 b: 162), not ‘ M. mouretus Blainville’ (Morrison, 1972: 53; a nomen nudum). Hubendickula was ranked as section in Siphonaria s. str. by Hubendick (1955: 5 – 6). The name Planesiphon is unavailable from Iredale (1940: 437, 441) as no type species was designated (Art. 13.1 of the Code). However, Zilch (1959: 86), subsequently made the name available referring to Iredale’s (1940) description while designating Planesiphon elegans as the type species. He treated the taxon as a subgenus of Siphonaria. The prior type species designation of S. elegans Iredale, 1940 by McAlpine (1952: 42) is not valid because Planesiphon was not treated as an accepted name (Art. 69.1 of the Code). Hubendick (1945, 1946, 1955) and Morrison (1963) treated Heterosiphonaria as a valid section of Siphonaria s. str. Benhamina has been ranked as a valid genus by Powell (1939: 217; 1946: 91; 1955: 120; 1979: 293, pl. 54, figs 12 – 13), Borland (1950: 385 – 393), Morton and Miller (1968: 302, 336, 338, 354, 378), and Morley (2004: 130), but as a section in Liriola by Hubendick (1945: 55; 1946: 9, 11, 18, 24 – 26, 64). Our results indicate that the monotypic genus Benhamina is unwarranted because its general morphological and molecular characteristics fall within the range of Siphonaria. Moreover, we synonymize Talisiphon Iredale, 1940 with Siphonaria, and its type species, S. virgulata, with S. funiculata. The type species of Simplisiphonaria Hubendick, 1945 is Siphonaria cookiana Suter, 1909 by original designation. This species is a junior objective synonym of Siphonaria australis Quoy & Gaimard, 1833 because of Boreham’s (1959) subsequent lectotype designation (a juvenile shell of S. australis from a mixed syntype lot; Jenkins, 1983: 1, 21, figs 3 e – g). The type species of the following genus-group taxa are herein considered as junior synonyms of Siphonaria species: Mestosiphon Iredale, 1940 (type species M. eumelas = S. atra); Ellsiphon Iredale, 1940 (type species E. marza = S. denticulata); Triellsiphon Iredale, 1940 (type species Triellsiphon acervus = S. atra); Legosiphon Iredale, 1940 (type species L. optivus = S. viridis); Ductosiphonaria Hubendick, 1945 (type species ‘ S. bifurcata’ [non Reeve, 1856] = S. zelandica); Planesiphon Zilch, 1959 (type species P. elegans Reeve, 1856 = S. zelandica). Siphonaria as delineated herein reveals considerable phylogenetic structure (Fig. 1). However, none of the principal clades retrieved herein is found to exhibit a consistent morphological feature that allows us to consistently distinguish its members from those of any other clade. Therefore, we treat all previously introduced genus-group names listed above as junior synonyms of Siphonaria. Given the lack of clearly identifiable diagnostic characteristics, we do not find it useful or even justifiable to maintain subgenera within Siphonaria for any of the species examined herein. The sole exception is Siphonaria thersites (type species of Liriola). This species is the sister group of all remaining Siphonaria species (Figs 1 – 4) and is distinguished by the combination of a very low-level occurrence intertidally, the animal not being fully enclosed by a reduced and often flattened shell, the BD in the RS totally enclosed within the CD and a relatively small and short SPM. Should these characters be revealed as synapomorphies of a distinct taxonomic lineage that incorporates additional extralimital species, then the name Liriola was available for such a natural group. However, none of the species from the IWP examined herein is member of this potential lineage.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF888207FF68FCC2FA0FF7F6.taxon	description	(Figs 8 – 10, 12 A – F, L – M)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF888207FF68FCC2FA0FF7F6.taxon	materials_examined	Material examined. Type material. Neotype of Siphonaria sipho Sowerby I, 1823, present designation, from Calalinan, Siquijor, Philippines, 9 ° 12.834 ’ N, 123 ° 30.043 ’ E; coll. B. W. Jenkins, 3 April 2019, PHS 02 - 5 (NHMUK 20210025 [M 412, SK 095], Fig. 8 A). Holotype of Siphonaria cornuta Gould, 1846 from ‘ Mangsi’ [= Mangsee] Islands (USNM 5850, Fig. 12 E). Four syntypes of S. siquijorensis Reeve, 1856 from Siquijor (NHMUK 1969166, Figs 8 K – M). Three syntypes of Siphonaria luzonica Reeve, 1856 from Puteao, Luzon, Philippines (NHMUK 1979163, Figs 8 B – D). Syntype of Siphonaria corrugata Reeve, 1856 from Luzon, Philippines (NHMUK 1979165, Fig. 8 E). Other, non-type material. Philippines, Siquijor: Sawang, 09 ° 08.249 ’ N, 123 ° 31.113 ’ E, PHS 02 - 3 (AM C. 585532 20 + p, C. 585108 p [M 404, SK 078], C. 585109 p [M 406, SK 074], C. 585110 p [M 415, SK 098], C. 585111 p [M 477, SK 284], C. 585112 p [M 479, SK 286], C. 585116 p [SK 277], C. 585117 p [SK 278]); Calalinan, 09 ° 12.834 ’ N, 123 ° 30.043 ’ E, PHS 02 - 5 (AM C. 585908 20 + p )); Lazy, 09 ° 07.544 ’ N, 123 ° 38.407 ’ E, PHS 03 - 1 (AM C. 595952 1 p). Cebu: Mactan Point, 10 ° 20.014 ’ N, 124 ° 02.723 ’ E, PHS 04 - 2 (AM C. 585946, 20 + p, C. 585119 p [M 413, SK 096], C. 585120 p [SK 094], AM C. 585946 10 p). Bohol: Caubian Is, 10 ° 17.117 ’ N, 124 ° 10.306 ’ E, PHS 04 - 3 (AM C. 595936 p [SK 561]). Luzon: San Agapito, E side Isla Verde, 13 ° 31.782 ’ N, 121 ° 05.499 ’ E, PHV 02 - 1 (AM C. 585862 p [M 407, SK 076]), NW Polillo Is, Bolunga District, nr Panukalan, E Quezon, 14 ° 59 ’ N, 121 ° 49 ’ E (WAM S 72342, 10 + p). Japan, Okinawa: Sun Marina Beach, Onna, 26 ° 27.842 ’ N, 127 ° 48.755 ’ E, JP 01 - 1 (AM C. 585618 9 p); Cape Maeda, 26 ° 26.573 ’ N, 127 ° 46.113 ’ E, JP 01 - 2 (AM C. 585619 9 p, C. 595925 p [M 597, SK 541], C. 595927 p [M 598, SK 542], C. 595928 p [M 599, SK 543]); Onna, 26 ° 26.113 ’ N, 127 ° 46.085 ’ E, JP 01 - 3 (AM C. 585620, 2 p); Moon Bay, Onna, 26 ° 26.653 ’ N, 127 ° 48.230 ’ E, JP 01 - 4 (AM C. 585625 4 p, C. 584909 p [M 506, SK 327], C. 584910 p [M 508, SK 323]); Tancha Bay, rocky point, 26 ° 27.941 ’ N, 127 ° 49.194 ’ E, JP 01 - 6 (AM C. 584922 p [M 494, SK 313], C. 584923 p [M 497, SK 316], C. 584924 p [M 503, SK 324], C. 584925 p [M 507, SK 322]). Honshu: Boso Peninsula, Tateyama, Arai Bch, seawall, 34 ° 59.838 ’ N, 139 ° 51.378 ’ E, JP 01 - 2 (AM C. 584934 1 p [M 488, SK 307], Fig. 12 A, L). Indonesia: Riau Islands, NE coast of Pulau Panjang, 04 ° 15.9 ’ N, 108 ° 12.27 ’ E (ZRC. MOL. 24910 [M 520], Fig. 8 P), 24911 p [M 521], Fig. 8 Q), 24907 p [M 517, SK 433], 24908 p [M 518, SK 434]); Anambas, SW Pulau Mubur, Teluk Air Bandung, eastern shore of Pulau Kecik, ZRC EA-ZJ 07 (ZRC. MOL. 24909 p [M 522]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF888207FF68FCC2FA0FF7F6.taxon	discussion	Taxonomic remarks. Siphonaria sipho is the type speciesof Siphonaria bysubsequentdesignation. Thename has widely been used through the taxonomic literature, but it has been treated inconsistently in the past resulting in a confused taxonomic history. Sowerby (1823) referred to ‘ Patella sipho ’ as a ‘ common name’, which, to our knowledge, was not described previously and therefore is considered as a nomen nudum. The original type material of S. sipho is considered lost; we found no references to types in previous systematic literature and no type material was located at the NHMUK after an exhaustive search (K. Way & J. Ablett, pers. com.; Art. 75.3.4 of the Code). The neotype of S. sipho (Fig 8 A), is herein designated to clarify the taxonomic status of this taxon and its type locality in accordance with Art. 75.3 of the Code. This neotype designation aims to remove prevailing uncertainty about the identity of this species and to clarify the taxonomic status of Siphonaria. By designating the neotype, we restrict the type locality of S. sipho to Calalinan, Siquijor, Philippines. Altogether four hand-drawn figures of S. sipho were published in earlier treatments. The original description of this species was published in 1823 while the original black and white drawings were published in 1824 (Sowerby I 1824: pl. 122, fig. 1; Fig. 8 O). Subsequently, the species was depicted by H. Adams & A. Adams (1856: pl. 84, fig. 10 b) and Reeve (1856: pl. 2, figs 9 a – d) based on specimens from the Philippines. While these figures differ in some respects, they provide a limited number of shell characteristics that aid the identification of this species. Although Sowerby’s (1823) figures of S. sipho are generalized, the proportions, evenness and nature of shell ribbing, and shape / shading of the interior shown in these figures, closely match the characteristics of topotypic specimens of S. luzonica Reeve, 1856 rather than any other Indo West-Pacific species. Based on this close similarity in shell features, we treat S. luzonic a as a junior synonym of S. sipho. Figures of S. sipho in Reeve (1856: pl. 2, fig. 9 a – b, [specimen 1] from Philippines) and H. Adams & A. Adams (1856: pl. 84, fig. 10 b; no location) closely match Sowerby’s figure of S. sipho. Reeve’s second figure of S. sipho (1856: pl. 2, fig. 9 c – d [specimen 2] from Philippines), however, differs from the first specimen. It is closely similar with Reeve’s (1856: pl. 6, fig. 29 a – b) coloured figures of S. luzonica and with Reeve’s (1856: pl. 6, fig. 31 a – b) coloured figures of S. corrugata. Our delineation of S. sipho is based on comparative analyses of the morpho-anatomy and mitochondrial genetics of the neotype and freshly collected topotypes of S. sipho depressior (Figs 8 I) and a geographic series of additional specimens (Tables S 1 – 2), including topotypes of S. exigua (Figs 8 J), and specimens closely matching the types of S. luzonica and S. corrugata, respectively (Figs 8 B – D, E). Sowerby I (1823) described Siphonaria exigua by reference to the unavailable name Lepas exigua published in Martini & Chemnitz (1769: 92, figs 88 – 89) (work on the Official Index of rejected and invalid works in zoological nomenclature). The original figure of “ Lepas exigua ” is similar to the shell depicted in Sowerby I (1823: pl. 143, fig. 4) by absence of a prominent siphonal ridge and the configuration of ribs. Therefore, we treat S. exigua Sowerby I, 1823 as a junior synonym of S. sipho. Gray (1824: 276) introduced S. radiata as a substitute name for all species figured by Sowerby’s (1823) (i. e. S. sipho, S. concinna, S. exigua and S. tristensis) instead of using one of these names stating that “ he [Sowerby] has … fallen into the common error of modern conchologists, of making too many species; for I have good reason to believe that all the specimens that he has figured, except S. tristensis, …. belong to one species ”. Siphonaria radiata Gray (1824: 276) therefore is an unnecessary replacement name and permanently invalid. Gray (1847 a: 181) subsequently reverted to using the name S. sipho as the type species of the genus. Siphonaria cornuta Gould, 1846 was described for shells collected at Mangsee Islands, Sulu Sea during the ‘ United States Exploring Expedition’. The shells were figured subsequently by Gould (1856). Gould (1846 b: 11) indicated that S. cornuta was allied to S. atra. Johnson (1964: 60) listed Gould’s (1856) figured specimen as the ‘ holotype’ and mentioned additional paratypes (3 paratypes MCZ 88100 ex BSNH 3763, original no. 792; 3 paratypes MCZ 216757 ex Smithsonian Inst.). As the original description does not contain an original type designation, Johnson’s (1964: 60) subsequent reference to the holotype qualifies as a lectotype designation (Art. 74.6 of the Code). The shell sculpture and geometry of the holotype of S. cornuta (i. e., elongate shell, low profile primary ribs, strong protruding shell edge ribbing, interior shell edge colouration and interstice secondary ribbing, Fig. 12 E) closely matches diagnostic features of S. sipho. For this reason, we treat S. cornuta as a junior synonym of S. sipho. Pilsbry (1895: 5) considered S. cornuta as a synonym of S. atra. However, the holotype of S. cornuta differs from the more circular geometry and less protruding edge ribbing of S. atra. Therefore, we reject Pilsbry’s synonymization. Siphonaria siquijorensis Reeve, 1856 was described from the Island of Siquijor, Philippines. One syntype is figured in Higo, Callomon & Gotō (2001: 142, fig. G 4974, “ BMNH 1969166 ”). While topotypic specimens matching the shell form of syntypes were not found in this study, the shell morphology of specimens of S. sipho from Okinawa, Japan (Fig. 12 B – C) match the shell sculpture and geometry of syntypes of S. siquijorensis (Figs 8 K – M). For this reason, we treat S. siquijorensis as a junior synonym of S. sipho. The record of S. siquijorensis in Hedley (1909: 369, Qld) is outside of the known distribution of S. sipho; likely a misidentification of either S. atra or S. opposita. Schrenck (1867: 306) described three infraspecific taxa (elatior, depressior and intermedia), all intended to describe the extent of intraspecific variation within this species, stating ‘ former and the latter of these forms are available to us from the Philippines, the second, ... from the fauna area in question, the North Japan Sea’. The subspecies depressior was recorded from ‘ Bai von Hakodate auf Jesso’ [Hakodate Bay, Hokkaido, Japan]. Hubendick (1946: 48) treated all three taxa as junior synonyms of S. sipho. Mitochondrial and comparative anatomical data presented here reveal that specimens matching these nominal taxa are well contained within the range of intraspecific morphological variation of S. sipho. Therefore, we maintain their treatment as synonyms of S. sipho. Reeve (1856) incorrectly listed several species as synonyms of S. sipho (i. e., S. crebricostata [nomen nudum; refer below], S. albicans [sic albicante = S. atra], S. zelandica [Australia], S. punctata [Mauritius], and S. plicata [Tonga]). Schrenck (1867: 306) and Hutton (1880: 36, adding S. inculta Gould) appear to have accepted Reeve’s synonymy. The listing of S. exigua in Adcock (1893: 11) as synonym of S. luzonica (from SA; misidentified S. zelandica) is erroneous. The identity of ‘ S. sipho Sow. ’ in Vernhout (1913: 252, from Ceram) is implausible for being well outside the known distribution of S. sipho. It possibly refers to S. costellata sp. nov. described herein. The identity of S. sipho in Hutton (1880: 36, 1883: 143, from New Zealand) is unclear and most likely a misidentification of S. australis (Jenkins, 1983: 11). The synonymy given by Hutton (1880) is incorrect and includes distinct species. Therefore, the stated distribution of S. sipho (‘ Philippine Islands, Indian Archipelago, Mauritius, Tonga, Chatham Islands, Auckland to Dunedin [NZ] ’) is also incorrect. Hubendick (1946: 47) treated S. sipho and S. exigua as intraspecific taxa within S. laciniosa, most likely based on the type specimen of Patella laciniosa Linneaus, 1758: 781, held at UUZM. Examination of type specimens and morpho-anatomy in this study reveal that Hubendick’s interpretation of S. sipho is based on misidentifications of S. javanica, S. viridis and S. sipho. His synonymization of S. sipho with S. laciniosa is rejected. Kuroda (1960: 43) also incorrectly synonymized S. sipho Sowerby with S. laciniosa. He listed ‘ S. laciniosa ’ from Okinawa (unfigured). We consider this record to be identical either with S. sipho or S. tanchaensis sp. nov., both found in Okinawa and exhibiting shells very similar to S. laciniosa. Morrison (1972) treated S. sipho as a junior synonym of S. laciniosa ‘ Lamarck, 1819 ’ along with 26 other nominal species based on similarity to a ‘ laciniosa ’ shells figured throughout the literature and a ‘ common reproductive development’. These synonymies are not supported by examination of type specimens and comparative morpho-anatomy and are therefore rejected.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF888207FF68FCC2FA0FF7F6.taxon	description	External morphology (Figs 8 T – U, 12 L – M). Foot sole dark grey, foot edge pale yellow, unpigmented; foot wall, pneumostome and cephalic lobes pale greyish yellow, covered in irregular blotches of black pigmentation; mantle thin, translucent, pale yellowish grey, edge thickened, lobed, with black pigmentation edge bands aligning with underside of rib interstices. Shell (Figs 8, 12 A – F; Table S 9). Highly variable, medium sized (max sl mean = 16.8 mm, SD = 2.9 mm, n = 9), circular ovate, low to commonly tall; thick, apex offset central sightly to posterior and left, apical sides straight to convex, sometimes strongly convex (Figs 8 G – H, J); protoconch homostrophic (n = 2; Figs 8 V), shell whorl dextral; shell exterior and edge usually uneven; growth striae prominent in bands; rib count (mean = 37, SD = 5; n = 9), 10 – 12 fairly evenly spread primary ribs, pale white, raised rounded ridges, widen to shell edge, protrude beyond shell lip to unevenly scallop and corrugate the edge, ends of primary ribs may be flared creating uneven roughness on rib ridges, between primary ribs brown flecks / bands with two to six finer whitish secondary ribs (sometimes none), rib interstices narrow, darker; paired primary ribs on siphonal ridge, no more prominent than other primary ribs. Interior shell margin white to cream, dark brown markings on shell edge align under rib interstices may extend over shell margin (as in Sowerby’s original figure), white rays align on shell margin under primary / secondary ribs, ADM distinct, varies from tan to dark brown; siphonal groove distinct, same colour as shell edge; spatula golden tan with central white to dark chocolate brown; ADM scar distinct, brown to white as spatula; cephalic muscle weakly convex; thickening and whitening of shell lip common, translucent, infills and reduces lip scalloping, spatula becomes whitened. Neotype (Fig. 8 A). Shell (sl = 18.7, sw = 14.4, sh = 6.3 mm) circular ovate, medium; thick, apex offset weakly to posterior and left, ~ 14 primary ribs, with 1 – 3 in between finer secondary ribs; siphonal ridge formed by adjacent dual primary ribs. Interior shell lip and margin white under ribs, dark brown rays on shell lip aligning under rib interstices; taller and slightly darker interior shell form of S. sipho. RS (Fig. 9 A). Reproductive system (Figs 9 – 10; n = 15). Positioned to the right side of coelom, against foot wall, on foot muscle and under the respiratory cavity; epiphallic parts positioned over BM and between RAM; GA very small, with singular GP through foot wall; AO large, base wide, elongated, tip bluntly pointed and solidly embedded in MG, joined to lower ED and upper GA; ED as long as AO, narrow, centrally bent, joins to outer side of GA; GA, AO, ED all white muscular fibrous tissue; EG soft whitish tissue, slightly folded, medium in size, joins end of ED; single flagellum (F 1), short, often looped, appears as an extension of broader ED; BD and CD connect closely but in opposing directions into GA between ED join and GP, BD narrower and longer, both smooth, whitish, featureless, pass closely together through outer side of RAM (BD over CD) into soft white folded tissues of MG; MG / AG complex may be large; CD connecting to ducts; BD with large distal loop often looping behind ED, MA often present; BC small, embedded in top of AG, spherical, thin whitish translucent test; HD short, small, folded, links ducts in soft white folded tissues of AG to yellowish granulated HG; AG similar size to HG, sides match curvature of inner foot wall. Spermatophore (Figs 9 C, G, 10 B). Thread-like (mean total length = 3.63 mm, SD = 1.06 mm, n = 2), translucent, test thin; head section, tip bluntly rounded, evenly cylindrical, elongate, containing a white gelatinous mass; taper region into the filamentous transparent flagellum is short; both sections smooth, featureless. Head shorter and thicker than flagellum (mean head length = 1.46 mm, SD = 0.11 mm, n = 2; mean of SPM length ~ 41 %, SD = 9 %; mean head width = 120 μm, SD = 20 μm, n = 2; mean flagellum width = 13 μm, n = 2, SD = 15 μm). Seven SPM tightly coiled in one bursa found in [SK 078]. Radula and jaw. Dentition formula 40: 1: 40 (Hubendick 1946: 47, misidentified as S. laciniosa).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF888207FF68FCC2FA0FF7F6.taxon	diagnosis	Comparative remarks. Siphonaria sipho (laciniosa group, unit 24) is genetically well-differentiated from other species by COI distances of ≥ 14.9 % (Table S 6). The sister species of S. sipho is S. rodrigoensis sp. nov. (Fig. 3). The nominal taxa S. luzonica, S. exigua, and S. corrugata are synonymized herein based on examination of topotypic samples. Siphonaria siquijorensis, S. exigua, and S. corrugata are treated as conspecific based on examination of specimens corresponding closely to the types. Shells matching Sowerby’s original figure (Fig. 8 O), Reeve’s figures of S. sipho and specimens matching Sowerby’s original figure of S. exigua are all conspecific. Anatomically the RS and SPM of all forms reveal some variation in RS structure (i. e., AO may be very wide and blunt (Fig. 9 D); and F 1 short and (un) looped (Figs 9 B, D – F). This variation is considered as infraspecific. Siphonaria sipho exhibits a wide range of shell variation, particularly in shell thickness and height, primary and secondary ribbing as well as internal and exterior colouration. This variation appears uncorrelated with shell size or geographical distribution. Shell variability ranges from a heavily ribbed, tall shape (resembling S. exigua; Fig. 8 J) to a low, finer ribbed shape with ribs aligning closely to the shell edge (i. e., not strongly protruding; resembling S. cornuta, S. luzonica and S. corrugata shell forms). Populations on Riau Island, Indonesia, represent a geographical outlier that is morphologically somewhat distinct but genetically not well-differentiated from other representatives of this species. The Riau Island specimens differ by having a smaller, lower shell, less raised ribbing, weaker edge scalloping, a smaller AO, shorter, narrower ED, and shorter F 1. Throughout the range of S. sipho we found seven congeners occurring in partial sympatry. Four of these are sympatric in the Philippines: The shell of S. bifurcata resembles finer ribbed forms of S. sipho but is distinguished by narrower primary ribs, fewer intervening secondary ribs, a weakly flared siphonal ridge edge, and paler interior colouration as well as by having a smaller AO, GA and a shorter ED. Siphonaria sirius has a lower shell height, a siphonal ridge formed by single rib, a darker exterior and interior, broader and white primary ribs as well as a smaller AO and bursal loop. Siphonaria caubianensis sp. nov. has a lower height, darker exterior and interior, strongly and unevenly scalloped edge, broader and white primary ribs, a larger AO, bursal loop, and smaller BC. Siphonaria alba has broader primary ribs, a shorter, wider ED, and a smaller AO. Four species occur in sympatry with S. sipho in Okinawa: Siphonaria camura sp. nov. and S. rucuana are smaller in shell size, the former having a fragile, evenly brown shell; the latter a combination of less raised shell, even ribbing, and darker interior colouration. Both have a smaller AO, shorter wider unlooped BD, and S. camura sp. nov. has a barbed SPM. Siphonaria subatra has a darker exterior and interior colouration, narrower ribbing, lower shell, a smaller BC, a slightly smaller AO. Siphonaria tanchaensis sp. nov. closely resembles S. sipho anatomically and morphologically; it has a similar shell geometry, colouration and ribbing, and internal colouration (i. e., spatula, shell lip including a thickened white lip form). However, has a taller shell, narrower ribs, and a paler, less banded inner shell lip, a smaller BC, ED and AO, and a shorter SPM. Siphonaria sipho co-occurs with S. japonica in Honshu, and has a lower, thinner, and more fragile shell, dark brown exterior and interior colouration, narrower ribbing, a smaller AO, larger BC, and a barbed SPM. Siphonaria radians, sympatric on Riau Islands, has a lower shell with a more posteriorly offset apex and a multi-ribbed siphonal ridge without a flared edge, a larger AO, longer wider twisted ED and a smaller BC. Specimens identified as S. sipho from Java, Bali, and PNG by Adam & Leloup (1939) are here attributed to S. viridis. The figure of S. sipho in Hirase (1941: 94, pl. 121, fig. 11) is probably a specimen of S. tanchaensis sp. nov. The RS figured in Hubendick (1945: 31, fig. 48) matches the RS of S. sipho. The RS figure in Hubendick (1945: 31, fig. 52) attributed to ‘ S. kurracheensis ’ differs from RS of S. kurracheensis and is probably of S. sipho. Specimens from Palawan figured as ‘ Siphonaria javanica ’ in Springsteen & Leobrera (1986: pl. 81, figs 21 – 22) are specimens of S. sipho. The record of ‘ S. sipho var. exigua ’ in Subba Rao & Dey (2000: 190) from Andaman and Nicobar Islands is based on a misidentification. Figured specimens of ‘ S. sirius ’ in Poppe (2010: pl. 913, figs 3 – 4) are here attributed to S. sipho. Instead, S. sirius typically possesses a single rib forming the siphonal ridge. Specimens figured as ‘ unit 25, laciniosa group’ in Dayrat e t al. (2014: fig. 4 F, H, I) from Okinawa, Taiwan, and the Philippines are all consistent with characters typical of S. sipho.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF888207FF68FCC2FA0FF7F6.taxon	distribution	Distribution and habitat. Philippines (Bohol, Balicasag, Bandian, Negros, Olango, Siquijor, Mactan, Isla Verde, Cebu, Mangsee Is), Taiwan, Japan (Okinawa, Boso Peninsula, Honshu), Indonesia (Riau and Anambas Islands) (Fig. 11). Commonly found in sheltered locations, such as hollows and crevices, across upper and mid littoral levels, on bare, exposed rocks. Home scars prominent.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF818202FF68FF02FE07FA56.taxon	description	(Figs 12 G – H, K, N – O, 15 A)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF818202FF68FF02FE07FA56.taxon	materials_examined	Material examined. Type material. Lectotype of Patella javanica Lamarck, 1819, present designation, from ‘ les côtes de Java’ [coasts of Java, Indonesia]) (MHNG MOLL- 50923, syntype number ‘ 2 ’, Fig. 12 G). Paralectotypes, same data as lectotype (MHNG MOLL- 50923, syntype number ‘ 1 ’, d, Fig. 12 H; syntype number ‘ 3 ’, d, Fig. 12 I). Other, non-type material. Timor-Leste: N of Dili: Dolokoan Beach, 8 ° 31.424 ’ S, 125 ° 37.091 ’ E, TL 01 - 1 (AM C. 584795 p [M 434, SK 145]; Fig. 12 J).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF818202FF68FF02FE07FA56.taxon	discussion	Taxonomic remarks. The type series for Patella javanica Lamarck, 1819 comprises three syntypes labelled 1 to 3. Syntypes # 1 and # 2 are siphonariids (Figs 12 G – H); however, syntype # 3 is a liotiid (Fig. 12 I). In the original description Lamarck (1819) stated that the shell had ‘ small white sides … fine and longitudinal streaks between its ribs’ while the interior was ‘ blackish, bordered with yellow and a white edging’. This description closely matches syntype 2 (Fig. 12 G), which is herein designated as the lectotype of Patella javanica for stabilisation of the name (Art. 74.1 of the Code). The figure of Patella javanica in Delessert (1841: pl. 23, fig. 3 a – c), matches syntype # 1 (Fig 12 H). Hubendick (1946: 38) treated S. javanica Lamarck, 1819 as variety of S. laciniosa. This treatment is rejected herein. Specimens figured in Hubendick (1946: 47) as S. laciniosa var. sipho (pl. 3, fig. 19, ‘ Java’) and S. stellata (pl. 3, fig. 23, ‘ Sunda Islands’), respectively, are of S. javanica. The ‘ very minute barbs’ on the SPM mentioned by Hubendick (1946: 48) have not been observed. Such barbs on SPM have been observed only in S. japonica, S. camura sp. nov. The RS figured as S. kurracheensis siquiorensis (sic!) in Hubendick (1945: 31, fig 52; from ‘ Edam Java’) matches the RS of S. alba as described herein, not S. kurracheensis nor S. javanica (Fig. 15 A). No type specimen of P. javanica was figured in Memod (1950). Morrison (1972: 51) in redefining selected species groups, incorrectly considered S. sipho as a synonym of S. javanica and incorrectly stated this taxon to be the type species of the genus. Further, Morrison (1972: 52) incorrectly treated S. exigua and S. sipho as synonyms of S. javanica based on similarity of the shell and a ‘ common reproductive development’. These synonymies are not supported by examination of type specimens and comparative morpho-anatomy. The figured specimen in Morrison (1972: 55, fig. 1, ‘ North Borneo’) matches the lectotype (Fig. 12 G). Reversing taxonomic precedence, Rehder (1980: 97) incorrectly stated ’ Patella javanica Lamarck, 1819 ’ (= S. javanica) to be a synonym of S. sipho. Records of ‘ S. javanica ’ in Tan & Chou (2000: fig. 116) and Tan & Woo (2010: 61) are misidentifications of S. viridis (refer Comparative remarks, S. viridis). The record of ‘ S. javanica’ in Tan & Kastro (2004: 50) from Anambas and Natuna Islands, Indonesia is incorrect and attributable to S. sipho.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF818202FF68FF02FE07FA56.taxon	description	External morphology (Figs 12 N). Foot sole centrally pale grey to dark yellow at foot edge, unpigmented; foot wall and cephalic folds dark yellow with irregular black flecks; mantle translucent dark grey to join of foot wall, yellow markings aligning with underside of ribs, black bands between on edge align under rib interstices, edge lobed and thickened to shell lip. Shell (Fig. 12 G – I, K; Table S 9). Medium sized (max sl mean = 19.3 mm, SD = 9.7 mm, n = 2), ovate; height tall; apex offset central, apical sides convex, posterior straight to concave, protoconch area dark brown; protoconch direction central to weakly heterostrophic (n = 1), shell whorl dextral; growth striae prominent, shell thick; rib count (mean = 54, SD = 1, n = 2), ~ 11 primary ribs pale white, fairly straight, rounded ridges, increasingly raised / broaden and protrude beyond shell lip (~ 1 mm) to unevenly scallop and corrugate the edge; 3 or more interspersed finer secondary ribs pale white with irregular black / brown flecks, rib interstices white, uneven axial brown streaks; paired fused primary ribs form siphonal ridge. Interior shell margin intensely white with fine dark brown rays on edge aligning under rib interstices, siphonal groove distinct; spatula dark chocolate brown; ADM scar indistinct and paler brown with some darker rays to shell margin, CMS convex, paler than shell lip; thickening of shell lip not observed. Reproductive system (Fig. 15 A; n = 1). Positioned to right side of coelom, against foot wall on foot muscle, under respiratory cavity; epiphallic parts positioned between RAM and BM; GA small, with singular GP through foot wall; AO large, very wide, elongated, bluntly pointed, joined to lower ED and upper GA; ED relatively long, wide, unfolded, joins to outer side of GA; GA, AO, ED all white muscular fibrous tissue; EG medium, joins end of much longer ED, soft whitish tissue, slightly folded; single, coiled end flagellum (F 1), appears as an extension of wider ED, lays over back of BM; BD and CD junctions into GA close between ED join and GP, both ducts narrow, very long, smooth, whitish, featureless, pass closely together through outer side of RAM (BD over CD) into soft white folded tissues of MG; MG / AG complex large; CD connecting to ducts; BD with large distal loop without MA, BC very small, embedded in top of AG, spherical, thin whitish translucent test; HD short, small, folded, links ducts in soft white folded tissues of AG to yellowish granulated HG; AG similar size to HG, sides match curvature of inner foot wall.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF818202FF68FF02FE07FA56.taxon	diagnosis	Comparative remarks. In the molecular phylogeny (Figs 1, 3), S. javanica (laciniosa group, unit 65) is represented by a single individual only. The species is genetically well-differentiated from other species by COI distances of ≥ 27.6 % (Table S 6). Throughout the range of S. javanica, we found eight congeners occurring in partial sympatry. Siphonaria viridis (sympatric in northern PNG and Timor-Leste) has a smaller shell with less raised ribs and less scalloped edge, a shorter and broader ED and slightly larger BC. Siphonaria atra (sympatric in northern PNG) has a lower, darker, unpatterned shell with less raised darker and narrower ribs and a shorter ED. Siphonaria normalis has a smaller shell with less raised ribs, a less to unscalloped edge, and a shorter ED and larger BC. Siphonaria opposita (sympatric on Timor-Leste) has a lower, paler, unpatterned shell with narrower ribs, less scalloped edge, flared siphonal ridge, a shorter ED, a more pointed AO, and a bursal loop on BD. Siphonaria alba has a lower darker, unpatterned shell, a twisted ED, and a larger BC. Siphonaria forticosta sp. nov. has a lower, darker shell with less raised ribs, weaker edge scalloping, a smaller AO and ED, a larger BC, and a longer F 1. Siphonaria planucosta sp. nov. has a smaller, lower, darker shell with less raised and even ribbing, a less scalloped edge, a shorter ED and AO, and a larger BC. Siphonaria campestra sp. nov. has a lower shell with less raised ribs, less scalloped edge, darker interior, and shorter ED, F 1, and AO. Siphonaria javanica is conchologically resembles several species of the laciniosa group but differs anatomically (i. e., RS structure mainly size of BC; epiphallus parts of ED, F 1). Hubendick (1945: fig. 53) figured the distal parts of the reproductive system of what he considered to be S. kurracheensis var. siquiorjensis from Java. However, this figure matches closely the anatomy of S. javanica (Fig. 15 A) and therefore we consider this treatment as a misidentification. Shells from Indonesia depicted as S. javanica by Dharma (1992: pl. 17, fig. 1) closely resemble S. sipho, while those depicted by Dharma (2005: pl. 79, figs 19 a – b) from East Java and Bali are correctly attributed to S. javanica. Specimens from Palawan identified as ‘ S. javanica ’ by Springsteen & Leobrera (1986: pl. 81, figs 21 – 22) are misidentified specimens of S. sipho. A specimen from Singapore figured by Tan & Chou (2000: fig. 116) as ‘ S. javanica ’ closely resembles S. viridis with respect to shell colouration, number and prominence of primary ribs, interior margin, size rather than S. javanica. There are currently no confirmed records of S. javanica from Singapore. A record of ‘ Siphonaria javanica ’ from Hainan (Hasegawa et al. 2001: 28) is likely a misidentification from outside of the known distribution of this species (Fig. 11). The correct identity of these specimens is unknown.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF818202FF68FF02FE07FA56.taxon	distribution	Distribution and habitat. Recorded from throughout Indonesia and Timor-Leste (Fig. 11). In this study collected on exposed to moderately exposed rocky boulder shores at upper littoral levels.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF83823DFF68FA62FF44F956.taxon	description	(Figs 13 A – L, Q – S, 15 B – E)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF83823DFF68FA62FF44F956.taxon	materials_examined	Material examined. Type material. Neotype of Patella japonica Donovan, 1824, present designation, from Headland S of Chitose Bch, Boso Peninsula, Japan, 34 ° 59.240 ’ N, 139 ° 58.304 ’ E; coll. B. W. Jenkins, 15 March 2020, JP 02 - 2 (AM C. 584938 [M 489, SK 308]; Figs 13 A, 15 D – E). Syntypes of Siphonaria cochleariformis Reeve, 1856 (NHMUK 1981015, Fig. 13 H – K). Syntypes of Siphonaria alterniplicata Grabau & King, 1928 (MBM 280642, 72 shells, Peitaino, July 1925; Fig. 13 F). Holotype of Siphonaria (Kerguelenella) corallina Christiaens, 1980 from Channel Rock, Hong Kong, China; on corals at 10 m depth (NHMUK 1977170, Fig. 13 G). Other, non-type material. Japan, Honshu: Boso Peninsula, Point S of Chitose Beach, 34 ° 59.240 ’ N, 139 ° 58.304 ’ E, JP 02 - 2 (AM C. 585394 9 p, AM C. 585113 p [SK 423], AM C. 585934 p [SK 425], AM C. 585935 p [SK 339], AM C. 584983 p [SK 338], AM C. 595953 p [SK 566]); seawall Arai Bch, Tateyama 34 ° 59.838 ’ N, 139 ° 51.378 ’ E, JP 02 - 1 (AM C. 585389 10 p, AM C. 585235 p [SK 340]). China: Hong Kong, Cape D’Aguilar 22 ° 12.28 ’ N, 114 ° 15.38 ’ E (ZRC. MOL. 2001 - 1768 21 p, ZRC. MOL. 24905 p [M 476, SK 283], ZRC. MOL. 24903 p [SK 403], ZRC. MOL. 24904 p [SK 404], ZRC. MOL. 24906 p [SK 405]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF83823DFF68FA62FF44F956.taxon	discussion	Taxonomic remarks. Type species of Sacculosiphonaria Hubendick, 1945 by original designation. This name became first available with publication of the original plate 79 (June 1, 1824) not the bound volume (3, 1825). Plates and texts were ‘ published monthly, bound yearly’ (Molluscabase Eds. 2022). The original description does not contain an original type designation. The original type material is considered lost as it could not be found at the NHMUK in 2022 (J. Ablett, pers. comm.). The neotype is designated herein to clarify the identity of this nominal species and to designate the type locality (Art. 75.3 of the Code). Our delineation of this species is based on comparative analyses of the morpho-anatomy and mitochondrial genetics of freshly collected topotypes (Figs 13 B – D) and geographic series of additional specimens (Tables S 1 – 2). Pilsbry (1920 a: 141) treated S. cochleariformis Reeve, 1856 as a junior synonym of S. japonica (Donovan, 1824). This was followed by Hubendick (1946: 43), Kuroda & Habe (1952), Kuroda (1960), and Dayrat et al. (2014). We maintain this synonymy but note that it requires corroboration through examination of topotypic material, which is currently unavailable. The types of S. cochleariformis (NHMUK 1981015, Figs 13 H – K) closely resemble the figures in the original description of Reeve (1856). These shells also match the original description and figures of P. japonica in Donovan (1824). Moreover, Reeve (1856) appeared to be unaware of the description of S. japonica. Hedley (1915: 752) correctly stated that the record of S. cochleariformis from NSW (Angas, 1867: 232) is by mistake. The type locality of S. alterniplicata can be inferred from the title of the work to be ‘ Peitaiho’ [China, Beidaihe District, 39 ° 50 ′ N, 119 ° 29 ′ E] (Grabau & King, 1928: 62) ’. The syntype of S. alterniplicata figured in Coan et al. (2015: 221, fig. 38). The proposed synonymy of S. alterniplicata (Fig. 13 F) remains to be tested by examination of topotypes. Siphonaria (Kerguelenella) corallina wasdescribedby Christiaens (1980 a: 80) based on five specimens collected from corals and barnacles at 10 m depth off Channel Rock, Hong Kong, with the largest specimen lodged as holotype in NHMUK (Fig. 13 G). Christiaens figured the shell and halfradularowfortheholotype (Christiaens, 1980 a: 80, fig. 18 A, C) indicating the radula dentition formula of 19.1.19. No other material is recorded. Unfortunately, no animals are available to confirm anatomical (mainly radula) or assess molecular characteristics. However, examination of the holotype indicates the specimen is a juvenile. It is brownish transparent and has wide and glassy growth lines on the shell consistent with early growth stages in siphonariids. The holotype can be aligned well with the early growth stages of adult specimens of S. japonica (Fig. 13 N), revealing that shell profile, apex, prominent ribs, and formation of dual siphonal ridge ribs match. By contrast, the holotype cannot be aligned with early growth stages in shells of other co-occurring species, such as S. alba (Fig. 13 M), S. subatra (Fig. 13 O) and S. rucuana (Fig. 13 P). While molecular sequences are required for validation, we postulate that the holotype of S. corallina is a juvenile of S. japonica. Regarding collected depth, it is not uncommon for Siphonaria to occur subtidally (e. g., S. lateralis has been recorded from ‘ bottom of a 6 m deep tide pool’ at the Snares Islands (AM C. 586004; coll. D. Horning) and at ‘ 3 to 4.5 m’ at Aerial Cove, MI, Australia (coll. J. Lowry). However, most specimens have been recorded intertidally.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF83823DFF68FA62FF44F956.taxon	description	External morphology (Fig. 13 Q). Foot sole, foot wall, mantle, cephalic folds and pneumostomal lobe all evenly orange in colour; mantle thin, narrower than foot wall, edge thickened lobed with black bands aligning with rib interstices; dark irregular black blotches of pigmentation on foot wall and cephalic lobes, fading to join with mantle, concentrated over centre of cephalic folds; pneumostome long between right adductor muscles and within mantle. Shell (Figs 13 A – L; Table S 9). Medium sized (max sl mean = 19.4 mm, SD = 1.7 mm, n = 9), circular ovate, height medium, shell thin; apex offset strongly posterior and weakly left, apical sides strongly convex, protoconch direction homostrophic (n = 1), shell whorl dextral; growth striae prominent uneven, radial colour banding faint, exterior pale brown; rib count (mean = 46, SD = 6, n = 5), wavy and bent, pale red brown / grey, ridges narrow, rounded, golden tan, variably raised, uneven, slightly broaden to align with unscalloped shell edge; ~ 20 primary ribs, 2 – 3 finer secondary ribs between primary ribs, interstices narrow with regular red brown and irregular dark grey / white markings; paired primary ribs form siphonal ridge, protrudes slightly beyond shell edge. Interior: shell lip with white rays aligning under primary / secondary ribs, fade over pale tan shell margin; ADM scar and spatula evenly dark brown, siphonal groove distinct; ADM not prominent, CMS straight; thickening / whitening of shell lip not apparent. The neotype (Fig. 13 A). Shell (sl = 20.5, sw = 16.8, sh = 7.5 mm) elongate ovate, medium; thin, apex offset weakly to posterior and left, exterior uneven, mottled brown with radial banding, interstices with dark grey and reddish flecks, ~ 42 ribs, siphonal ridge clear, formed by adjacent dual ribs. Interior shell lip dark brown strongly corrugated with pale rays aligning under ribs, margin whitish tan, RS (Fig. 15 D) and SPM (Fig. 15 E). Reproductive system (Figs 15 B, D; n = 4). Positioned to the right side of coelom, against foot wall on foot muscle, under the respiratory cavity; epiphallic parts positioned between RAM and BM; GA small, with singular GP through foot wall; AO small, elongated, narrow, centrally bent, bluntly pointed, joined to lower ED and upper GA; ED long, wide, centrally coiled / folded, joins to outer side of GA; GA, AO, ED all white muscular fibrous tissue; EG large elongated, slightly shorter than ED, soft whitish tissue, slightly folded, joins ED; single, long, narrow, coiled flagellum (F 1), appears as an extension of wider ED, lays over back of BM; BD and CD junctions into GA close between ED join and GP but in opposing directions, both ducts narrow, long, smooth, whitish, featureless, pass closely together through RAM (BD over CD) into soft white folded tissues of MG; MG / AG complex medium; CD connecting to AG / MG ducts, BD with folded distal loop and MA, BC very large, embedded in top of MG against AG, elongated spherical, thin whitish translucent test; HD short, large, coiled, links ducts in soft white folded tissues of AG to yellowish granulated HG; AG smaller than HG, sides match curvature of inner foot wall. Spermatophore (Figs 15 C, E). Cylindrical, thread-like (length = 13.9 ± 1.5 mm, n = 3), test thin, translucent; head tip tapered bluntly rounded, section containing a white gelatinous core, tapers to a thin flagellum and tip; both sections smooth, 8 – 12 short barbs on upper flagellum pointing towards head (n = 3); head section smooth, longer thicker than flagellum (head length = 8.3 ± 0.2 mm, n = 3; 60 % of SPM length, SD = 6 %; flagellum length = 5 ± 1.5; head section width = 195 ± 12 μm; flagellum width = 33 ± 12 μm), 10 SPM tightly coiled in white gelatinous mass in BC of one specimen (ZRC. MOL. 24905 [M 476, SK 283]). Radula and jaw. Dentition formula “ varies at any rate between 32: 1: 32 and 40: 1: 40 ” (Hubendick 1946: 43).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF83823DFF68FA62FF44F956.taxon	diagnosis	Comparative remarks. Siphonaria japonica (normalis group, unit 2) forms a distinct mitochondrial clade together with its sister species, S. camura sp. nov. (Figs 1, 4). The minimum distance between S. japonica and S. camura n. sp. is 11.1 % in COI (Table S 8). Throughout the range of S. japonica we found four congeners occurring in partial sympatry. Three are sympatric with S. japonica on Honshu: Siphonaria acmaeoides has a more solid, whitish, smoother shell with a more central apex, a smaller AO and BC, and a shorter F 1 and short drop-like SPM. Siphonaria sirius has more prominent and raised ribbing with stronger edge scalloping, a larger AO, and SPM without barbs. For comparison with S. sipho refer to that species. Siphonaria camura sp. nov. has a smaller, paler shell with less raised ribbing, strongly offset and hooked apex with weaker edge scalloping, smaller AO and F 1, and a larger BC. Shell, RS, and SPM of ‘ S. cochleariformis’ depicted by Hubendick (1945: 26, fig. 34 – 36, 28, fig. 43; 1946: 14, fig. 20) from Japan, subsequently reproduced by Berry (1977: 210, fig. 19), correspond well with S. japonica (both taxa are synonyms). However, we found that type specimens of S. cochleariformis (Fig. 13 H – K) differ from typical S. japonica in being more solid, whitish, unpatterned, and having a thickened margin and raised ribs. However, no topotypic material of S. cochleariformis is available to study and we cannot address its taxonomic status. Figures of the SPM of ‘ S. kurracheensis ’ in Hubendick (1945: 31, fig. 51) are attributed to S. japonica; the number of barbs corresponds well with features typical of this species whereas flagellum barbs are absent in S. kurracheensis. Figured specimens of S. japonica in Yoo (1976: pl. 19, figs 1, 4) from Korea are misidentifications; fig. 2 and possibly fig. 3 are correctly identified. A record of ‘ S. japonica ’ from Hainan Island (Hasegawa et al. 2001: 29) is plausible but requires confirmation by dissection and / or DNA sequencing.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF83823DFF68FA62FF44F956.taxon	distribution	Distribution and habitat. Recorded from Hong Kong, Honshu, Taiwan (Fig. 11). In this study, commonly found on exposed rocky shores at various locations on Honshu, in crevices and hollows across the upper littoral level.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFBC8239FF68F962FD42FCB6.taxon	description	(Figs 14 A – D, N, Q – R, 15 F – H)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFBC8239FF68F962FD42FCB6.taxon	materials_examined	Materialexamined. Typematerial. Neotypeof Siphonaria obliquata Sowerby I, 1825, present designation, from near quarry, NZ, Dunedin, W side Blackhead, 41 ° 30.34 ′ S, 171 ° 56.76 ′ E; coll. K. Walton, 6 Dec. 2020 (NMNZ M. 331450 [M 515, SK 421], Fig. 14 A). Five syntypes of S. scutellum Deshayes, 1841 from Chatham Islands (MNHN IM- 2000 - 5117, Fig. 14 C). Other, non-type material. NZ, South Island: same data as neotype (NMNZ M. 331114 p); Gentle Annie Point, N of Mokihinui River Mouth, West Coast 45 ° 55.78 ′ S, 170 ° 25.72 ′ E KW 20 - 103 (NMNZ M. 331115 6 p, p [M 516, SK 422]; Fig. 14 B); between Buttler and Gray Rivers, W Coast, 41 ° 43.8 ’ S, 171 ° 29.31 ’ E, NZS 01 (AM C. 585927 p [SK 394], C. 585928 p [SK 401]). Point Elizabeth, 42 ° 24 ’ S, 171 ° 13 ’ E (AM C 265378) (Fig. 14 D).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFBC8239FF68F962FD42FCB6.taxon	discussion	Taxonomic remarks. Siphonaria obliquata is the type species of Benhamina Finlay, 1926 by original designation. The original type material is considered lost as it could not be found at the NHMUK in 2020 (J. Ablett, pers. comm.). The neotype is designated herein to ensure an unambiguous identification of this species, to stabilize its nomenclature, and to designate a correct type locality (Art. 75.3 and 76.1 of the Code). Sowerby (1825: Appendix 7), originally stated the origin of this species to be “ Van Diemen’s Land [Tasmania] ”. However, this probably incorrect since Siphonaria obliquata has never been found in Tasmania or elsewhere in Australia. The type locality is corrected to South Island, NZ (see above for details). Our delineation of this species is based on comparative analyses of the morpho-anatomy and mitochondrial genetics of freshly collected topotypes of S. obliquata and of S. sculellum and geographic series of additional specimens (Tables S 1 – 2). Reeve (1856) correctly stated that S. obliquata occurs in NZ and that S. scutellum is its junior synonym. Stearns (1894: 405, 430) reversed the priority of both species when treating them as synonyms referencing “ Carpenter’s Reeve-Cuming List’ (Stearns (1894: 405). He also incorrectly states the distribution to be Galapagos. This was subsequently followed by Dall (1870: 39), who incorrectly added occurrence on the “ W coast [of S America] N of Panama ”. Hubendick (1946: 24) correctly treated S. scutellum as a synonym of S. obliquata, but incorrectly listed it as a synonym of S. australis (Hubendick 1946: 49). Burch (1945: 16) listed ‘ S. scutella Deshayes’ as a valid species from Galapagos, which is rejected herein.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFBC8239FF68F962FD42FCB6.taxon	description	External morphology (Fig. 14 N). Animal extends beyond shell coverage; foot sole swollen smooth, foot wall, mantle, cephalic folds and pneumostomal lobe evenly pale grey / cream, paler at edge foot / wall; blotches of black pigmentation on centre of cephalic folds, faintly on foot wall; mantle narrower than width of foot wall, non-translucent, covers exposed inner shell lip, edge thickened, lobed, vertical bands of black pigmentation aligned with shell rib interstices; single genital pore distinct, located on foot wall to right anterior of right cephalic fold; two small black epithelial eye spots centralised on two centrally touching cephalic folds; pneumostomal lobe long under the mantle between the right ADMs. Shell (Fig. 14 A – D, R; Table S 9). Large sized (max sl mean = 48.1 mm, SD = 8.9 mm, n = 24), elongate ovate; height low to medium, often with lateral profile of shell edge arched profile; apex offset weakly posterior and central (usually eroded), apical sides anteriorly strongly convex, posteriorly straight to concave; protoconch direction heterostrophic (n = 2; Fig. 14 R), shell whorl dextral; exterior uneven, growth striae prominent, concentric brown tonal bands prominent, shell thick (often> 2 mm); apical rib count (mean = 57, SD = 10.2, n = 24), ribs pale white, ridges rounded, interstices darker brown; 20 – 25 primary ribs, adapical, slightly more prominent than interspersed finer secondary ribs, often bifurcate, curved, increasingly raised and broaden to shell lip, shell edge weakly corrugated; siphonal ridge often indistinct, formed by paired primary ribs, wide at shell edge. Interior smooth, shell lip brown to black with white rays corresponding under ribs, margin white to tan, ADM scar paler, siphonal groove apparent, shallow, same colour as shell edge; spatula tan to fawn, often with irregular brown to grey markings; ADM scar distinct, CMS straight to convex, paler than shell lip; thickening of shell lip translucent, infills and reduces lip scalloping, spatula becomes whitened. Shell lip tends to thicken in adult specimens. The neotype (Fig. 14 A). Shell (sl = 35.5, sw = 22.6, sh = 11.2 mm) elongate ovate, medium; thick, apex offset weakly to posterior and left, exterior uneven, mottled brown with radial banding, ~ 45 ribs, siphonal ridge not prominent, formed by non-adjacent dual ribs. Interior shell lip dark brown weakly corrugated with pale rays aligning under ribs, margin whitish tan, RS (Fig. 15 F) and SPM (Fig. 15 G). Neotype specimen [M 515] grouped within unit 96 (S. obliquata). Reproductive system (Figs 15 F – H; n = 3). RS positioned within right side of coelom under the respiratory cavity, hermaphroditic glands positioned to posterior against right foot wall and over foot sole, epiphallic parts positioned between sides of BM and RAM; AO, GA and ED merged, very large rounded, tapered to ED; ED wide, short, bent at join to distinct EG, large rounded fold, with orange striated prostate, curved continuation at top of ED, flagellum (F 1) indistinct, twisted; epiphallic parts all muscular tissue; CD and BD both wide, short, featureless, jointly connect into AO on under side, BD over CD (in RS of neotype, BD appears absent but is enclosed within side tissue of CD (Fig. 15 F), pass between outer side of RAM and inner foot wall; BD significantly wider, slightly curved; CD twisted immediately prior to entering MG / AG complex close to BD; BC large rounded, positioned between footwall and AG, test thick, opaque, filled with orange-brown gelatinous mass; curved elongated SV embedded in AG close to top of BC; single large GP with lobes at end of GA; HD prominent, brownish, folded and heavily lobed, links AG to yellow-tan, finely granulated HG; MG and larger AG small folded soft white tissue, MG at anterior of AG, sides moulded to curvature of inner foot wall at right posterior quarter of coelom. Spermatophore (Fig. 15 G). Short elongate, test thin, featureless, translucent (length = 4.38 mm, n = 1), head bluntly pointed; flagellum short; both sections smooth, featureless, short clear webbing spans the bend at taper between head and flagellum; head much larger than flagellum (head length = 3.0 mm, n = 1, ~ 76 % length of SPM; head width = 790 μm; flagellum width = 96 μm); three SPM found in neotype BC ([M 515]), 2 appear to be the white core remnants of larger SPM’s decomposing; while the shape / size is consistent, no test or flagellum present. Radula and jaw. Dentition formula 68: 1: 68 (258 rows) and description of jaw in Hutton (1883: 141, as ‘ scutellum ’); see Cottrel (1911: 586, fig. 6) for a description, dentition formula “ about 64: 1: 64 ” (Hubendick 1946: 25).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFBC8239FF68F962FD42FCB6.taxon	diagnosis	Comparative remarks. Siphonaria obliquata (lateralis group, unit 96) represents the sister group of a clade formed by three species, S. tasmanica, S. lessonii, and S. funiculata (Figs 1, 4). From these species it differs by 16 S distances of ≥ 7.6 % (S. funiculata), ≥ 7.9 % (S. lessonii), and ≥ 8.9 % (S. tasmanica) (no COI sequences available). Within the distributional range of S. obliquata, we found three sympatric congeners: Siphonaria australis and S. propria are sympatric on the South Island of NZ, and have smaller, more circular ovate shells, with more scalloped shell edges, prominent raised ribbing, wider HDs, smaller ephiphallic parts and BCs, and threadlike SPM. Siphonaria lateralis is sympatric on Auckland Island, NZ, and differs by having a much smaller, evenly brown, and fragile shell, a smaller BC, a larger, wider BD, and more bulbous SPM. Anatomically, the structure of RS resembles that of S. lateralis and S. tasmanica, S. funiculata, S. lessonii (Güller et al. 2015: 85, fig. 4) and S. fuegiensis (Güller et al. 2015: 92, fig. 9), all of which are relatively closely related. Hutton (1883: 141, pl. 17, figs B – D) and Cottrel (1911: 587 – 590, figs 2 – 4, 6 – 7, pl. 29, fig. 3) described the anatomy of S. obliquata; radula and jaw, RS. The RS depicted herein corresponds well with the RS depicted in Hutton (1883: 141), Cottrell (1911: 590), and Hubendick (1945: 18, fig. 14), but also reveals some intraspecific variation (e. g., details of epiphallic parts, of HD and CD in Cottrel and width of CD = “ spov ” in latter). The SPM shown herein (Fig. 15 G) corresponds well with that shown by Hubendick (1945: 14, fig. 11), reproduced by Berry (1977: 210, fig. 19). The egg mass has been described by Mestayer (1920: 171). Borland (1950: 385) described the distribution, behaviour and ecology of S. obliquata.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFBC8239FF68F962FD42FCB6.taxon	distribution	Distribution and habitat. Recorded from North and South Islands as well as Chatham, Stewart and Snares Islands, NZ, between latitudes of 34 ° S and 48 ° S (Fig. 16). In this study found to be common on exposed rocky intertidal shores in sheltered positions, such as crevices, vertical faces, mostly shaded from the midday and afternoon sun, across upper and mid littoral levels.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFB8823BFF68FA42FB3CFD96.taxon	description	(Figs 17 A – B, U – V, 18 A – B)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFB8823BFF68FA42FB3CFD96.taxon	materials_examined	Material examined. Type material. Neotype of Pileopsis radiata, present designation, Malaysia, Port Dickson, 02 ° 31.234 ’ N, 101 ° 48.026 ’ E; coll. B. W. Jenkins, 15 Feb 2015, PD 01 - 1 (AM C. 585861, Fig. 17 A). Other, non-type material. Malaysia: Bak Bak Beach, Kudat, Sabah, 07 ° 00 ′ N, 116 ° 46 ′ E (AM C. 585236 20 p, C. 595941 d [SK 523]); Bukit Keluang, Terengganu, 5 ° 47.81 ’ N, 102 ° 36.43 ’ E (ZRC 1999 - 1978 12 p; ZRC. MOL. 24900 p [M 528, SK 444], ZRC. MOL. 24901 p [M 596, SK 528]); Port Dickson, 02 ° 31.234 ’ N, 101 ° 48.026 ’ E PD 01 - 1 (AM C. 585959 3 p, C. 585821 7 d, C. 585921 p [SK 337 protoconch H 2]); Air Papan, Mersing, Johor, 2 ° 30.98 ′ N 103 ° 50.1 ’ E (ZRC 1999 - 1755 20 p, ZRC. MOL. 24895 p [M 527]; ZRC. MOL. 24896 p [M 526]); Tg. Bidara, Malacca, 02 ° 17.568 ’ N, 102 ° 5.207 ’ E (ZRC 1999 - 1746 9 p; ZRC. MOL. 24891 p [SK 348], Fig. 17 B; ZRC. MOL. 24892 p [M 525], ZRC. MOL. 24893 p [M 595], ZRC. MOL. 24894 p [M 594]); Pulau Langkawi, 6 ° 18 ′ N, 99 ° 52 ′ E (AM C. 595974 20 + p, C. 585098 p [SK 525], C. 585697 p [SK 524]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFB8823BFF68FA42FB3CFD96.taxon	discussion	Taxonomic remarks. Blainville (1826: 462) introduced the new name Pileopsis radiata, and subsequently transferred it to Siphonaria in Blainville (1827 a: pl. 2, figs 4,4 a; 1827 b: 294). The original type material is considered lost; no types were found in the MNHN (Philippe Bouchet, pers. comm.). The neotype is designated herein in accordance with Art. 75.3 of the Code to clarify the identity of this nominal taxon as well as to restrict its type locality. Blainville’s name is not invalidated by S. radiata Gray, 1824, which is not an available name (for details see under S. sipho). Siphonaria radiata (Blainville, 1826) is a senior secondary homonym of S. radiata Sowerby I, 1835 (not reviewed herein) and S. radiata A. Adams & Reeve, 1850, which has been replaced by S. radians H. Adams & A. Adams, 1855. Hubendick (1946: 49) incorrectly considered S. radiata (Blainville, 1827) a synonym of S. australis.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFB8823BFF68FA42FB3CFD96.taxon	description	External morphology. Foot sole grey, paler to foot edge; foot wall bluish to inner, evenly dark cream, with irregular black pigmented blotches, paler to foot, pustules prominent; cephalic folds thick, narrow, irregular black pigmentation darker over centre of cephalic folds; mantle wide thin translucent edge lobed thickened band; eye spots prominent, pneumostome small, under mantle. Shell (Figs 17 A – B, V; Table S 9). Small sized (max sl mean = 13.29 mm, SD = 1.55 mm, n = 42), circular ovate, height medium to tall; apex slightly curved, apex offset slightly to posterior and left, apical sides convex, posterior strongly concave, protoconch direction weakly heterostrophic to central (n = 1; Fig. 17 V), shell whorl dextral, shell thick; growth striae prominent, even, unraised; radial colour bands indistinct; rib count (mean = 30, SD = 3.1, n = 42), primary ribs distinct from secondary ribs; ~ 16 pale brown to off white primary ribs, ridges raised, rounded; siphonal ridge formed by paired adjacent primary ribs, most primary ribs broaden weakly, project beyond shell lip (especially siphonal ridge) with ends slightly raised to scallop and corrugate shell edge; 0 – 2 finer secondary ribs between primary ribs, rib interstices darker. Interior shell margin and spatula dark brown to golden tan, often some calcification; off white to cream rays on shell margin align under primary / secondary ribs, extend to spatula; siphonal groove distinct, paler than margin; ADM scar indistinct, similar to margin and spatula; CMS straight; whitening and thickening of shell lip observed. The neotype (Fig. 17 A). Shell (sl = 15.1, sw = 10.8, sh = 5.5 mm) circular ovate; thick, apex offset weakly offset to posterior and left, apical sides straight, 32 ribs, ~ 17 whitish primary ribs, with 0 – 2 in between secondary ribs, rib interstices dark, siphonal ridge formed by adjacent dual primary ribs. Interior evenly dark brown, white rays on shell lip under ribs. Reproductive system (Fig. 18 A; n = 4). Positioned within coelom under the respiratory cavity, hermaphroditic glands positioned to posterior against right foot wall and over foot sole, epiphallic parts between RAM and extending over BM; ED distinct entry to top of small GA, AO small, short, bent, tip blunt, bulbous, appears part of GA under ED; ED thick, short, centrally bent; AO, GA and ED all muscular white tissue; EG very large, elongated, 2 lobes, folded, soft white tissue; single thick elongated blunt flagellum F 1 appears as extension of ED at EG join; BD and CD connect closely together into GA, both ducts smooth, short, broad, slightly bent, pass together through outer edge of RAM (BD over CD) connecting into MG; BC medium to large, bulbous, white opaque test, embedded along with part of BD in soft white folds of MG; HD short, broad, coiled, links soft white folded AG to small yellowish granulated HG; AG larger than HG, both with outer sides curved reflecting the close positioning to curvature of inner foot wall at right posterior quarter of coelom; SV embedded in AG close to BC. Spermatophore (Fig. 18 B). Broad head with short flagellum (length = 4.6 ± 0.57 mm, n = 2); head section cylindrical, bulbous, centrally bent, tip rounded; test thin, smooth, featureless, translucent encasing a white opaque central core; short looped tapering section merges head to filamentous flagellum; head slightly shorter, wider than translucent flagellum (head length = 2.28 ± 0.67 mm, n = 3; flagellum length = 2.38 mm, n = 1; length / head length ~ 52 %; head width = 172 ± 17 μm; flagellum width = 26 ± 0 μm, n = 3); 5 SPMs tightly coiled in cream gelatinous mass in BC of one specimen (Fig. 18 B).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFB8823BFF68FA42FB3CFD96.taxon	diagnosis	Comparativeremarks. Siphonariaradiata (normalis group, unit 12) is the sister species of S. planucosta sp. nov. (Figs 1, 4 C). Both species are separated from each other species by distances of at least 11.5 % in COI (Table S 8). The lowest interspecific genetic distances were observed between S. radiata and S. madangensis sp. nov. (unit 88) = 10.7 %, S. costellata sp. nov. (unit 13) = 11 %, and S. normalis (unit 14) = 11.4 %) (Table S 8). We found a sympatric congener in Sabah: Siphonaria kudatensis (for comparison refer to S. kudatensis). Siphonaria radians has a lower, slightly larger shell with finer ribbing, a less prominent siphonal ridge, weaker edge scalloping, a larger AO, longer twisted ED, longer narrower BD with distal loop and a smaller BC. Shell geometry and sculpture of S. radiata resemble that of S. costellata sp. nov. (unit 13) and S. normalis (unit 14). However, S. radiata possesses slightly fewer and broader primary ribs, fewer secondary ribs, narrower interstices, more prominent siphonal ridge and more scalloped edge than these two species. The interior colouration also differs being generally paler browns to white with prominent red-brown rays extending to spatula in S. radiata. The SPM of S. radiata resembles that of S. normalis and S. gemina sp. nov. The record of ‘ S. cf kurracheensis’ in Way & Purchon (1981: 321) is a misidentification and likely of S. radiata. The specimen figured as ‘ normalis group, unit 12 ’ in Dayrat et al. (2014: 260, fig. 3 N) corresponds well with the present species.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFB8823BFF68FA42FB3CFD96.taxon	distribution	Distribution and habitat. Recorded from W coast of Thailand and Port Dickson, Malacca Strait, Malaysia (Fig. 16). In this study, found on exposed and bare rocky shores, mid littoral level (Fig. 17 U).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFBA8237FCCAFD22FEBCFDD6.taxon	description	(Figs 14 E – I, P, S – T, 18 C – E)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFBA8237FCCAFD22FEBCFDD6.taxon	materials_examined	Material examined. Type material. Probable holotype of S. crenata Blainville, 1827 (type locality unknown; Savigny collection) (MNHN IM 2000 - 35937, Fig. 14 E). Holotype of Siphonaria rosea Hubendick, 1943 from “ Insel Kharg, Persischer Meerbusen ” [Kharg Island, Persian Gulf, Iran]; coll. G. Thorson, 1937 (UUZM 1577, Fig. 14 H). Other, non-type material. Saudi Arabia: Persian Gulf, 28 ° N, 50 ° E (AM C. 69719 d, Fig. 14 I); Pakistan: Karachi, French Beach, 24 ° 50.367 ’ N, 66 ° 49.387 ’ E PA 01 - 1 (AM C. 585851 p [M 242, SK 233], Fig. 14 G, C. 585892 p [M 239], C. 595917 p [SK 534]), PA 01 - 2 (AM C. 585853 p, C. 585338 p [SK 153], Fig. 14 F; C. 585853 p [SK 302]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFBA8237FCCAFD22FEBCFDD6.taxon	discussion	Taxonomic remarks. Savigny (1817: pl. 1, figs 1 – 4; pl. 3, figs 1 – 5) published engravings without captions of two distinct siphonariids (Fig. 14 O – P herein) (see Bouchet & Danrigal, 1982 for bibliographic details). The first reference to Savigny’s figures appeared in Blainville (1825: 655): Under “ et corrections ” for the genus Siphonaria, he referenced Savigny’s figure (“ Ègypt. Gastéropod., pl. 3, fig. 1 – 5 ”) [= S. crenata] as ‘ figured the animal of a species of this genus’ without assigning a name. Later, Blainville (1827 b: 295) introduced the name S. crenata with reference to these figures stating “ Savigny AEgypt. Zoolog.; Gastropodes, pl. 3, fig. 3 – 35. ” without mentioning Savigny’s plate 1. Specimen 4 in the Savigny collection (MNHN IM 2000 - 35937, Fig. 14 E) resembles the specimen figured by Savigny (1817: pl. 3, fig. 3.5; Fig. 14 P herein) with respect to shell edge and prominence of ribbing rather than any other of the figured shells. Bouchet & Danrigal (1982: 15) mentioned three syntypes of S. savignyi to be held by the MNHN (most likely MNHN IM 2000 - 35934, Fig. 14 L; 2000 - 35935, Fig. 14 K; 2000 - 35936, Fig. 15 J). The last two shells closely match in Savigny’s (1817) figures pl. 1, fig. 1 (Fig. 15 K) and pl. 1 figs 2 – 4 (Fig. 14 J) respectively. Our delineation of S. crenata is based on comparative analyses of the morpho-anatomy and mitochondrial genetics of freshly collected specimens of S. crenata (Fig. 14 F) and S. rosea (Fig. 14 G) and geographic series of additional specimens (Tables S 1 – 2). Hubendick (1946: 50) listed among some ‘ transitionalforms’inan‘Indian-WestPacificform-group’, a ‘ S. zanda <> S. rosea ’ form (Hubendick 1946: 50, pl. 5, figs 1, 2). This shell is probably a specimen of ‘ S. rosea’ (= S. crenata, Figs 14 E – F). Hubendick (1946: 54) proposed S. crenata to be a possible synonym of S. kurracheensis. However, type specimens of S. kurracheensis differ from Hubendick’s (1946: 54) interpretation of this taxon, and of the specimens from ‘ Persian Gulf / Suez’ figured as ‘ S. kurracheensis ’ by Hubendick (1946: pl. 2, figs 36 – 40; except fig. 38 from ‘ Java Sea’), are consistent with typical characteristics of S. crenata but not S. kurracheensis. A shell figured as ‘ Siphonaria savignyi ’ from Woody Point, Moreton Bay, Qld, Australia by Hubendick (1955: 2 (MV F 13951 )) is a misidentification and attributed here to S. opposita. Morrison (1972: 61) stated that ‘ S. basseinensis’ mentioned by Tillier & Bavay (1905: 176) is a record of S. crenata; however, this statement is not supported herein following examination of the type specimens of S. crenata (Fig. 14 E) and of S. basseinensis Melvill, 1893 (Fig. 14 M). This nominal species is not assessed in the present study. Morrison (1972: 56 – 58) treated Siphonaria rosea and 29 other nominal species as synonyms of Siphonaria laciniosa based on similarity in shell shape and “ a common reproductive development ”. These synonymies are not accepted herein following examinations of type specimens and comparative morpho-anatomy. Bouchet & Danrigal (1982: 15) incorrectly considered a shell figured by Reeve (1856: pl. 5, species 20) as S. kurracheensis to be identical with S. savignyi.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFBA8237FCCAFD22FEBCFDD6.taxon	description	External morphology. Foot wall, pneumostome, cephalic folds and mantle evenly cream, foot edge paler, foot sole darker; faint black pigmentation over upper foot wall and mid centre of cephalic folds; mantle narrower than foot wall, thin translucent, white banded edge thickened strongly lobed, overlaid with black bands aligning to rib interstices; pneumostomal lobe thin and within mantle between the right ADMs, closes the pneumostomal and anal openings at the mantle edge; genital pore inconspicuous, located on foot wall posterior to right cephalic fold; two small black epithelial eye spots centralised on two thick centrally touching cephalic folds. Shell (Fig. 14 E – G, T; Table S 9). Medium to large sized (max sl mean = 15.2 mm, SD = 1.1, n = 4), ovate; height medium to low; apex offset sightly posterior and central (usually eroded), apical sides strongly convex, protoconch direction weakly heterostrophic (n = 2; Fig. 14 T), shell whorl dextral; growth striae prominent in bands, shell thickness thick; rib count (mean = 32.3, SD = 1.1, n = 4), primary ribs pale white, fairly straight, increasingly raised and protrude beyond shell lip to unevenly scallop and corrugate the edge; 1 – 2 interspersed pale white finer secondary ribs, rib interstices darker; paired primary ribs on siphonal ridge, no more prominent than other primary ribs. Interior shell margin dark brown to tan, white rays align on shell margin under primary / secondary ribs, siphonal groove distinct, same colour as shell edge, points to right anterior; spatula dark chocolate brown to mottled tan even whitish (Fig. 14 F – G); ADM scar distinct, CMS straight, paler than shell lip; thickening of shell lip translucent, infills and reduces lip scalloping, spatula becomes whitened. Reproductive system (Fig. 18 C – E; n = 4). Positioned within right side of coelom, against foot wall on foot muscle, under the respiratory cavity; epiphallic parts positioned between RAM and BM; GA small indistinct with singular GP through foot wall; AO large wide bluntly pointed, joined to upper GA; ED short wide thickened, bent, joins to GA; GA, AO, ED all white muscular fibrous tissue; EG large, soft whitish tissue, folded, joins ED; single short flagellum (F 1) shorter narrower than ED, lays over EG, appears as an extension of ED. BD and CD connect closely in opposing directions to GA between AO and GP, both ducts narrow elongated bent smooth whitish, pass together through RAM (BD over CD) into soft white folded tissues of MG, BC partly embedded in folds, small flat bulbous, thin whitish translucent test (0 SPM in brownish gelatinous mass of BC); MG / AG complex relatively small; HD short narrow coiled, links ducts in soft white folded tissues of AG to yellowish granulated HG; AG / MG larger than HG, sides match curvature of inner foot wall.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFBA8237FCCAFD22FEBCFDD6.taxon	diagnosis	Comparative remarks. Siphonaria crenata (atra group, unit 43) is member of Clade H in the atra group (Figs 1, 2) forming a subclade together with S. belcheri and S. madagascariensis. It is well-differentiated from other species by COI distances of ≥ 21.4 % (Table S 5). Within its range, we found four species in partial sympatry with S. crenata (e. g., in Karachi, Pakistan): Siphonaria asghar has a smaller, taller, paler shell with a less distinct siphonal ridge and weaker edge scalloping, smaller AO, shorter wider BD without distal loop, and a smaller BC. Siphonaria belcheri has a smaller, taller shell with less raised ribbing, weaker edge scalloping, darker reddishbrown interior, and slightly longer BD. Siphonaria kurracheensis has a smaller shell with less edge-protruding ribbing and weaker edge scalloping, and a smaller AO. Siphonaria perexigua sp. nov. has a smaller, taller shell with a less prominent siphonal ridge, less raised ribbing, weaker edge scalloping, a paler interior, smaller AO, longer ED, and a relatively larger BC. Hubendick’s (1943: 3, fig. 9) illustration of the RS of S. rosea from the Persian Gulf corresponds well with the RS of S. crenata shown herein (Figs 18 C – E) in terms of having a long thin BD, short CD, large EG, short ED and F 1, and a large, twisted AO. Moreover, figured shells of ‘ S. rosea ’ in Hubendick (1943: 1, fig. 1 a – b, 1946: 91, pl. 4, fig. 12 – 15 from the same locality are consistent with typical features of S. crenata as herein delimited. Shells of ‘ S. savignyi’ figured in Hubendick (1946: 92, pl. 4, fig. 25 – 29) from ‘ Port Tewfick, Red Sea’ and ‘ Gulf of Suez’ are also likely specimens of S. crenata. Figured specimens of ‘ S. kurracheensis ’ in Bosch et al. (1995: 186, fig. 863) and Ali et al. (2011: 1086, fig. 1 B) are herein considered to represent S. crenata as well for corresponding shell size, ribbing, and extension of siphonal ridge.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFBA8237FCCAFD22FEBCFDD6.taxon	distribution	Distribution and habitat. This species is recorded from the Red Sea and the Persian Gulf through to Karachi, Pakistan (Fig. 28). In this study found in sheltered positions on exposed rocky intertidal shores, upper littoral level (Fig. 14 S).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFB68233FF68FDE2FDF0F8F6.taxon	description	(Figs 17 C – G, O – Q, 20 A – C)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFB68233FF68FDE2FDF0F8F6.taxon	materials_examined	Material examined. Type material. Lectotype of Siphonaria diemenensis Quoy & Gaimard, 1833, present designation, from ‘ le canal de d’Entrecasteaux, á l’ î le de van Diémen’ [d’Entrecasteaux Channel, Tasmania] (MNHN-IM- 2000 - 35952, Fig. 17 C). Paralectotype, same data as lectotype (MNHN-IM- 2000 - 5059). Other, non-type material. Australia, NSW: Terrigal, The Skillion, 33 ° 27.008 ’ S, 151 ° 27.122 ’ E, NSW 08 - 2 (AM C. 585631 5 p); Wy-ar-gine Point Balmoral, 33 ° 49.159 ’ S, 151 ° 15.195 ’ E, NSW 06 - 5 (AM C. 585630 6 p); Laings Point Sydney Harbour, 33 ° 50.419 ’ S, 151 ° 16.638 ’ E, NSW 06 - 3 (AM C. 585404 17 p); Bombo Kiama, 34 ° 39.232 ’ S, 150 ° 51.649 ’ E, NSW 03 - 1 (AM C. 585682 7 p); Murunna Point Camel Head, 36 ° 22.720 ’ S, 150 ° 04.766 ’ E, NSW 02 - 1 (AM C. 585402 10 p, C. 585001 p [SK 037]); Oman Point Eden, 37 ° 04.634 ’ S, 149 ° 53.445 ’ E, NSW 01 - 1 (AM C. 585527 20 + p). Vic: Bastion Head Mallacoota, 37 ° 34.429 ’ S, 149 ° 45.927 ’ E, V 09 - 1 (AM C. 585611 4 p); Cape Conran, 37 ° 48.798 ’ S, 148 ° 43.608 ’ E, V 08 - 2 (AM C. 585543 20 + p, C. 585293 p [M 196]); Frankston, 38 ° 09.236 ’ S, 145 ° 06.457 ’ E, V 06 - 1 (AM C. 585356 p); Point Lonsdale (nr Queenscliff), 38 ° 17.276 ’ S, 144 ° 36.977 ’ E, V 05 - 1 (AM C. 585436 10 p); Port Fairy, 38 ° 23.692 ’ S, 142 ° 14.260 ’ E, V 01 - 1 (AM C. 585694 7 p); Roadknight Point, 38 ° 25.707 ’ S, 144 ° 11.102 ’ E, V 04 - 1 (AM C. 585716 8 p, C. 585286 p [M 168], C. 585287 p [M 169]); WestHeadFlinders, 38 ° 28.883 ’ S, 145 ° 01.727 ’ E, V 06 - 3 (AM C. 585542 20 + p, (AM C. 585290 p [SK 003], C. 585291 p [M 198]); Loutit Bay Lorne, 38 ° 31.190 ’ S, 143 ° 59.429 ’ E, V 03 - 2 (AM C. 585541 20 + p); San Remo, 38 ° 31.489 ’ S, 145 ° 21.858 ’ E, V 07 - 1 (AM C. 585357 p [SK 029]); Crofts Bay, 38 ° 35.363 ’ S, 142 ° 50.633 ’ E, V 01 - 3 (AM C. 585680 6 p); Marengo Rocks Apollo Bay, 38 ° 46.772 ’ S, 143 ° 39.997 ’ E, V 03 - 1 (AM C. 585433 10 + p). Tas: Georges Bay, Burns Bay, 41 ° 16.62 ’ S, 148 ° 28.92 ’ E (TMAG E 41975 3 p); Beaumaris, Shelly Point, 41 ° 26.136 ’ S, 148 ° 16.638 ’ E (TMAG E 41965 4 p); Denison Beach, Porch Rocks, 41 ° 47.334 ’ S, 148 ° 16.158 ’ E (TMAG E 41963 2 p); Bicheno, 41 ° 52.837 ’ S, 148 ° 18.525 ’ E, T 02 - 1 (AM C. 585692 7 p); Bicheno Courland Bay, 41 ° 55.866 ’ S, 148 ° 18.414 ’ E (TMAG E 41960 p); Swansea Spiky Beach, 42 ° 11.142 ’ S, 148 ° 4.104 ’ E (TMAG E 41961 7 p); Louisville, Alginate Bay, 42 ° 32.4 ’ S, 147 ° 54.6 ’ E (TMAG E 24490 p); Maria Island, Darlington Bay, 42 ° 34.8 ’ S, 148 ° 3.6 ’ E (TMAG E 16769 2 p); Maria Island, Hopground Beach, 42 ° 35.4 ’ S, 148 ° 3.6 ’ E (TMAG E 08499 3 p); Earlham, foreshore, 42 ° 39.6 ’ S, 147 ° 57 ’ E (TMAG E 08098 6 p); Dodges Ferry, 42 ° 51.083 ’ S, 147 ° 36.981 ’ E, T 03 - 1 (AM C. 585428 10 + p); Dodges Ferry, Red Ochre Beach, 42 ° 51.6 ’ S, 147 ° 36.6 ’ E (TMAG E 08843 p); Park Beach, Dodges Ferry, 42 ° 51.716 ’ S, 147 ° 36.665 ’ E, T 03 - 4 (AM C. 585260 p [M 170], C. 585261 p [SK 016]); Carlton Beach, Spectacle Island, 42 ° 52.044 ’ S, 147 ° 36.024 ’ E (TMAG E 41955 4 p); Primrose Sands, 42 ° 53.4 ’ S, 147 ° 40.2 ’ E (TMAG E 04957 p); Lauderdale, Roches Beach, 42 ° 54.6 ’ S, 147 ° 0.3 ’ E (TMAG E 03265 p); Lagoon Bch (near Saltwater River), 42 ° 56.903 ’ S, 147 ° 39.962 ’ E, T 03 - 2 (AM C. 585253 p [SK 015], C. 585254 p [SK 185]); Taroona Beach, 42 ° 57.18 ’ S, 147 ° 21 ’ E (TMAG E 41969 14 p); Kingston Beach, 42 ° 58.8 ’ S, 147 ° 19.2 ’ E (TMAG E 02012 p); Eaglehawk Neck, eastern side, 43 ° 0.6 ’ S, 147 ° 0.6 ’ E (TMAG E 01661 3 p); Blackmans Bay, 43 ° 0.6 ’ S, 147 ° 19.8 ’ E (TMAG E 15878 8 p); Saltwater River, 43 ° 01.083 ’ S, 147 ° 43.578 ’ E, T 03 - 11 (AM C. 585725 7 p); Tasman Arch, 43 ° 02.033 ’ S, 147 ° 56.963 ’ E, T 03 - 3 (AM C. 585771 5 p, C. 585256 p [M 175], C. 585257 p [M 176]); Pirates Bay, Fossil Island, 43 ° 1.8 ’ S, 147 ° 57 ’ E (TMAG E 01299 p); Tinderbox Beach, 43 ° 3.6 ’ S, 147 ° 19.8 ’ E (TMAG E 05223 3 p); North Bruny Island, Dennes Point, 43 ° 3.87 ’ S, 147 ° 21.066 ’ E (TMAG E 41968 3 p); Nubeena, Parsons Bay, 43 ° 6 ’ S, 147 ° 6 ’ E (TMAG E 05994 p); Kettering, Oyster Cove, 43 ° 6.6 ’ S, 147 ° 16.2 ’ E (TMAG E 05350 3 p); Nubeena, White Beach, 43 ° 7.2 ’ S, 147 ° 43.8 ’ E (TMAG E 06016 p); North Bruny Island, Simmonds Bay, 43 ° 7.8 ’ S, 147 ° 21.6 ’ E (TMAG E 02677 2 p); Fortescue Bay, 43 ° 8.4 ’ S, 147 ° 57.6 ’ E (TMAG E 05581 2 p); Three Hut Point d’Entrecasteaux Channel, 43 ° 16.195 ’ S, 147 ° 14.414 ’ E, T 04 - 3 (AM C. 585482 18 p, C. 584797 p [SK 047], C. 585269 p [M 112]); Huon Point d’Entrecasteaux Channel, 43 ° 17.471 ’ S, 147 ° 05.778 ’ E, T 04 - 1 (AM C. 585430 10 + p); South Bruny Island Coal Point, 43 ° 19.8 ’ S, 147 ° 19.8 ’ E (TMAG E 35239 2 p); South Bruny Island, Adventure Bay, Cemetery Bluff, 43 ° 20.388 ’ S, 147 ° 19.44 ’ E (TMAG E 41956 2 p); Moss Glen, 43 ° 31.910 ’ S, 146 ° 53.641 ’ E, T 05 - 1 (AM C. 585270 p [M 107], C. 585271 p [M 115]); Fishers Point, 43 ° 34.260 ’ S, 146 ° 55.244 ’ E, T 05 - 4 (AM C. 585570 3 p); Flensing Rock, 43 ° 34.291 ’ S, 146 ° 54.856 ’ E, T 05 - 2 (AM C. 585714 8 p, C. 585272 p [SK 013]); Pancake Bay, 43 ° 34.673 ’ S, 146 ° 55.293 ’ E, T 05 - 5 (AM C. 585273 p [M 113]; Cape Portland, Petal Point, 40 ° 46.566 ’ S, 147 ° 56.604 ’ E (TMAG E 41972 3 p); Herbies Landing 40 ° 50.112 ’ S, 147 ° 38.724 ’ E (TMAG E 41966 4 p); Bridport — beach 40 ° 59.808 ’ S, 147 ° 23.526 ’ E (TMAG E 41957 p); S of Granite Point Bridport, 40 ° 59.739 ’ S, 147 ° 23.468 ’ E, T 01 - 1 (AM C. 585550 22 p, C. 595920 p [SK 552]); Tamar River, Kelso, foreshore, 41 ° 6 ’ S, 146 ° 6 ’ E (TMAG E 16716 2 p); Aikenhead Point, Devonport, 41 ° 09.997 ’ S, 146 ° 21.927 ’ E, T 01 - 2 (AM C. 585691 7 p); Goat Island, foreshore of island and nearby mainland, 41 ° 8.214 ’ S, 146 ° 8.196 ’ E (TMAG E 41962 p); Somerset, foreshore, 41 ° 2.178 ’ S, 145 ° 49.962 ’ E (TMAG E 41975 14 p); Somerset, 41 ° 01.168 ’ S, 145 ° 47.632 ’ E, T 01 - 3 (AM C. 585461 14 p, C. 585252 p [M 117]); Wynyard, Inglis River mouth, 40 ° 58.8 ’ S, 145 ° 43.8 ’ E (TMAG E 02094 2 p); Boat Harbour, beach, 40 ° 55.8 ’ S, 145 ° 37.2 ’ E (TMAG E 07953 p); Rocky Cape, Castle Rock Bay, 40 ° 53.238 ’ S, 145 ° 31.356 ’ E (TMAG E 41971 6 p); Rocky Cape, Picnic Beach & rocks to S, 40 ° 52.026 ’ S, 145 ° 29.052 ’ E (TMAG E 41958 p); Little Peggs Beach 40 ° 51.084 ’ S, 145 ° 21.384 ’ E (TMAG E 41970 p); Stanley, Godfreys Beach, 40 ° 45.138 ’ S, 145 ° 17.67 ’ E (TMAG E 41959 p); Marrawah, Green Point, 40 ° 54 ’ S, 144 ° 54 ’ E (TMAG E 09069 4 p); Lucas Point Pilot Bay Macquarie Harbour, 42 ° 12.241 ’ S, 145 ° 12.005 ’ E, T 06 - 1 (AM C. 585479 17 p); Trial Harbour, 41 ° 55.758 ’ S, 145 ° 10.434 ’ E (TMAG E 41964 5 p); King Island: E coast, 39 ° 54.6 ’ S, 144 ° 6.6 ’ E (TMAG E 04045 p); Fitzmaurice Bay, 40 ° 3.708 ’ S, 143 ° 52.896 ’ E (TMAG E 41967 p); Lavinia Beach, beach nr Lake Martha Lavinia, 39 ° 39.018 ’ S, 144 ° 4.692 ’ E (TMAG E 41973 2 p). SA: Cape Northumberland, 38 ° 03.503 ’ S, 140 ° 40.378 ’ E, SA 15 - 1 (AM C. 585452 12 p); Cape Northumberland Port Macdonnell, 38 ° 03.308 ’ S, 140 ° 39.398 ’ E, SA 15 - 2 (AM C. 585676 6 p, C. 595918 p [SK 085]); Cape Thomas, 37 ° 04.461 ’ S, 139 ° 44.659 ’ E, SA 14 - 1 (AM C. 585426 10 + p); Fisheries Bay Lands End, 35 ° 37.999 ’ S, 138 ° 06.921 ’ E, SA 13 - 2 (AM C. 585464 15 p); Groper Bay nr West Cape, 35 ° 14.108 ’ S, 136 ° 49.883 ’ E, SA 10 - 1 (AM C. 585722 9 p); Pondalowie Bay, 35 ° 13.989 ’ S, 136 ° 49.892 ’ E, SA 10 - 2 (AM C. 585712 8 p, C. 585215 p [SK 014]); Port Neill, 34 ° 07.102 ’ S, 136 ° 21.271 ’ E, SA 06 - 1 (AM C. 585423 20 + p); Port Moonta, 34 ° 03.273 ’ S, 137 ° 33.592 ’ E, SA 09 - 1 (AM C. 585386 10 p); Salmon Point, 33 ° 38.547 ’ S, 134 ° 51.916 ’ E, SA 04 - 2 (AM C. 585549 22 p); Wellesley Point, 33 ° 38.483 ’ S, 134 ° 51.963 ’ E, SA 04 - 1 (AM C. 585422 10 + p); Whyalla, 33 ° 02.539 ’ S, 137 ° 35.511 ’ E, SA 07 - 1 (AM C. 585721 9 p); Rocky Point, 32 ° 12.250 ’ S, 133 ° 14.861 ’ E, SA 02 - 4 (AM C. 585421 10 + p); Cape Thevenard, 32 ° 08.703 ’ S, 133 ° 38.553 ’ E, SA 03 - 3 (AM C. 585708 8 p); Ceduna, 32 ° 07.438 ’ S, 133 ° 40.260 ’ E, SA 03 - 2 (AM C. 585451 12 p); Port Le Hunte Point Sinclair, 32 ° 05.554 ’ S, 132 ° 59.476 ’ E, SA 02 - 2 (AM C. 585447 11 p); Cactus Beach Point Sinclair, 32 ° 05.135 ’ S, 132 ° 58.943 ’ E, SA 02 - 3 (AM C. 595959 11 p).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFB68233FF68FDE2FDF0F8F6.taxon	discussion	Taxonomic remarks. The lectotype (Fig. 17 C) is designated herein for the stabilisation of the name and to ensure the unambiguous identity of this taxon (Art. 74.1 of the Code). Our delineation of this species is based on comparative analyses of the morpho-anatomy and mitochondrial genetics of freshly collected topotypes and geographic series of additional specimens (Tables S 1 – 2). We also demonstrate that previous treatments of S. denticulata as a synonym or variety of S. diemenensis are incorrect and that S. diemenensis and S. denticulata are distinct species. Menke (1844: 54) regarded S. diemenensis (as “ diemensis ”) as ‘ closely related and associated’ with S. javanica (as “ iavana ” and “ iavanica ”), which is incorrect. The shell of S. diemenensis has more numerous and narrower ribs than that of S. javanica. Both species also differ in RS structure: AO larger, BD longer, narrower, F 1 shorter in S. diemenensis than S. javanica. Both species have non-overlapping distributions in cool temperate waters of SE Australia (Fig. 16) and in the Tropical Philippines (Fig. 11), respectively. Tryon & Pilsbry’s (1891: 25) reference to Patella diemensis Philippi, 1848 from Hobarttown, Tasmania, as a possible siphonariid is incorrect. This taxon is a junior synonym of Patella peronii Blainville, 1825 (Coan & Kabat, 2017: 110). Hutton (1878: 41) stated that he earlier misidentified specimens of ‘ Benhamina obliquata ’ (= S. obliquata) from NZ as S. diemenensis. Subsequently, Suter (1909 b: 33, 1913: 599) and Hubendick (1946: 24) agreed. Hutton (1878: 10, 42) also incorrectly listed S. denticulata as a synonym of S. diemenensis and was uncertain whether S. diemenensis inhabited NZ. Herein, we establish that S. diemenensis is not present in NZ. Verco (1907: 104) incorrectly considered Trimusculus albida (Angas, 1878) as a possible white form of S. diemenensis. According to Oliver (1915: 546), Suter’s (1907) specimens of S. diemenensis were a misidentification of S. raoulensis. Hubendick (1946: 38) treated S. denticulata Quoy & Gaimard, 1833, S. scabra Reeve, 1856, S. exulum Hanley, 1858 (including ‘ Ellsiphon exulorum’ Iredale, 1940) and S. raoulensis Oliver, 1915 as synonyms of S. diemenensis. Hubendick (1946: 38 – 39) also treated another five taxa as varieties of the former. We consider that four of these names are incorrectly assigned to S. diemenensis (i. e., denticulata Quoy & Gaimard, 1833, scabra Reeve, 1856, exulum Hanley, 1858, and perplexa Oliver, 1915). Siphonaria exulum, S. perplexa and S. macauleyensis are all conspecific with S. e xulum. Hubendick’s (1946: 38) inclusion of S. javanica Blainville, 1827 and S. alternicosta Potiez & Michaud, 1838 as possible synonyms of S. diemenensis are also rejected. McAlpine (1952: 41) erroneously treated S. scabra as a junior synonym of S. diemenensis. Yet, these are distinct species. Grove et al. (2006: 60) incorrectly listed S. denticulata, S. javanica Blainville 1827 (not Lamarck, 1819), S. exulum and S. scabra as synonyms of S. diemenensis.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFB68233FF68FDE2FDF0F8F6.taxon	description	External morphology (Fig. 17 O). Foot sole smooth, reddish orange, darker at foot edge; foot wall dark yellowish grey with evenly spread white subepithelial pustules becoming more vivid and dense close to the foot sole and around pneumostomal lobe; genital pore inconspicuous, located on foot wall posterior to right cephalic fold; fringing mantle reasonably wide, thin, yellowish, translucent to transparent, extends to shell edge, outer edge lobed, weakly banded yellow and black reflecting corrugations and inner colouration of shell lip and ribs; pneumostomal lobe large and within mantle between the right ADMs, closes the pneumostomal and anal openings at the mantle edge; two small black epithelial eye spots centralised on two thick centrally touching orange grey cephalic folds that darken to their outer edge, covered with white mucous cells similar (but smaller) to those of the foot wall tissue. Shell (Figs 17 C – G, Table S 9). Small to medium sized (max sl mean = 17.7 mm, SD = 3.1 mm, n = 8), ovate; height tall; apex central, usually eroded, apical sides strongly convex, protoconch direction homostrophic (n = 6; Fig. 17 P), shell whorl dextral; growth striae indistinct with weak external colour banding aligning with growth stages; rib count (mean = 29, SD = 5.6, n = 8), primary ribs off white, strongly raised, rounded and straight, extend past a fairly even shell lip to scallop the edge; very few secondary ribs, rib interstices dark, strongly prominent, paired primary ribs over an indistinct siphonal ridge; interior shell margin coloured white aligning under primary ribs, very dark brown to black aligning under rib interstices, colour extends from shell lip to spatula region, spatula yellowish brown; ADM scar indistinct, paler colour than spatula, CMS straight. Shell thickening not displayed. Reproductive system (Figs 20 A – C; n = 2). Positioned within coelom under the respiratory cavity, over foot muscle and against right side of foot wall. GA, EG and ED positioned between BM and RAM; single GP inconspicuous, opening from an extremely small GA through foot wall posterior to right cephalic fold; AO short, wide, bluntly pointed; join of AO, GA and ED distinct, all whitish fibrous muscular tissue; ED very long, thin, bent, twisted; EG wide, lobed, with single curved, indistinct flagellum (F 1); BD and CD very thin and long, jointly connect into GA between ED, AO and GP; BD noticeably longer than CD with a prominent central loop over epiphallic parts before joining BC (no distal loop), both ducts smooth, featureless, pass together outside RAM connecting into curved MG, BD ventral to CD; BC bulbous elongated, thin test, internally embedded in folds of MG (0 to 5 SPM in BC); HD long, thick, coiled, links ducts in folds of small AG to elongated narrow yellowish granulated HD; AG larger than HG, MG and AG folded, soft white tissue; SV embedded on left side of AG under BC; outer edge of AG and HG sides match curvature of inner foot wall. Spermatophore (Fig. 20 B). Thread-like (length = 9.35 ± 3.61 mm, n = 2), translucent, test thin; head section, tip bluntly rounded, weakly bulbous, elongate, containing a white gelatinous mass; tapers quickly into consistently filamentous transparent flagellum; both sections smooth, featureless. Head shorter and thicker than flagellum (head length = 4.7 ± 0.7 mm, n = 2; mean ~ 20 % of SPM length; head width = 260 ± 42 μm; flagellum width = 45 ± 21 μm, n = 2). Typically, tightly coiled in bursa, embedded in white gelatinous mass. The SPM (Fig. 20 B) reasonably matches the SPM of S. diemenensis figured in Hubendick (1945: 23, fig, 29) apart from width of head section. Radula (Figs 83 A – D). Dentition formula 35: 1: 35 with 141 transverse rows, n = 1, AM C. 201772); single central rachidian tooth flanked squarely by 35 half row laterals, of which 3 are inner, 7 mid and 25 outer laterals (n = 1); central tooth with narrow pointed mesocone, inner lateral mesocones predominantly bicuspidate but with an irregular few unicuspidate, endo and ectocones absent (Figs 83 A – B); mid laterals with pointed mesocone; single ectocone, protrudes at acute angle halfway along the tooth’s length; outer laterals with a broad ‘ chisel’ shaped pointed mesocone flanked by small, single, pointed ecto and endocones (Fig. 83 C), angle of separation of each cone from the mesocone varies, basal plate of outer laterals typically as broad as the tooth’s length.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFB68233FF68FDE2FDF0F8F6.taxon	diagnosis	Comparative remarks. Siphonaria diemenensis (lateralis group, unit 7) represents the sister group of a clade containing the three species S. jeanae, S. propria, and S. australis (Figs 1, 4). It is well-differentiated from these species by COI distances of ≥ 18.8 % (S. jeanae), ≥ 19.5 % (S. propria), and ≥ 19.9 % (S. australis) (Table S 8). From other species it differs by COI distances of ≥ 20 % (Table S 8). Within its range we found eight partially sympatric congeners. With four of these species, it co-occurs in Sydney, NSW, Australia: Siphonaria pravitas sp. nov. has a lower shell with stronger raised ribs and edge scalloping, no BD distal loop, larger BC, wider F 1, shorter SPM. Siphonaria scabra has a paler shell with finer ribbing and coarser exterior, a longer AO, BC and F 1, and a bursal loop. Siphonaria denticulata has a larger, lower shell with less prominent ribbing and a more scalloped edge, a smaller, pointed AO, shorter and wider ED, no bursal loop and a larger BC. Siphonaria emergens has a smaller, elongate, paler brown shell with a strongly offset apex, less prominent ribbing and edge scalloping. A further five congeners are sympatric with S. diemenenesis in SE Australia: Siphonaria funiculata has a paler, taller shell with less raised ribs broadening to a fainter scalloped edge, a larger AO, shorter BD and ED, absent to smaller F 1, and a larger, drop-like SPM. Siphonaria stowae has a smaller, elongate, and paler shell with a strongly offset apex, less prominent ribbing and edge scalloping, a smaller AO, shorter ED, larger BC, and shorter SPM. Siphonaria tasmanica has a lower, bluish coloured shell with a more offset apex, fainter edge scalloping and less raised ribs broadening to edge, a smaller AO and BD, shorter ED, and short, drop-like SPM. Siphonaria jeanae has a smaller, has a smaller, lower, bluish coloured shell, a smaller AO, shorter ED, and short, drop-like SPM. Siphonaria zelandica has a paler, lower shell with less prominent and finer, narrower ribbing, a shorter ED and BD, and short, drop-like SPM. Exterior shell characteristics of S. diemenensis resemble those of S. denticulata. Hubendick’s (1946) treatment of S. scabra and S. denticulata as varieties of S. diemenensis is rejected as these are accepted herein as distinct species. A specimen figured as ‘ S. diemenensis ’ by Davey (1998: 113, text-fig.) is identified herein as S. funiculata. Our description of the radula matches that of Hutton (1883: 124, pl. 17, figs E – G), but the dentition formula is more than twice the count of 12: 1: 12 given by Hubendick (1946: 38). The RSs figured herein (Figs 20 A, C) closely matches figures by other authors (e. g., Hubendick 1945: fig. 23; McAlpine, 1952: 43, fig. 2). The RS and SPM (Fig. 20 B). Hubendick; ’ s (1946: 14, fig. 19) illustrations of RS and SPM of ‘ S. bifurcata ’ from ‘ Port Jackson’ (= Sydney) are herein attributed to S. diemenensis with which the largely correspond apart from a narrower and shorter SPM head section. The specimen figured as ‘ unit 7 ’ in Dayrat et al. (2014: fig. 3 H) from Portland, Vic closely resembles S. diemenensis. This unit was subsequently referred to as ‘ Siphonaria sp. (Australia) ’ in González-Wevar et al. 2018. The record of ‘ S. diemanensis Quoy & Gaimard 1835 ’ (sic) in Smythe (1979: 69) from the United Arab Emirates is incorrect and well outside the known distribution of this species.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFB68233FF68FDE2FDF0F8F6.taxon	distribution	Distribution and habitat. Endemic to cooltemperate waters of south coast Australia, between just north of Sydney through to west of Cape Northumberland, South Australia, including Tasmania (Fig. 16). In this study found on exposed places on intertidal rocky shores, common across upper and mid littoral, associated with mussel and barnacles, home scars common.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFB2822DFF68F8C2FD09FA76.taxon	description	(Figs 15 I – J, 17 H – J, R – T)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFB2822DFF68F8C2FD09FA76.taxon	materials_examined	Material examined. Type material. Lectotype of Siphonaria australis Quoy & Gaimard, 1833 from ‘ détroit de Cook, à la Nouvelle-Zélande’ [Cook Strait, New Zealand] MNHN IM 2000 - 5036 (Fig. 17 G); paralectotype (MNHN-IM- 2000 - 5037), same data as lectotype. Holotype of Siphonaria inculta Gould, 1846 USNM 5857, possible paratype MCZ 169190 (Fig. 17 K), same data as holotype (figured in Gould, 1852: 358, fig. 465, 465 a, 465 b and in Jenkins, 1983: 21, pl. 3 b). Lectotype of Siphonaria cancer Reeve, 1856, NHMUK 198112 (figured in Jenkins, 1983: 21, pl. 3 c), four paralectotypes NHMUK 198112, same data as lectotype (figured in Jenkins, 1983: 21, pl. 3 d). Other, non-type material. NZ; North Island: Big Bay, Manakau Harbour, Auckland, 37 ° 02.58 ’ S 174 ° 38.72 ’ E (AM C. 265856 p [SK 511 protoconch I 6], Fig. 17 S). South Island: N end Seaview Marina, Lower Hutt, 41 ° 14.85 ’ S, 174 ° 54 ’ E (NMNZ M. 331451, 2 p). Stewart Is: Watercrest Bay, 46 ° 54.223 ’ S, 168 ° 07.195 ’ E STI 01 - 1 (AM C. 585953 4 p, C. 585247 p [M 480, SK 289] Fig. 17 H); E of Ringaringa Bch, 46 ° 54.544 ’ S, 168 ° 08.646 ’ E STI 02 - 1 (AM C. 585952 6 p, C. 585249 p [M 481, SK 088], Fig. 17 I).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFB2822DFF68F8C2FD09FA76.taxon	discussion	Taxonomic remarks. The lectotypes of S. australis and S. cancer have been designated by Jenkins (1983). Our delineation of S. australis is based on comparative analyses of the morpho-anatomy and mitochondrial genetics of freshly collected topotypes of S. australis (Figs 17 I – J), S. inculta, S. cancer and S. cookiana (Tables S 1 – 2). Siphonaria spinosa Reeve, 1856 is a junior synonym of S. aspersa Krauss, 1848 as based on examination of the types (Jenkins, 1983: 29). Reeve (1856: pl. 7, fig. 32 a, b) incorrectly stated this taxon to originate in NZ. Siphonaria inculta Gould, 1846 b was first figured in Gould (1852: 358, fig. 465 a, b) and is considered a junior synonym of S. australis. Reference to ‘ S. cancer Reeve ’ in Swinhoe (1865: 165) from Formosa [Taiwan] is based on a misidentification. Reference to S. zelandica, as a synonym of S. sipho, in Schrenck (1867: 306) is erroneous and a probable misidentification of S. australis. Hutton (1878 a: 41) considered S. cancer to be a probable variety of ‘ S. zelandica ’, but is S. australis. He also (p. 42) misidentified specimens from ‘ New Zealand’ (probably of S. australis) as ‘ S. diemenensis’. Hubendick’s (1946: 49) interpretation of S. australis is partly confused: He incorrectly included S. zelandica and S. lentula (as lentulus) in the synonymy of S. australis, which is rejected herein. The nominal species S. scutellum (= S. obliquata) and P. innocuus (= S. exulum) are here removed from the synonym of S. australis, but the synonymy of S. inculta and S. cancer is maintained. Morrison (1972: 56 – 58) treated S. cancer as a synonym of S. laciniosa based on conchological similarity and ‘ common reproductive development’. This is rejected herein based on examinations of types and comparative morpho-anatomy. Raven & Bracegirdle (2010: 46) incorrectly treated S. zelandica as a synonym of S. australis. The re-description of this species by Jenkins (1983) forms the foundation for the delimitation of this species in the present study.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFB2822DFF68F8C2FD09FA76.taxon	description	External morphology (Fig. 17 R). Foot sole smooth, pale cream / yellowish, paler to foot edge; foot wall pale grey-cream with irregular dark / black markings along foot wall & concentrated over cephalic lobes; genital pore conspicuous, located on foot wall posterior to right cephalic fold; fringing mantle thick, cream, translucent to transparent, extends to shell edge, outer edge with thick lobes, strong dark bands aligning with shell lip ribinterstices; pneumostomal lobe large and within mantle between the right ADMs, closes the pneumostomal and anal openings at the mantle edge; two small black epithelial eye spots centralised on two thick centrally touching cephalic folds that darken to their outer edge. Shell (Figs 17 G – I; Table S 9). medium sized, ovate (max sl mean = 17.99 mm, SD = 1.93 mm, n = 99), asymmetrical, thick, height low to medium, edge often very uneven, exterior grey / light brown, uneven; rib count (mean = 28.5, SD = 4.4, n = 99), raised irregularly spaced radial ribs, broaden to edge, ridges rounded to flat; apex weakly offset to posterior and left, protoconch direction homostrophic (n = 1, Fig. 17 S), body whorl dextral; apical sides convex increasingly to shell margin, growth striae indistinct discontinuous, radial ribs project unevenly weakly beyond shell edge, particularly in larger specimens, dual primary ribs form siphonal ridge, project furthest; rib interstices brown, furrowed often fairly deep. Interior glossy, uneven, light brown to tan, margin chocolate brown with white rays aligning under ribs; ADM scar paler, spatula pale blue / yellowish tan, siphonal groove usually mottled dark brown or grey, CMS straight to convex. Reproductive system (Fig. 15 I; n = 3). Positioned within coelom under the respiratory cavity, hermaphroditic glands positioned to posterior against right foot wall & over foot sole, epiphallic parts positioned between BM and to side of RAM; GA bulbous, GP distinct, protrudes; AO indistinct, merges with bulbous GA; ED relatively short, narrow, twisted close to GA; EG large, unfolded, rounded; single long, uncoiled flagellum F 1, extends from end of wider EG; AO, GA and ED all muscular white tissue; BD and CD junctions into GA side-by-side & close to opposing ED connection; BD longer, slightly narrower than CD without distal loop, both ducts, fused, smooth unfeatured, pass together through outer side of RAM connecting into folds of MG (BD above CD); BC translucent test, midsized, elongate bulbous, embedded in AG; large heavily coiled HD, connects soft white folded tissues of AG to smaller elongated brownish finely granulated HG; MG at distal end of AG; dark SV centrally embedded within AG near top of BC, AG larger than HG, sides match curvature of inner foot wall at right posterior quarter of coelom. Spermatophore (Fig. 15 J). Body cylindrical, thread-like (length = 6.1 ± 2.4 mm, n = 2), test thin, translucent; head tip tapered bluntly rounded, section containing a white gelatinous core, tapers to a thin flagellum & tip; both sections smooth, featureless; head longer, thicker than flagellum (head length = 5.27 ± 2.19 mm, ~ 86 % of SPM length; head width = 89 μm ± 7 μm, flagellum width = 24 ± 10 μm, n = 2). Typically, SPM tightly coiled in BC, embedded in white gelatinous mass. Radula (adapted from Jenkins 1983: 17). Mean dentition formula 35: 1: 35 (SD = 7.01) with 127 (SD = 7.33, n = 5) transverse rows; 35 half row laterals, of which 20 mid (SD = 4.98) and 15 outer (SD = 3.58) laterals (n = 5); central tooth single pointed mesocone, inner laterals absent, mid laterals with bluntly pointed mesocone and broad pointed ectocone (Jenkins, 1983, pl. 6 d, e), ectocone strong, protrudes at acute angle halfway along the tooth’s length; outer lateral with a ‘ chisel’ shaped pointed mesocone flanked by small, irregularly pointed, single ecto and endocones (Jenkins 1983: pl. 6 f), angle of separation of each cone from the mesocone varies. Dentition formula: 36: 1: 36 (Hutton, 1883: 142), 30: 1: 30 (Hubendick 1946: 49).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFB2822DFF68F8C2FD09FA76.taxon	diagnosis	Comparative remarks. Siphonaria australis (lateralis group, unit 86) is the sister species of S. propria (Figs 1, 4). Both species form a subclade in Clade D together with S. jeanae and S. diemenensis. From its closest relative, S. propria, this species differs by COI distances of ≥ 10.4 %. From other species, S. australis differs by distances of ≥ 19 % (Table S 8). Within its range S. australis is found in sympatry with two congeners: For comparison with S. obliquata refer to comparative remarks under that species. Siphonaria propria has a slightly smaller shell with a more prominent siphonal ridge, posteriorly left offset apex, regular ribbing and a consistently internal tan to off white siphonal groove, as well as a smaller to inconspicuous AO and longer F 1. Some authors were uncertain of the identity of S. australis. Specimens figured in Hubendick (1946: pl. 3, figs 28 – 31) have been correctly identified as S. australis; however, the corresponding description (Hubendick 1946: 49) incorrectly includes specimens from Port Jackson [Sydney], Australia (likely S. denticulata). Hubendick (1946) incorrectly included Port Jackson and Norfolk Island in the distribution of this species. The radula description matches that of Hutton (1883: 124, pl. 17, figs E – G).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFB2822DFF68F8C2FD09FA76.taxon	distribution	Distribution and habitat. Recorded from New Zealand, North and South Islands, Chatham, Stewart, and Snares Islands, between latitudes of 34 ° and 48 ° S) (Fig. 16). In this study found to be common in sheltered positions (e. g., crevices, vertical faces) on exposed to semi-exposed rocky intertidal shores across upper and mid littoral levels (see also Jenkins 1983: 14).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFAC822EFF68FA42FB65FBD6.taxon	description	(Figs 17 L – N, 18 F – G)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFAC822EFF68FA42FB65FBD6.taxon	materials_examined	Material examined. Type material. Lectotype of Siphonaria capensis Quoy & Gaimard, 1833, present designation, from ‘ Cap de Bonne-Espérance, baie de la Table’ [Table Bay, Cape of Good Hope, South Africa] (MNHN IM 2000 - 38235 Fig. 17 L); Seven paralectotypes, same data as lectotype (MNHN-IM- 2000 - 5042). Other, non-type material. Mozambique: Inhaca, Ponta do Farol, 25 ° 58.2 ’ S, 32 ° 59.4 ’ E MM 6 (MNHN IM- 2019 - 1477 p [M 584], Fig. 17 N; IM- 2019 - 1481 p [M 585], Fig. 17 M).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFAC822EFF68FA42FB65FBD6.taxon	discussion	Taxonomic remarks. The lectotype is designated herein for the stabilisation of the name and to ensure the unambiguous identity of this taxon (Art. 74.1 of the Code). Krauss (1848: 58) introduced Siphonaria capensis var. lineolata; however, this name is preoccupied by S. lineolata Sowerby I, 1835 (Hubendick 1946: 33; White & Dayrat, 2012: 65) and S. lineolata Orbigny, 1841 (White & Dayrat, 2012: 65). Hubendick (1946: 33) introduced S. capensis forma kraussi as a replacement name. The taxonomic status of this taxon remains uncertain, but it is synonymized herein because of Hubendick’s treatment as an infrasubspecific taxon (forma). Hubendick (1946: 33) synonymized S. jonasii Dunker, 1846 (probably a juvenile), S. placentula Menke, 1853, and S. venosa Reeve, 1856 with S. capensis. He also attributed the record of ‘ S. pectinata ’ by Martens (1874) to S. capensis. Our delineation of this species is based on comparative analyses of the morpho-anatomy and mitochondrial genetics of freshly collected topotypes (e. g., Fig. 17 M-N) and geographic series of additional specimens (Tables S 1 – 2). The taxonomic status of S. placentula, S. oculus, S. venosa and S. jonasii has not been assessed herein. Therefore, Hubendick’s (1946) treatment of these taxa stands. Trew (1983: 2) incorrectly treated S. capensis as a synonym of S. funiculata. Chambers & McQuaid (1994 a, b) reported that S. capensis deposits benthic egg masses from which planktonic veliger larvae hatch.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFAC822EFF68FA42FB65FBD6.taxon	description	External morphology (preserved). Animal yellowish / cream. Irregular, widely spaced dark / black markings / spots along foot wall and cephalic lobes; genital pore conspicuous, located on foot wall posterior to right cephalic fold; mantle edge thick, large lobes. Shell (Figs 17 L – N; Table S 9). Small to medium sized, ovate, thick, medium height (max sl mean = 23.1 mm, SD = 3.3 mm, n = 13), exterior grey / light brown, fairly even, apical sides convex, almost straight on posterior; apex offset to left and posterior, often eroded, protoconch direction undetermined, body whorl dextral, growth striae clear with faint external colour banding aligning with growth stages; primary ribs touching, raised fairly evenly spaced, straight, broaden to shell edge, ridges rounded to flat, rib count (mean = 42.6, SD = 5.8, n = 13), few to no secondary ribs; ribs project weakly beyond shell edge, siphonal ridge indistinct, weakly raised, formed by multiple ribs; rib interstices dark brown, weakly furrowed; shell edge weakly corrugated and scalloped. Interior glossy, edge and margin chocolate brown to black at shell edge with white / cream rays aligning under ribs, extending and narrowing from shell edge over margin to ADM scar, spatula pale blue / tan, siphonal groove apparent, same as interior colour, CMS straight. Reproductive system (Fig. 18 F; n = 3). Positioned within coelom under the respiratory cavity and intestine, over foot muscle, against right side of foot wall; epiphallic parts positioned over BM and between RAM; single conspicuous GP, opening from GA through foot wall posterior to right cephalic fold, junction of AO, GA and ED close distinct; AO large broad and flat, larger than GA, thicker than ED; GA bulbous proximal to ED and BD / CD junctions, ED relatively long and twisted; EG very large, broad, lobed, flagellum (F 1) indistinct; BD and CD closely coupled, of similar length and thickness, connect in parallel close together into GA between ED and GP, BD without distal loop or MA; both ducts smooth, featureless, pass together partially through RAM connecting into soft curved folds of MG, CD wider and dorsal to BD; CD connects to small, bulbous, thin test BC, partially embedded into folds of MG; HD short thick, brown markings, coiled, links AG to elongated narrow yellowish granulated HD; HG larger than AG, MG and AG folded, soft white tissue; SV embedded on left side of AG. Spermatophore (Fig. 18 G). Very small, drop-like with short flagellum (length = 0.41 ± 0.037 mm, n = 3); head section bulbous, rounded (head length = 0.31 ± 0.04 mm, head ~ 76 % of SPM length, head width = 230 ± 37 μm; flagellum width = 17 ± 0 μm, n = 3), body and flagellum test fur-like, opaque, tapering to a pointed end; both sections featureless; 4 SPM in brown gelatinous mass in one BC (MNHN IM- 2019 - 1481 [M 585]) embedded in white gelatinous mass. Radula. Dentition formula: 42: 1: 42 (Hubendick 1946: 34).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFAC822EFF68FA42FB65FBD6.taxon	diagnosis	Comparative remarks. Analyses of mtDNA sequences supported a sister relationship of S. capensis (unit 96) with unit 4 (= S. pectinata from Spain and Ghana + S. naufragum from Trinidad and Tabago; not reviewed herein, see Giribet & Kawauchi 2016). Distances between units 96 and 4 were ≥ 10.7 % in COI (Table S 8). We found two sympatric congeners in Mozambique: Siphonaria carbo has a darker shell with a more central apex and finer ribbing, an indistinct AO, larger, elongate BC and longer, larger SPM. Siphonaria plana has a smaller, paler shell with coarser ribbing and a flared siphonal ridge, smaller AO, twisted ED, BD with bursal and distal loops, a larger BC and a longer, thread-like SPM. For comparisons with South African siphonariids, such as S. serrata (Fischer, 1807), S. concinna Sowerby I, 1825, S. oculus Krauss, 1848 and S. carbo Hanley, 1858, which occur within the distributional range of S. capensis, refer to the studies of Allanson (1958), Chambers & McQuaid (1994 a) and Teske et al. (2007, 2011). The RS figured in Hubendick (1945: 18, fig. 20) closely resembles the RS of S. capensis figured herein (Fig. 18 F) except for one detail in the junctions of the CD / BD / GP / GA, which Hubendick portrays as wider and with ducts more separated. Apart from the absence of an AO, the RS of S. capensis figured in Allanson (1958: 165, fig. 10) matches well the RS figured herein (Fig. 18 F). Figured specimens of ‘ S. capensis’ and ‘ S. capensis v. kraussi’ in Hubendick (1946: pl. 1, figs 40 – 42 from Cape, South Africa; pl. 2, figs 1 – 4 from Port Natal, respectively) corresponds well with shells shown herein (Fig. 17 L – N). Similarly, specimens figured in Krauss (1848: 58, pl. 4, fig. 2), Braga (1956: 7, pl. 1 fig. 1), Allanson (1958: 150, 157, pls 1 a – b, 2 a – b), Kilburn & Rippey (1982: 134, pl. 32, fig. 13) and Chambers & McQuaid (1994 a: 256, figs 1 E, 3 A) correspond well with typical features of this species.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFAC822EFF68FA42FB65FBD6.taxon	distribution	Distribution and habitat. Endemic to SE coast of Africa (Fig. 28). During this study, collected on intertidal rocks in Baie de Maputo, Mozambique.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFAF822AFCCAFBE2FECCF7F6.taxon	description	(Figs 19 A – F, M – P, T – U, 21)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFAF822AFCCAFBE2FECCF7F6.taxon	materials_examined	Material examined. Type material. Lectotype of Siphonaria atra Quoy & Gaimard, 1833, present designation, from l’ile de Vanikoro [Vanikoro Island, Solomon Is]; coll. Expédition Durville, 1826 (MNHN IM- 2000 - 35950, Fig. 19 A). Two paralectotypes, same data as lectotype (MNHN IM- 2000 - 5034). Lectotype of Siphonaria albicante Quoy and Gaimard, 1833, present designation, from l’ile de Vanikoro et celle de la Nouvelle Iriande, au havre Carteret [Vanikoro Island, Solomon Is and New Ireland, PNG, at Carteret Harbour; thought to be Carteret Islands, PNG] (MNHN IM 2000 - 38234, Fig. 19 M). Paralectotype, same data as lectotype (MNHN IM 2000 - 5026, Fig. 19 N). Holotype of Triellsiphon acervus Iredale, 1940 from Canala, NC; coll. 1928. (AM C. 103716, Fig. 19 D). Fifteen probable paratypes (AM C. 140260), same data as holotype. Probable holotype of Mestosiphon eumelas Iredale, 1940 from Snapper Island, [near Port Douglas], N Qld [Australia]; coll. 1928 (AM C. 103711, Fig. 19 C). Fifteen probable paratypes (AM C. 140260), same data as probable holotype. Other, non-type material. Solomon Islands: Santa Cruz Ids, Vanikoro off Sunday Reef, 11 ° 42 ’ S, 166 ° 50 ’ E (AM C. 52095 d). NC: Tiari, 20 ° 15.692 ’ S, 164 ° 24.664 ’ E NC 04 - 3 (AM C. 585008 p [M 361]); S of Pouebo, 20 ° 25.950 ’ S, 164 ° 39.251 ’ E NC 04 - 2 (AM C. 585005 p [M 383]); Hienghène, 20 ° 41.210 ’ S, 164 ° 59.108 ’ E NC 04 - 1 (AM C. 585003 p [M 391, SK 173], C. 585004 p [M 392]); Ponerihouen, 21 ° 05.644 ’ S, 165 ° 26.646 ’ E NC 03 - 1 (AM C. 584977 p [SK 059], C. 584978 p [SK 155], C. 584980 p [M 354], C. 584981 p [M 356], C. 584982 p [M 359] C. 584983 p [M 405, SK 072], C. 585329 p [SK 040]); Poindimie, 21 ° 55.901 ’ S, 165 ° 19.672 ’ E NC 03 - 2 (AM C. 585967 20 + p, C. 584993 p [M 351]); Ouassé nr Canala, 21 ° 30.346 ’ S, 166 ° 03.732 ’ E NC 02 - 1 (AM C. 585377 10 + p, C. 584973 p [M 373], C. 584974 p [M 374], C. 584975 p [M 376], M 378, C. 584976 p [M 411]; Bonhomme de Bourail, La Roche Percee, 20 ° 36.487 ’ S, 165 ° 27.423 ’ E NC 06 - 3 (AM C. 585011 p [M 369]); Presqu’ile de Ouano La Foa, 20 ° 51.434 ’ S, 165 ° 48.479 ’ E NC 06 - 4 (AM C. 595912 8 p); Port Quenghi nr marina, 21 ° 54.405 ’ S, 166 ° 03.880 ’ E NC 07 - 1 (AM C. 585018 p [M 379], C. 585019 p [M 380]). PNG: New Ireland, Channel betw. Manne I. and New Ireland mainland, 02 ° 43 ’ S, 150 ° 43.1 ’ E (MNHN IM- 2013 - 55761 p [M 531], IM- 2013 - 55762 p [M 538], IM- 2013 - 55763 p [M 529], IM- 2013 - 55764 p [M 535]; Biliau I., 05 ° 11.8 ’ S, 145 ° 48.2 ’ E PM 38 (MNHN IM- 2013 - 15193 p [M 559]). Australia, Qld: Somerset Bay, Cape York, 10 ° 44.310 ’ S, 142 ° 35.637 ’ E, Q 47 - 2 (AM C. 585177 d [R 12001]); Capt Billy Landing, 11 ° 38.019 ’ S, 142 ° 51.472 ’ E, Q 46 - 1 (AM C. 584791 p [M 006, SK 250]); Lizard Is, 14 ° 40.908 ’ S, 145 ° 27.007 ’ E; Q 40 - 1 (AM C. 584801 4 p); Q 40 - 5 (AM C. 585166 p [SK 218]); Cape Kimberley, 16 ° 16.535 ’ S, 145 ° 28.737 ’ E, Q 35 - 1 (AM C. 585412 10 + p, C. 585167 p [M 454, SK 178], C. 585503 p [SK 179]); Low Isles, 16 ° 23.085 ’ S, 145 ° 33.596 ’ E, Q 34 - 1 (AM C. 585704 4 p, C. 585160 p [M 132], C. 585873 p [M 136]); Port Douglas, 16 ° 28.697 ’ S, 145 ° 27.859 ’ E, Q 33 - 2 (AM C. 585158 p [M 134]); Red Cliff Pt, N of Cairns, 16 ° 41.294 ’ S, 145 ° 35.080 ’ E, Q 33 - 3 (AM C. 595969 3 p); Gribble Pt Mission Bay Yarrabah, 16 ° 53.781 ’ S, 145 ° 51.852 ’ E, Q 32 - 1 (AM C. 585504 2 p); Mourilyan Harbour, 17 ° 35.951 ’ S, 146 ° 07.583 ’ E, Q 29 - 1 (AM C. 585410 8 p, C. 585534 5 p, C. 585151 p [M 004], C. 585152 p [M 401], C. 585153 p [SK 124], C. 585154 p [SK 217]); Horseshoe Bay Magnetic Is, 19 ° 06.755 ’ S, 146 ° 51.875 ’ E, Q 27 - 2 (AM C. 585345, p); Picnic Bay Magnetic Is, 19 ° 10.757 ’ S, 146 ° 50.555 ’ E, Q 27 - 3 (AM C. 585563 9 p, C. 595970 2 p); Mackay breakwater wall, 21 ° 06.415 ’ S, 149 ° 14.033 ’ E, Q 14 - 2 (AM C. 585502 p; C. 585639, 5 p); Wreck Pt Yeppoon, 23 ° 08.736 ’ S, 150 ° 45.865 ’ E, Q 08 - 4 (AM C. 585562 3 p, C. 585132 p [M 080]); Bagara Hervey Bay, 24 ° 49.180 ’ S, 152 ° 28.011 ’ E, Q 06 - 1 (AM C. 585699 8 p); Drury Pt Scarborough, 27 ° 12.255 ’ S, 153 ° 07.022 ’ E, Q 03 - 7 (AM C. 585685 7 p). NT: S Turtle Beach 12 ° 18.852 ’ S, 136 ° 55.959 ’ E NT 04 - 2 (AM C. 585666 6 p, C. 585073 p [M 013], C. 585074 p [M 096]; Cape Wirawawoi Nhulunbuy, 12 ° 09.513 ’ S, 136 ° 46.904 ’ E NT 05 - A (AM C. 585593 4 p); Smith Pt, 11 ° 07.360 ’ S, 132 ° 08.134 ’ E NT 21 - 2 (AM C. 585667 6 p); Nightcliff Darwin, 12 ° 22.836 ’ S, 130 ° 50.402 ’ E NT 23 - 1 (AM C. 585088 d). WA: Tait Point 1 Mission Bay, 14 ° 05.442 ’ S, 126 ° 41.143 ’ E, WA 04 - 1 (AM C. 584716 5 p; WAM S 74089 5 p); Walsh Point, 14 ° 33.998 ’ S, 125 ° 51.060 ’ E, WA 13 - 1 (AM C. 584697 3 p; WAM S 74090 3 p); Caffarelli Is, 16 ° 01.991 ’ S, 123 ° 18.625 ’ E, WA 19 - 1 (AM C. 584717 5 + p, C. 585012 p [SK 034], WAM S 74092 5 p); Conilurus Is, 16 ° 08.872 ’ S, 123 ° 35.205 ’ E, WA 18 - 2 (AM C. 584721 5 p, WAM S 74091 4 p); Catamaran Bay formal, 16 ° 27.622 ’ S, 123 ° 00.242 ’ E, WA 22 - 3 (AM C. 584703 4 p, WAM S 74094 3 p); Catamaran Bay, 16 ° 27.622 ’ S, 123 ° 00.242 ’ E, WA 22 - 1 (AM C. 584737 27 p, C. 585300 p [SK 189], WAM S 74093 5 p); Gantheaume Point, 17 ° 58.384 ’ S, 122 ° 10.677 ’ E, WA 26 - 2 (AM C. 584718 5 p; WAM S 74095 5 p).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFAF822AFCCAFBE2FECCF7F6.taxon	discussion	Taxonomic remarks. The largest syntype is herein designated as the lectotype of Siphonaria atra (MNHN IM- 2000 - 35950) to ensure an unambiguous identity of this species and for the stabilisation of the name (Art. 74.1 of the Code). While topotypic specimens from Vanikoro are currently unavailable, the identity of S. atra (Fig. 19 B) and its junior synonyms Triellsiphon acervus (Fig. 19 F) and S. albicante (Fig. 19 O) are each established based on the examination of specimens from NC that closely match the types. Furthermore, our delineation of this species is also based on comparative analyses of the morpho-anatomy and mitochondrial genetics of freshly collected topotypes of M. eumelas (Fig. 19 P) and Triellsiphon acervus (Fig. 19 E) as well as geographic series of additional specimens (Table S 1). The shells figured as ‘ Siphonaria atra ’ by Reeve (1856: pl. 3, fig. 14) exhibit a mixture of features typical for two species, S. sirius and S. atra. The description in Reeve mentioned features characteristic of S. sirius (i. e., shell pattern and single siphonal rib) but not S. atra. Furthermore, the initial two locations listed in Reeve (1956) are within the known distribution of S. sirius and well outside the known distribution of S. atra. Only Vanikoro, the type locality, is within the range of S. atra. Reeve (1956) incorrectly treated S. coreensis as a synonym of S. atra. In several cases, the details of type locality in the original designations of Iredale (1940), differed from what appeared on the specimen labels. Iredale (1940: 438) specified ‘ type’ in the original descriptions for T. acervus and M. eumelas. The figured specimens of T. acervus (Iredale, 1940: pl. 34, figs 22 – 23; Fig. 19 D herein) and M. eumelas (pl. 34, figs 5 – 6; Fig. 19 C herein) are considered as probable holotypes. Hubendick (1946: 50) listed amongst a number ‘ transitional’ forms in an ‘ Indian-West Pacific form-group’ a ‘ S. zanda <> S. atra ’ form. The figured specimen (Hubendick 1946: 50, pl. 5, figs 3 – 4) is as a specimen of S. atra. Overall, it appears that the interpretation of S. atra in Hubendick (1946: 52) is uncertain. The specimens assigned to this species by him were from Mindanao, the Java Sea, and Osagawara [= Ryukyu, Japan] (Hubendick 1946: pl. 4, figs 5 – 7). None of these localities are within the known distribution of S. atra (Fig. 16) and therefore these are likely specimens of S. alba and S. subatra, respectively. Similarly, ‘ S. atra ’ recorded from Nagasaki in Lischke (1871: 105) and from Okinawa in Kuroda (1960: 43) are likely S. subatra. A record of ‘ S. albicans ’ in Hutton (1880: 36) from NZ is likely based on a misidentification of S. australis. Mestosiphon eumelas has been synonymized with S. atra by Hubendick (1946: 52, 1955: 128), which is followed herein. Cernohorsky (1972: 210) also treated M. eumelas a synonym of S. atra; however, his figure of ‘ S. atra ’ (pl. 60, fig. 1, Naviti Is, Fiji) shows a specimen of S. vudaensis sp. nov. Morrison (1972: 56 – 58) treated S. atra as a synonym of S. laciniosa based on similarity in the shell and for a ‘ common reproductive development’. This synonymy is rejected herein based on examination of type specimens and comparative morpho-anatomy. The unfigured records of ‘ S. atra ’ in Chambers (1980: 141, 143), Christiaens (1980 a: 78) and Chambers, McQuaid & Kirby (1998: 51) from Hong Kong are here attributed to either S. alba or S. sirius.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFAF822AFCCAFBE2FECCF7F6.taxon	description	External morphology (Fig. 19 U). Foot sole, foot wall, foot edge, cephalic folds and pneumostomal lobe evenly cream; mantle thick translucent in larger specimens, thin in smaller specimens, edge thickened, whitish, lobed, with very light black pigmentation at mantle edge aligning with large primary rib interstices; genital pore inconspicuous, located on foot wall to right anterior of right cephalic fold; two small black epithelial eye spots centralised on two centrally touching, centrally black pigmented cephalic folds; pneumostomal lobe long, under the mantle, unpigmented, behind right cephalic fold. Shell (Figs 19 A – F, M – P, T; Table S 9). Shell morphology displays wide intraspecific variability, medium sized (max sl mean = 21.7 mm, SD = 3.3 mm, n = 23), ovate; height low to medium; apex offset sightly posterior and central, apical sides convex, protoconch direction homostrophic (n = 3) shell whorl dextral; growth striae prominent in bands, shell thickness thick; exterior uneven, dark purple brown to pale tan, radial colour banding often present, protoconch area pale, central band darker and shell edge dark brown; rib count (mean = 49, SD = 6, n = 23), primary rib ridges pale, ribs fairly straight, increasingly raised and strongly protrude beyond shell lip (often> 1 mm) to unevenly scallop and corrugate the edge; siphonal ridge often greatest protrusion, formed by paired primary ribs; 1 – 2 finer interspersed secondary ribs between primary, rib interstices darker. Interior shell margin dark chocolate brown to white, white rays on shell margin aligned under primary / secondary ribs dark chocolate brown to white, siphonal groove distinct, often same colour as shell margin, white forms with fringing dark brown markings; spatula dark chocolate brown to golden brown even whitish; ADM scar distinct, CMS straight, darker than shell lip; thickening of shell lip common, overcoats brown markings on shell margin with white, infills and reduces lip scalloping, and often spatula becomes whitened. Reproductive system (Fig. 21; n = 19). HG / AG / MG complex positioned within right side of coelom, against foot wall over foot muscle, under the respiratory cavity; epiphallic parts positioned between BM and RAM; GA small, with singular GP through foot wall; AO large, broad, bluntly pointed, joined to upper GA; ED short, broad, centrally slightly bent, joins to side of GA; GA, AO, ED all white muscular fibrous tissue; EG large, longer than ED, soft whitish tissue, folded, joins ED; single very broad flagellum (F 1), similar length and width to ED, appears as an extension of ED. BD and CD connect side-by-side into GA between ED join and GP, both ducts short, straight, smooth, thickened, whitish, featureless, pass closely together through RAM (BD over CD) into soft white folded tissues of MG; MG / AG complex relatively large; CD connecting to ducts, BC embedded in folds of AG / MG complex close to embedded blackish SV; BD with short distal twisted loop, similar thickness but longer than CD; BC relatively small, bulbous, thin whitish translucent test, HD short, wide, coiled, links ducts in soft white folded tissues of AG to yellowish granulated HG; outer edge of MG lobbed; AG usually larger than HG; genital pore inconspicuous, located on foot wall posterior to right cephalic fold. Spermatophore (Figs 21 C, G). Over half of length comprises a translucent cylindrical head section, tip bluntly rounded, central white core; flagellum very thin, transparent, tapering to a thread-like end; both sections smooth, featureless; test thin, translucent (head length = 10.58 ± 1.73 mm; head width = 188 ± 44 μm n = 2; flagellum width = 33 μm, n = 1), possibly longer as flagellum appears incomplete); very short taper region into the filamentous transparent flagellum; three and 5 SPMs respectively tightly coiled and embedded in brown gelatinous mass in BC of 2 specimens [M 391, M 380]. Radula. Dentition formula: 35: 1: 35 (Hubendick 1946: 52).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFAF822AFCCAFBE2FECCF7F6.taxon	diagnosis	Comparative remarks. Siphonaria atra (atra group, unit 41) forms a clade with unit 40 (S. hienghenensis sp. nov. + unidentified species from PNG and Qld) and S. alba (unit 39) (Figs 1, 2). It differs from these two units by COI distances of ≥ 11.1 % (S. hienghenensis sp. nov.) and ≥ 8.3 % (S. alba) (Table S 3). The maximum distance among sequences in unit 41 (= S. atra) is 12.8 % (Tabe S 2). This is the highest intraspecific genetic divergence observed among all Siphonaria species examined herein. This exceptionally high divergence can partly be attributed to a single GenBank sequence from PNG (UF 332973), which is considered to likely represent a separate, unidentified species. When this sequence is excluded from S. atra, the maximum COI distance within unit 41 is 9 %. Siphonaria alba is indeed morphologically similar; however, it has a shell with a slightly less scalloped edge, a longer ED, larger BC, and longer SPM. Both species occur in sympatry in Timor-Leste and Western Australia. Siphonaria hienghenensis sp. nov. has a paler, grey-brown shell with a weaker scalloped edge, and a smaller AO. Both species are found in sympatry in New Caledonia. Throughout the range of S. atra we found twenty-two congeners within partly sympatric distributions. Eight congeners are sympatric with S. atra in NC: Siphonaria namukaensis sp. nov. has a smaller, paler, slightly taller shell with a weaker scalloped edge, a broader ED, and a longer SPM. Siphonaria normalis has a smaller, paler shell with a less prominent siphonal ridge and a weaker scalloped edge, a smaller AO and larger BC. Siphonaria ouasseensis sp. nov. has a smaller more evenly ribbed shell with a stronger offset apex, weaker scalloped edge, and a smaller AO. Siphonaria caledonica sp. nov. has a smaller shell with a greater offset apex and weaker scalloped edge, a longer ED, and longer SPM. Siphonaria bourailensis sp. nov. has a smaller shell with a greater offset apex and weaker scalloped edge, and a shorter F 1. Siphonaria poindimiensis sp. nov. has a smaller, taller, and darker shell with external patterning, a larger, broader ED, and a longer SPM. Siphonaria viridis has a smaller, taller, and darker shell with external patterning, a larger, broader ED, and longer SPM. Ten congeners are sympatric with S. atra in tropical Qld, NT and WA, Australia (along with S. normalis and S. viridis). Siphonaria costellata sp. nov. has a shell with a greater offset apex, weaker scalloped edge, a smaller AO, longer BC, and shorter BD. Siphonaria denticulata has a larger, paler, taller shell with narrower ribbing and without a scalloped edge, a smaller AO, and longer SPM. Siphonaria gemina sp. nov. has a smaller, paler shell with a weaker scalloped edge, a smaller AO, and a shorter BD. Siphonaria jiigurruensis sp. nov. has a smaller, taller shell with a greater offset apex, patterned exterior, a longer, broader ED, more bulbous AO, and a longer SPM. Siphonaria oblia has a far smaller, fragile and taller shell with a well offset apex, weaker scalloped edge, a longer, wider ED and smaller F 1 (Jenkins, 2018: 278, figs 3 C – D). Siphonaria opposita has a lower shell with paler interior, weaker scalloped edge, a larger BC, and longer ED and BD with bursal loop. Siphonaria restis sp. nov. has paler, taller shell with raised and uneven ribbing, a weaker scalloped edge, a smaller AO, longer ED and larger BC. Siphonaria scabra has a paler, taller shell with a weaker scalloped edge, a smaller AO, longer ED and larger BC. Siphonaria zelandica has a smaller, lower, thinner, paler shell with a weaker scalloped edge, a smaller AO, without distal loop on BD, and a shorter drop-like SPM. Siphonaria forticosta sp. nov., along with S. alba, is sympatric in TL. It has a paler shell with weaker splayed siphonal ridge, a smaller AO, and larger BC. For a comparison with S. javanica found in sympatry in PNG refer to comparative remarks under that species. Wide intraspecific variability occurs in shell morphology of S. atra (particularly characteristics of shell thickness, ribbing and interior colouration) irrespective of shell size or geographical distribution. Variability in shell morphology ranges from a dark (i. e., S atra, S. eumelas) to pale (i. e., S. albicante, T. acervus) forms. The RS anatomy of both forms is consistent, however (Figs 21 A, F dark shell form, Fig. 21 B, pale shell form). Shell geometry, sculpture and colouration of S. atra resemble those of 13 other species in the atra group in the IWP region; S. subatra (unit 38), co-occurring S. forticosta sp. nov. (unit 71), S. hienghenensis sp. nov. (unit 40) and S. bifurcata (unit 45), S. denticulata (unit 33), S. scabra (unit 50), S. pravitas sp. nov. (unit 51), S. crenata (unit 43), S. tenebrae sp. nov. (unit 96), S. vudaensis sp. nov. (unit 37), S. alba (unit 39) and S. sirius (unit 31). However, all these species exhibit differences in the structure of RSs, some have finer shell features and all of them are genetically well-differentiated. Figured specimens of ‘ S. atra’ in Adam & Leloup (1939: 9, pl. 2, figs 3 a, b) match the shell size, sculpture, and ribbing of S. atra, and align with its distributional range and radula dentition count (p. 10, text fig. 2 a, b). Figured specimens of ‘ S. (S.) atra ’ in Hubendick (1946: 91, pl. 4, fig. 5 – 7) from Mindanao are likely specimens of S. alba, not of S. sirius (with a single siphonal rib) or S. atra (with narrower, less protruding primary ribbing). A shell figured as S. atra in Tan & Chou (2000: fig. 117) corresponds with features of S. alba (i. e., shell exterior ribbing, interior colouration). Records of ‘ S. atra ’ in Habe (1962: 96, pl. 44, fig. 17; 1964: 145, pl. 44, fig. 17) from Honshu, subsequently referenced also by Maes (1967: 119) are likely misidentifications of S. subatra. We could not confirm any records of S. atra from Japan or the Philippines. The records of ‘ S. atra ’ in Maes (1967: 119) and Wells (1994: 18) are here attributed to S. alba. Way & Purchon (1981: 321) recorded ‘ S. atra ’ from West Malaysia and Singapore, and Tan & Low (2014: 267) recorded ‘ S. atra ’ from CKI, however, we are unable to confirm the occurrence of S. atra in these regions. Figured specimens of the ‘ atra group, unit 41 ’ in Dayrat et al. (2014) from Lizard Island (fig. 5 W) and PNG (fig. 5 X) are morphologically consistent with S. atra.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFAF822AFCCAFBE2FECCF7F6.taxon	distribution	Distribution and habitat. Recorded from throughout large parts of Tropical IWP including Santa Cruz Islands, American Samoa, Solomons Islands, NC, PNG, northern Australia (Cape York Peninsula through to Kimberley, WA), and Timor-Leste (Fig. 16). In this study found to be common on exposed rocky shores across upper littoral levels (Fig. 19 Q).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFAB8225FCCAF8C2FAF1FD96.taxon	description	(Figs 19 G – L, R, 20 D – H)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFAB8225FCCAF8C2FAF1FD96.taxon	materials_examined	Material examined. Type material. Lectotype of Siphonaria denticulata Quoy & Gaimard, 1833, present designation, from ‘ partie sud de la Nouvelle-Hollande, au port Western, et probablement aussi au port du Roi-Georges’ [southern part of Australia, at Western Port, Vic, Australia, and probably also at King George Harbour; see comments below] (MNHN IM 2000 - 35951, Fig. 19 G). Two paralectotypes, same data as lectotype (MNHN IM 2000 - 5062). Probable holotype of Ellsiphon marza Iredale, 1940 from Keppel Bay [Qld, Australia]; coll. H. Bernhard, 1940 (AM C. 103715, Fig. 19 K). Probable paratypes of Ellsiphon marza Iredale, 1940. Similar data to holotype label (refer Taxonomic remarks) (MV F 13843, 3 d); Tampian Heads, Yeppoon, Keppel Bay; coll. H. Bernhard Dec 1938 (AM C. 58529, 26 d); Four probable paratypes; coll. 1940 (AM C. 106867). Holotype of Siphonaria currumbinensis Hubendick, 1955 from Currumbin [28 ° 07.83 ′ S, 153 ° 29.98 ’ E, Qld, Australia]; coll. R. Endean & R. Kenny, 20 - 6 - 1952 (MV F 15562, Fig. 19 I). Probable paratypes, same data as holotype (MV F 13953, 2 d, 3 p; MV F 13948, 2 d, 3 p). Other, non-type material. Australia; Qld: Wreck Pt Yeppoon, 23 ° 08.736 ’ S, 150 ° 45.865 ’ E, Q 08 - 4 (AM C. 585471 3 p, C. 584743 p [SK 030 protoconch B 7 jaw], C. 585133 p [M 081], C. 585134 p [SK 073]); Double Head Yeppoon, 23 ° 09.908 ’ S, 150 ° 47.638 ’ E, Q 08 - 3 (AM C. 585561 3 p); Bagara Hervey Bay, 24 ° 49.180 ’ S, 152 ° 28.011 ’ E, Q 06 - 1 (AM C. 585961 p); Alexandra Mooloolaba, 26 ° 40.342 ’ S, 153 ° 06.822 ’ E, Q 04 - 1 (AM C. 585383 10 p); Nth Reef Scarborough, 27 ° 11.432 ’ S, 153 ° 06.755 ’ E, Q 03 - 5 (AM C. 585560 4 p); Drury Pt Scarborough, 27 ° 11.836 ’ S, 153 ° 06.955 ’ E, Q 03 - 7 (AM C. 585408 10 + p); Scarborough Pt Scarborough, 27 ° 12.168 ’ S, 153 ° 06.980 ’ E, Q 03 - 8 (AM C. 585638 5 p); Currumbin Point, 28 ° 07.523 ’ S, 153 ° 29.203 ’ E, Q 02 - 2 (AM C. 585536 20 + p, C. 585122 p [M 073], C. 585123 p [M 074], C. 585124 p [M 183], C. 585125 p [M 184]). NSW: Brunswick Heads, 28 ° 32.297 ’ S, 153 ° 33.444 ’ E, NSW 12 - 1 (AM C. 585480 16 p, C. 585066 p [M 077], C. 585067 p [M 079]); Evans River at South Evans Head, 29 ° 6.82 ’ S, 153 ° 25.93 ’ E (AM C. 339263 p); Wooli Wooli River at Wooli, 29 ° 53.27 ’ S, 153 ° 15.98 ’ E (AM C. 339285 4 p; AM C. 339287 2 p); Redbank (Corindi) River at Red Rock, 29 ° 59.03 ’ S, 153 ° 13.55 ’ E (AM C. 339288 3 p); Urunga mangroves on S side of town just beyond footbridge, 30 ° 29.92 ’ S, 153 ° 1.16 ’ E (AM C. 355024 8 p); Camden Haven Inlet, 31 ° 38.37 ’ S, 152 ° 49.63 ’ E (AM C. 339289 p); Fingal Bay nr Port Stephens, 32 ° 44.087 ’ S, 152 ° 10.402 ’ E, NSW 09 - 2 (AMC. 5856837 p), 32 ° 44.990 ’ S, 152 ° 10.481 ’ E, NSW 09 - 1 (AM C. 585065 p [M 151]); Bolton Point Lake Macquarie, 33 ° 0.389 ’ S, 151 ° 36.889 ’ E, NSW 08 - 3 (AM C. 585737 2 p, C. 585058 p [M 452]; C. 585059 p [M 453]); Bateau Bay Beach, 33 ° 23.003 ’ S, 151 ° 29.197 ’ E, NSW 08 - 6 (AM C. 595940 p [SK 520); Crackneck, Bateau Bay, 33 ° 23.316 ’ S, 151 ° 29.139 ’ E, NSW 08 - 5 (AM C. 585063 p [SK 295], C. 585064 p [SK 296]); Broken Head Terrigal, 33 ° 26.796 ’ S, 151 ° 27.030 ’ E, NSW 08 - 1 (AM C. 585528 20 + p, C. 585050 p [M 001]); Terrigal The Skillion, 33 ° 27.008 ’ S, 151 ° 27.122 ’ E, NSW 08 - 2 (AM C. 585558 3 p, C. 585053 p [M 226], C. 585054 p [M 225]); Brisbane, WAter, 33 ° 29.82 ’ S, 151 ° 18.18 ’ E (AM C. 355025 8 p); Fairlight Sydney Harbour, 33 ° 47.971 ’ S, 151 ° 15.990 ’ E, NSW 06 - 4 (AM C. 585590 4 p); Fairlight, North Harbour, 33 ° 47.986 ’ S, 151 ° 16.837 ’ E, NSW 06 - 1 (AM C. 585589 6 p); 100 m NW of Spit Bridge, Port Jackson, 33 ° 48.210 ’ S, 151 ° 14.664 ’ E (AM C. 546770 2 p); Spit Bridge, Port Jackson, 33 ° 48.270 ’ S, 151 ° 14.520 ’ E (AM C. 546768 p); Wy-ar-gine Point Balmoral, 33 ° 49.159 ’ S, 151 ° 15.195 ’ E, NSW 06 - 5 (AM C. 585476 17 p, C. 585043 p [M 158], C. 585044 p [M 159], C. 585045 p [M 160], C. 585046 p [M 161], C. 595923 p [SK 031]); Laings Point Sydney Harbour, 33 ° 50.419 ’ S, 151 ° 16.638 ’ E, NSW 06 - 3 (AM C. 585589 5 p, C. 585034 p [M 193], C. 585041 p [M 165], C. 585042 p [M 167]); Bombo Kiama 34 ° 39.232 ’ S, 150 ° 51.649 ’ E, NSW 03 - 1 (AM C. 585403 10 + p, C. 585333 p [SK 043]); Murunna Point Camel Head, 36 ° 22.720 ’ S, 150 ° 04.766 ’ E, NSW 02 - 1 (AM C. 585401 10 + p). Victoria: Bastion Head Mallacoota, 37 ° 34.429 ’ S, 149 ° 45.927 ’ E, V 09 - 1 (AM C. 585787 5 p, C. 585294 p [M 194], C. 585295 p [M 195]); Cape Conran, 37 ° 48.798 ’ S, 148 ° 43.608 ’ E, V 08 - 2 (AM C. 585292 p [M 197]). Tas: T 01 - 1 S of Granite Point Bridport, 40 ° 59.739 ’ S, 147 ° 23.468 ’ E (AM C. 584834 p [M 173]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFAB8225FCCAF8C2FAF1FD96.taxon	discussion	Taxonomic remarks. The largest syntype with the clearest external sculpture (Fig. 19 G) is herein designated as the lectotype of S. denticulata for the stabilisation of the name and to ensure the unambiguous identity of this taxon (Art. 74.1 of the Code). Our delineation of this species is based on comparative analyses of the morpho-anatomy and mitochondrial genetics of freshly collected topotypes of S. denticulata (AM C. 585303 [SK 043], C. 585044 [M 159], C. 585053 [M 226]), Fig. 19 H), Ellsiphon marza (AM C. 584743 [SK 030], C. 585133 [M 081], Fig. 19 L) and S. currumbinensis (AM C. 585123 [M 074], Fig. 19 J) and geographic series of additional specimens (Table S 1). Quoy & Gaimard’s (1833: 340) statement ‘ probably also at King George Harbour’ is not considered as part of the type locality. While there is also a King George Harbour in Tas, the King George Harbour referred to by Quoy & Gaimard (1833) is probably that near Albany, WA. However, we have not found this species in WA. The type locality for E. marza is Keppel Bay and Caloundra as stated in the original description, yet some paratypes are from additional sites (Tampian Heads and Yeppoon). Noosa, Qld is mentioned by Hubendick as a site for ‘ S. currumbinensis ’ The locality on the holotype label (MV F 15562) is misspelled as Corrumbin (sic Currumbin) and the name is misspelled as S. corrumbinensis. Hutton (1873: 55) stated that S. denticulata and S. diemenensis occur in NZ; however, this is likely based on misidentifications of S. australis and / or S. propria.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFAB8225FCCAF8C2FAF1FD96.taxon	description	External morphology (Fig. 19 R). Foot sole smooth, evenly pale yellow grey; foot wall pale yellow grey with evenly spread white subepithelial pustules becoming more vivid and dense around pneumostomal lobe; foot wall shows black blotches; fringing mantle translucent, covers shell mantle, outer edge lobed, thickened, with black pigmented blotches aligning with rib interstices; pneumostomallobewithinmantlebetweentherightADMs, closes the pneumostomal and anal openings at the mantle edge; two small black epithelial eye spots centralised on two thick brownish yellow centrally touching cephalic folds; genital pore inconspicuous, located on foot wall to right anterior of right cephalic fold. Shell (Figs 19 G – L; Table S 9). Medium to large sized (max sl mean = 24.46 mm, SD = 2.04 mm, n = 20), ovate; height medium; apex offset sightly posterior and central (usually eroded), apical sides strongly convex, protoconch direction undetermined, shell whorl dextral; growth striae prominent in bands, shell thickness thick; rib count (mean = 42.6, SD = 4.5, n = 20), primary ribs pale white, fairly straight, increasingly raised and protrude beyond shell lip to unevenly scallop and corrugate the edge; 1 – 2 interspersed pale white finer secondary ribs, rib interstices darker; paired primary ribs on siphonal ridge, no more prominent than other primary ribs. Interior shell margin dark brown to tan, white rays align on shell margin under primary / secondary ribs, siphonal groove distinct, same colour as shell edge, points to right anterior; spatula dark chocolate brown to mottled tan even whitish; ADM scar distinct, CMS straight, paler than shell lip; thickening of shell lip translucent, infills and reduces lip scalloping, spatula becomes whitened. The shells of S. denticulata, S. scabra, S. pravitas sp. nov. and S. diemenensis may appear similar and may be misidentified; external ribbing is similar. Differences exist in the sharpness and intensity of primary ribs and rib interstice colouration. In addition, the spatula colouration differs; S. denticulata and S. scabra dark chocolate to tan-brown, S. diemenensis and S. pravitas sp. nov. are more golden brown. Reproductive system (Figs 20 D, F, G; n = 3). Positioned within coelom under the respiratory cavity, over foot muscle, against right side of foot wall; epiphallic parts positioned between BM and RAM; single GP inconspicuous, opening from GA through foot wall posterior to right cephalic fold. Join of AO, GA and ED distinct, all whitish fibrous muscular tissue; AO elongated, bluntly pointed, folded, larger than GA, thicker than ED; ED relatively long, thick and twisted; EG broad, lobed, bluntly pointed with two curved, broad, flagellum (F 1 and F 2; F 2 twice length of F 1); BD and CD connect close together into GA between ED, AO and GP; BD noticeably longer and thinner than CD with a prominent twisted loop immediately before BC (no distal loop or MA), both ducts smooth, featureless, pass together through RAM connecting into soft curved folds of MG, BD dorsal to CD; BC large, bulbous, thin test, internally embedded into folds of MG; HD short thick, coiled, links AG to elongated narrow yellowish granulated HD; HG larger than AG, MG and AG folded, soft white tissue; SV embedded on left side of AG under BC; AG and HG sides match curvature of inner foot wall at right posterior quarter of coelom. Spermatophore (Figs 20 E, H). Head section over half of total length (head length = 8.08 ± 0.42 mm; ~ 43 % of SPM length; head width = 119 ± 3 μm; flagellum width = 33 ± 1 μm, n = 3), cylindrically elongate, tip bluntly rounded, containing a white gelatinous mass; tapers quickly into consistently filamentous transparent flagellum; both sections smooth, translucent, test thin, featureless. multiple tightly coiled SPM in bursa, embedded in brown gelatinous mass. Seven and two SPM found in BC of two specimens [SK 043, SK 030]. Radula and jaw (Figs 83 E – H). Mean dentition formula 31: 1: 31 (SD = 8.1), mean transverse rows 138 (SD = 20.1, n = 3, AM C. 201775, C. 201765, C. 316303); single central rachidian tooth flanked squarely by 31 half row laterals, 0 – 4 are inner (SD = 4.98), 12 – 14 mid (SD = 4.98) and 15 – 19 outer (SD = 3.58) laterals (n = 5); central tooth with narrow unicuspid mesocone less than half length of basal plate, lower than laterals; inner laterals absent or few (Figs 83 E – F), mid lateral mesocones generally unicuspid, however, maybe irregularly bicuspidate (Figs 83 E – F), mid laterals with pointed ectocone; ectocone strong, protrudes at acute angle halfway along the tooth’s length; outer laterals typically with a ‘ chisel’ shaped mesocone flanked by small, pointed single ecto and endocones, endocone positioned higher on side of tooth than ectocone, angle of separation of each cone from the mesocone varies (Fig. 83 G). Jaw located inside front of buccal cavity, orange-brown, arch shaped with ‘ shingle’ - arranged cone-like rods, ~ 140 rods wide (~ 1.9 mm) by ~ 16 rods deep (mean length = 50 μm, SD = 6, mean width = 12 μm, SD = 3, n = 15); tip bluntly rounded [SK 030].	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFAB8225FCCAF8C2FAF1FD96.taxon	diagnosis	Comparative remarks. Siphonaria denticulata (atra group, unit 33) is the sister taxon of a clade containing two species, S. opposita (unit 34) and S. plana (unit 35) (Figs 1 – 4). It differs from these species by COI distances of ≥ 14.8 % (unit 35) and ≥ 15.8 % (unit 34) (Table S 3). We found eight congeners with sympatric occurrences in SE Australia. For comparisons with S. atra and S. diemenensis refer to comparative remarks under these species. Siphonaria stowae has a smaller, more elongate, paler shell with a strongly offset apex, less prominent ribbing and edge scalloping, a smaller AO, shorter ED larger BC, and shorter SPM. Siphonaria emergens has a smaller, elongate, paler brown shell with a strongly offset apex, less prominent ribbing and edge scalloping. Siphonaria funiculata has a paler, taller shell with less raised ribs broadening to shell edge, a fainter scalloped edge, a larger AO, shorter BD and ED, absent to smaller F 1, and larger, drop-like SPM. Siphonaria pravitas sp. nov. has a lower shell with greater raised ribbing and edge scalloping, no BD distal loop, a larger BC, wider F 1, and shorter SPM. Siphonaria scabra has a paler shell with finer ribbing and coarser exterior, a longer AO, BC and F 1, and a bursal loop. Siphonaria zelandica has a paler, lower shell with less prominent and finer narrower ribbing, a shorter ED and BD, and a short. drop-like SPM. Menke (1851: 38) referred to the name S. denticulata for a specimen from Mexico. This reference was subsequently considered as a misidentification of S. lecanium var. palmata Carpenter, 1857 (Carpenter 1857: 183). Hubendick (1946: 40) treated this taxon as a junior synonym and subspecies of S. maura Sowerby I, 1835. Erroneous records of ‘ S. denticulata ’ from NZ (Hutton, 1873: 55) and Mozambique (Braga, 1956: 7, pl. 1, fig. 2) are herein attributed to S. propria Jenkins, 1983 and possibly S. ferruginea Reeve, 1856, respectively. We have not located any samples of this species from NZ nor Mozambique in any of the examined museum collections. The treatment of S. denticulata as a synonym or variety of S. diemenensis by several authors is refuted herein and both species are accepted as distinct species. Hedley (1915; 1918; M 96) and Angas (1867: 232) incorrectly recorded S. denticulata from Port Jackson (= Sydney) and NSW. A specimen figured as ‘ S. marza’ in Hubendick (1946: 61, pl. 5, figs 27 – 28) from Port Jackson [Sydney Harbour]) is herein identified as S. denticulata. McAlpine (1952: 40, 42, fig. 1) misidentified specimens of S. scabra as ‘ E llsiphen ’ (sic Ellsiphon) denticulatus. The RS figured herein (Figs 20 D, G) matches closely the figure in McAlpine (1952: 43, fig. 1; as Ellsiphon denticulatus) and that shown in (Fig. 20 F, from Currumbin) closely matches the distal RS figure of ‘ S. currumbinensis ’ from in Hubendick (1955: fig. 4). A specimen from NZ labelled as ‘ S. denticulata ’ in Davey (1998: 117) is herein identified as S. funiculata. Specimens stated to resemble either S. diemenensis, S. denticulata or S. australis by Dayrat et al. (2014) as ‘ unit 7 ’ are herein identified as S. diemenensis. By contrast, the figured specimens of ‘ atra group, unit 33 ’ in Dayrat et al. (2014: fig. 5 J, K) are morphologically consistent with S. denticulata.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFAB8225FCCAF8C2FAF1FD96.taxon	distribution	Distribution and habitat. Endemic to temperate eastern and southeastern Australia, from Keppel Bay, Qld, south to Cape Conran, Vic, and Bridport, Tas (Fig. 16). In this study found in sheltered to moderately exposed sites on hard substrates (e. g., rocky shores, wooden pilings); upper and mid littoral levels; home scars prominent.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFA48223FCCAFD22FBCAFBF6.taxon	description	(Figs 22, 23, 24)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFA48223FCCAFD22FBCAFBF6.taxon	materials_examined	Material examined. Type material. Lectotype of Siphonaria viridis Quoy & Gaimard, 1833, present designation, from ‘ rade d’Amboine, aussi la Nouvelle-Guinée’ [on rocks, Amboina] (MNHN-IM- 2000 - 35958; Fig. 22 A). Three paralectotypes MNHN-IM- 2000 - 5042, same data as lectotype. Holotype of Parellsiphon zanda Iredale, 1940 AM C. l 03706 (from GBR and Michaelmas Cay, N Qld, [Australia]; coll. May 1926; Fig. 22 B). Lectotype of Siphonaria punctata Quoy & Gaimard, 1833, present designation, from Île-de-France [Mauritius, Indian Ocean] (MNHN-IM- 2000 - 35957, Fig. 23 A). Two paralectotypes, same data as lectotype (MNHN-IM- 2000 - 5053). Syntype of Parellisiphon promptus Iredale 1940 AM C. 103707 (from North West Island [23 ° 18 ’ S 151 ° 42 ’ E], Capricorn Group, [Qld, Australia]; coll. May 1931; Fig. 22 C). Holotype of Legosiphon densatus Iredale, 1940 AM C. 103716 (from Port Douglas, N Qld, [Australia]. Holotype of Legosiphon optivus Iredale, 1940 AM C. 103719 (from Magnetic Island, Qld, [Australia]; coll. A. F. Basset Hull, May 1924; Fig. 22 H). Seventeen probable paratypes of Legosiphon optivus Iredale, 1940 AM C. 153469, same data. Syntype of Legosiphon mirificus Iredale, 1940 AM C. 103718 (from Magnetic Island, near Townsville, Qld, [Australia]; coll. A. F. Basset Hull, May 1924; Fig. 22 G). Five probable paratypes of Legosiphon mirificus Iredale, 1940 same data as probable holotype, AM C. 153362). Probable holotype of Legosiphon densatus Iredale, 1940 from Port Douglas, N Qld, [Australia]; coll. Nov 1929, (AM C. 103717, Fig. 22 D). Two questionable paratypes, same locality data, Nov 1928 (AM C. 106998). Other, non-type material. Mauritius: Trou-auxBiches, 20 ° 02.488 ’ S, 57 ° 32.353 ’ E, MRU 04 - 1 (AM C. 585972 20 + p, C. 585834 p [M 246], C. 585835 p [M 247], C. 585836 p [M 248]). PNG: New Ireland, NW side of Big Nusa Island, 02 ° 34.1 ’ S, 150 ° 46.7 ’ E KM 12 (MNHN IM- 2013 - 55336 p [M 534]); Lemus Island, 02 ° 38 ’ S, 150 ° 37.5 ’ E KM 24 (IM- 2013 - 53728 p [M 533]; IM- 2013 - 53874 p [M 530]); Ta Island, 02 ° 41.6 ’ S, 150 ° 44.3 ’ E KM 22 (MNHN IM- 2013 - 51004 p [M 537]; IM- 2013 - 55767 p [M 536]; IM- 2013 - 55768 p [M 539]; IM- 2013 - 55765 p [M 540]; IM- 2013 - 55766 p [M 541]; IM- 2013 - 55770 p [M 542]; IM- 2013 - 55769 p [M 543]; IM- 2013 - 50993 p [M 544]); Rempi Area, S Dumduman Is., 05 ° 00,2 ’ S, 145 ° 47,6 ’ E PM 12 (MNHN IM- 2013 - 11999 p [M 561]); Riwo waters, 05 ° 08.9 ’ S, 145 ° 48.2 ’ E PM 40 (MNHN IM- 2013 - 15249 p [M 553]); Biliau Island, 05 ° 11.8 ’ S, 145 ° 48.2 ’ E, PM 38 (MNHN IM- 2013 - 14828 p [M 549], PM 39 (IM- 2013 - 15190 p [M 556]). Australia, Qld: Cape York Peninsula, Bamaga, Umagico, 10 ° 53.125 ’ S, 142 ° 20.799 ’ E, Q 50 - 1 (AM C. 585181 [M 179], C. 585599 4 p, C. 585180 p [M 104], C. 585182 p [M 433, SK 140 (RS), C. 595938 p [SK 539], C. 595939 p [SK 540], C. 585184 p [SK 352], C. 585185 p [SK 353]). Bathurst Head, 14 ° 15.912 ’ S, 144 ° 11.598 ’ E, Q 41 - 2 (AM C. 585973 10 p); S of Bathurst Head, 14 ° 17.583 ’ S, 144 ° 11.845 ’ E, Q 41 - 1 (AM C. 585948 4 p); Lizard Island, 14 ° 40.908 ’ S, 145 ° 27.007 ’ E, Q 40 - 1 (AM C. 585642 5 p); Cape Kimberley, 16 ° 16.535 ’ S, 145 ° 28.737 ’ E, Q 35 - 1 (AM C. 585903 2 p, C. 585173 p [M 084], C. 585174 p [M 085]); Low Isles, 16 ° 23.085 ’ S, 145 ° 33.596 ’ E, Q 34 - 1 (AM C. 585384 10 p, C. 585161 p [M 137], C. 585162 p [M 138], C. 585163 p [M 139], C. 585165 p [M 432, SK 137]); Port Douglas, 16 ° 28.697 ’ S, 145 ° 27.859 ’ E, Q 33 - 2 (AM C. 585962 6 p, C. 585963 5 p, C. 585157 p [SK 217], C. 585159 p [SK 219], C. 585871 p [M 131, SK 218], C. 585872 p [M 133]); Bingil Bay, E of El Arish, 17 ° 49.563 ’ S 146 ° 06.131 ’ E (AM C. 420154 2 d). Magnetic Is., Balding Bay, 19 ° 06.651 ’ S, 146 ° 52.260 ’ E, Q 27 - 1 (AM C. 585904 15 p), Horseshoe Bay, 19 ° 06.755 ’ S, 146 ° 51.875 ’ E, Q 27 - 2 (AM C. 585900 6 p), Geoffrey Bay, 19 ° 09.142 ’ S, 146 ° 52.125 ’ E, Q 27 - 4 (AM C. 585902 7 p C. 585533 d [R 4108], C. 585869 p [M 482], C. 585870 p [M 483]); Rocky Bay, 19 ° 10.43 ’ S, 146 ° 50.73 ’ E (AM C. 459332 1 d). Hamilton Is., Catseye Bay, 20 ° 20.831 ’ S, 148 ° 58.002 ’ E, Q 18 - 1 (AM C. 585463 15 p). NT: N Turtle Beach, 12 ° 18.816 ’ S, 136 ° 55.930 ’ E NT 04 - 1 (AM C. 585069 p [M 086], C. 585070 p [M 148], C. 585071 p [M 149], C. 585072 p [SK 079], C. 585079 p [SK 077]). Nhulunbuy, Cape Wirawawoi, 12 ° 09.513 ’ S, 136 ° 46.904 ’ E NT 05 - 1 A (AM C. 585594 4 p), East Woody Islet 12 ° 09.695 ’ S, 136 ° 45.075 ’ E NT 05 - 2 (AM C. 585632 5 p, C. 585078 p [M 462, SK 196]). Cobourg Peninsula, Kuper Pt 3, 11 ° 10.688 ’ S, 132 ° 13.711 ’ E NT 21 - 7 (AM C. 585390 10 + p, C. 585086 p [M 093], C. 585087 p [M 142]), Kuper Pt 2, 11 ° 10.877 ’ S, 132 ° 13.554 ’ E NT 21 - 6 (AM C. 585084 d [M 035], C. 585085 p [M 150]), Smith Pt, 11 ° 07.360 ’ S, 132 ° 08.134 ’ E NT 21 - 2 (AM C. 585634 5 p), Smith Pt 2, 11 ° 07.466 ’ S, 132 ° 08.538 ’ E NT 21 - 3 (AM C. 585081 p [M 463, SK 198], C. 585317 p [M 034]); Sandy Is Pt, 11 ° 07.862 ’ S, 132 ° 11.187 ’ E NT 21 - 1 (AM C. 585468 15 + p, C. 585080 p [M 023], C. 585842 p [M 095]). Cox Peninsula, 12 ° 24.824 ’ S, 130 ° 40.921 ’ E NT 25 - 1 (AM C. 585407 10 + p); Luxmore Hd, Melville Is, 11 ° 20.639 ’ S, 130 ° 23.149 ’ E NT 24 - 1 (AM C. 585595 4 p). WA: Caffarelli Is, 16 ° 01.991 ’ S, 123 ° 18.625 ’ E, WA 19 - 1 (AM C. 584739 10 + p, C. 584659 p [M 465, SK 200], C. 584660 p [M 466, SK 201], C. 584661 p [SK 290], C. 584662 p [SK 291], C. 584784 p [SK 221], C. 585297 p [SK 220], C. 585298 p [SK 360]); Conilurus Is, 16 ° 08.875 ’ S, 123 ° 35.234 ’ E, WA 18 - 1 (AM C. 585654 5 p); Catamaran Bay, 16 ° 27.622 ’ S, 123 ° 00.242 ’ E, WA 22 - 3 (AM C. 584663 p [M 026]); Emeriau Point Middle Lagoon, 16 ° 46.351 ’ S, 122 ° 34.200 ’ E, WA 23 - 1 (AM C. 584674 d [M 396]; C. 585884 d [M 393]); Broome, between Crab Creek and Broome, 17 ° 58 ’ S, 122 ° 14 ’ E (WAM S 76939 11 p, p [SK 065]); Cape Latouche Treville nr Gourdon Bay, 18 ° 27.101 ’ S, 121 ° 48.911 ’ E, WA 27 - 2 (AM C. 585795 20 p, C. 584675 p [SK 149], C. 585306 p [M 066], C. 585307 p [M 067]); Hearson Cove, Dampier, 20 ° 38.079 ’ S, 116 ° 48.031 ’ E, WA 33 - 2 (AM C. 585309 p [SK 064]); Gnoorea Point 2 20 ° 50.333 ’ S, 116 ° 20.825 ’ E, WA 36 - 2 (AM C. 585885 d (M 105 )); Tantabiddi, 21 ° 54.739 ’ S, 113 ° 58.706 ’ E, WA 42 - 1 (AM C. 584676 p [M 205]; C. 585886 p [M 206]). Timor-Leste: Dolokoan Beach, N of Dili, 8 ° 31.424 ’ S, 125 ° 37.091 ’ E TL 01 - 1 (AM C. 585745 10 + p, C. 584796 p [M 478, SK 285], C. 584821 p [SK 299], C. 584822 p [SK 300], C. 585388 p [SK 558], C. 585880 p [M 442], C. 585881 p [M 443], C. 585882 p [M 444], C. 585883 p [M 450], C. 595948 p [SK 563]). Indonesia, Bali: Tanah Lot IND 03 8 ° 37.25967 ′ S, 115 ° 5.22967 ’ E (ZRC. MOL. 24913 p [M 473]). Singapore: St Johns Island, Peninsula Lokos 01 ° 12.824 ’ N, 103 ° 51.076 ’ E SI 04 - 1 (AM C. 585224 p [M 324], C. 585225 p [M 325]); Lazarus Island channel headland, 1 ° 13.085 ’ N, 103 ° 51.429 ’ E SI 04 - 4 (AM C. 585604 10 p, C. 585164 p [SK 166], C. 585243 p [M 330], C. 585244 p [SK 167], C. 585245 p [M 329], C. 585246 p [M 331], C. 585486 p [M 336]); Lazarus Island causeway, 01 ° 13.288, 103 ° 51.195 ’ E SI 04 - 3 (AM C. 585982 4 p, C. 585240 p [M 328], C. 585241 p [M 333], C. 585242 p [SK 143]); Lazarus Island, 01 ° 13.355 ’ N, 103 ° 51.148 ’ E SI 04 - 2 (AM C. 585228 p [SK 184]. Malaysia: Palau Tioman 2 ° 49 ′ N, 104 ° 11 ′ E ZRC 1999 - 1781 7 p, ZRC. MOL. 24897 p [SK 374], ZRC. MOL. 24898 p [SK 373]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFA48223FCCAFD22FBCAFBF6.taxon	discussion	Taxonomic remarks. The lectotype of S. viridis is designated herein for the stabilisation of the name and to ensure the unambiguous identity of this taxon (Art. 74.1 of the Code). In addition, we designate the syntype of S. punctata with the clearest external sculpture (Fig. 23 A) as the lectotype of S. punctata for the stabilisation of the name (Art. 74.1 of the Code). The labels associated with the type specimens give ‘ Île-de-France’ [Mauritius]. Paetel (1889: 429) listed S. punctata ‘ Quoy. I. Maurit. ’ as an accepted species. However, Reeve (1856), Hanley (1858 b: 152) and Hutton (1880: 36) treated S. punctata as a synonym of S. sipho. Finally, Hubendick (1946: 47) listed S. punctata as a synonym of S. laciniosa. We recognise S. punctata as a synonym of S. viridis mainly based on genetic evidence. We found that topotypic specimens that closely match the shell morphology of the types fall within unit 25 (Fig. 3: genetic vouchers M 246 – M 248 from Mauritius) and exhibit morphological and anatomical features that are consistent with that of S. viridis. The type locality of P. zanda given on the specimen label of the types (‘ GBR and Michaelmas Cay, N Qld’ [Australia]) differs from that given in the original description (‘ Low Isles, N Qld’). The original descriptions of P. zanda and P. promptus each appear to be based on a single specimen. Iredale (1940: 438) referred to ‘ type’ in the original descriptions of P. zanda, L. densatus and L. optivus but not in that of L. mirificus nor P. promptus. Therefore, we consider the single type specimens of P. zanda, L. densatus, and L. optivus as holotypes, but the types of L. mirificus and P. promptus as syntypes. No additional types are known to exist. Hubendick (1946: 47) incorrectly considered S. viridis, L. optivus and L. densatus as synonyms of S. laciniosa based on shell colouration and shape. The interpretation of S. zanda in Hubendick (1946: 52) is likely based on a misidentification. While reference to the original designation is correct, the specimens examined and figured are from the Java Sea and Madagascar (Hubendick 1946: pl. 4, figs 8 – 11) and clearly differ from the type (Fig. 22 B) (refer to Comparative remarks). Hubendick (1946: 53) incorrectly considered L. mirificus as a possible synonym of ‘ S. ferruginea’ (= S. plana). The specimens figured in Hubendick (1946: pl. 4, fig. 16 – 19) do not resemble the type of L. mirificus. Morrison (1972: 56 – 58) synonymized ‘ Parellsiphon zanda ’ and ‘ P. promptus ’, with S. laciniosa based on similarities in shell shape and ‘ common reproductive development’. These synonymies are rejected herein based on examination of type specimens and comparative morpho-anatomy. Our delineation of this species is based on comparative analyses of the morpho-anatomy and mitochondrial genetics of freshly collected topotypes of all synonymized taxa and geographic series of additional specimens (Tables S 1 – 2).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFA48223FCCAFD22FBCAFBF6.taxon	description	External morphology (Figs 22 P, 23 D). Foot sole, foot wall, pneumostome lobe and cephalic folds evenly grey or cream in colour, darker to centre of foot sole, paler to foot edge, cephalic folds cream to mouth; irregular black blotches cover all but foot sole, concentrated to centre of cephalic folds; mantle narrow than foot wall, evenly grey, edge thickened, with white band, lobed; pneumostome lobe large, under mantle. Shell (Figs 22 – 23; Table S 9). Small to medium sized (max sl mean = 13.8 mm, SD = 3.6 mm, n = 21), elongate ovate; height tall; medium thickness; apex offset weakly posterior and left (frequently eroded), apical sides weakly concave, protoconch direction homostrophic (n = 6; Fig. 22 T, W), shell whorl dextral; growth striae prominent, irregular, uneven; rib count (mean = 37, SD = 7.6, n = 21), primary ribs prominent, white, protrude slightly beyond shell edge, often wider than rib interstices, ridge edge rounded, fused pair highlight siphonal ridge; few secondary ribs, 0 – 3 secondary ribs between anterior primary ribs, dark brown / black radial bands in rib interstices overlaying interstices and secondary ribs; shell edge fragile, weakly scalloped and corrugated at primary rib end. Interior shell lip to margin dark brown and white, white rays align on shell margin under furrowed primary / secondary ribs, dual brown rays aligning under interstices extend from lip over margin to merge into a single brown ray to evenly brown spatula; siphonal groove distinct furrow; ADM scar distinct, CMS straight. Growth thickening and whitening of the shell lip, with retention of axial brown markings, is common in larger specimens. The holotype specimen is an example of shell thickening. Reproductive system (Fig. 24; n = 21). Positioned within right side of coelom, against foot wall on foot muscle, under the respiratory cavity; epiphallic parts positioned between BM and RAM. GA small, with singular GP through foot wall; AO very large, wide, bluntly pointed, joined to upper GA, muscle attachment at side; ED elongated, thickened, centrally bent, joins to lower side of GA; GA, AO, ED all white muscular fibrous tissue; EG large, soft whitish tissue, slightly folded, joins ED; single narrow flagellum (F 1), similar length and width to ED, appears as an extension of ED. BD and CD connect in opposing directions into GA between ED join and GP, CD join is bulbous, both ducts long straight smooth whitish, pass together on outer side of RAM (BD over thicker CD) into soft white folded tissues of MG; BD long narrow with prominent distal loop with MA to inner anterior foot wall, bent and twisted before BC; CD long, wider than BD; CD connecting to MG ducts, BC embedded in folds close to embedded blackish SV; BC relatively small bulbous, thin whitish translucent test, 2 SPM in BC of topotypic specimen (AM C. 585165 Q 34 - 1 [M 432, SK 137]); MG / AG complex relatively large; HD short narrow coiled, links ducts in soft white folded tissues of AG to yellowish granulated HG; AG / MG larger than HG, sides match curvature of inner foot wall. Spermatophore (Figs 24 B, D, F, I). Thread-like; head section (head length = 9.74 mm, head width = 94.6 μm, n = 1, flagellum incomplete) cylindrical, bulbous, bent, rounded tip, tapers to flagellum; test thin, smooth, featureless, translucent encasing a white opaque central core; a short tapering section merges head to filamentous flagellum; head section wider than translucent flagellum (complete flagellum unavailable). We found two SPMs tightly coiled in brown gelatinous mass in bursa [M 432, SK 137], two SPM coiled, embedded in white gelatinous mass in each [SK 079, SK 077], a single SPM in white gelatinous mass in one bursa [SK 149]. Radula. Dentition formula: 20 - 25: 1: 25 - 20 (Adam & Leloup, 1939: 8, as ‘ S. sipho ’), 40: 1: 40 (Hubendick 1946: 47).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFA48223FCCAFD22FBCAFBF6.taxon	diagnosis	Comparativeremarks. Inourmolecularphylogenetic tree (Figs 1 – 4), Siphonaria viridis (laciniosa group, unit 25) is closely related to S. umagicoensis sp. nov. and S. punctata. These species are separated from each other by average genetic p-distances in COI of 6.5 % to 8.1 %. All three species together form a well-differentiated clade. Sequences from Sulawesi and Bali clustering within this clade and attributed to “ unit 25 ” by Dayrat et al. (2014) are of uncertain identity and require further scrutiny for lack of morpho-anatomical study. Siphonaria viridis has a variable shell (Figs 22 – 23) across its wide distribution throughout tropical Australia, PNG and Singapore (Fig. 25). Within this range, we found twenty-three sympatric congeners. Three congeners are sympatric in NC: Siphonaria hienghenensis sp. nov. has a slightly larger, darker shell with a flared siphonal ridge, a smaller AO, and shorter SPM. Siphonaria poindimiensis sp. nov. has a taller, darker shell with fused siphonal ribs, stronger edge scalloping, and longer ED. Siphonaria bourailensis sp. nov. has a shell with a more posteriorly offset apex, uneven ribbing, stronger edge scalloping, a smaller AO, and shorter SPM. Nine congeners are sympatric with S. viridis in PNG and TL: Siphonaria atra has a larger, lower shell with stronger edge scalloping, flared siphonal ridge, and larger BC, and shorter SPM. Siphonaria opposita has a lower, pale to dark brown shell, a larger BC, pointed AO, and longer ED. Siphonaria madangensis sp. nov. has a slightly smaller and lower shell with greater raised ribbing and stronger edge scalloping, a smaller, less prominent AO, larger BC, and shorter, bulbous, threadlike SPM. Siphonaria nusalikensis sp. nov. differs by being much smaller shell, with a darker interior, and BD without distal loop. Siphonaria planucosta sp. nov. has smaller, lower, darker shell with unpatterned interstices, a smaller AO, larger BC, and shorter SPM. Siphonaria forticosta sp. nov. has a larger shell with darker spatula, and a smaller AO. Siphonaria alba has larger, lower shell with stronger edge scalloping, a flared siphonal ridge, and a longer SPM. Siphonaria campestra sp. nov. has smaller shell without exterior patterning, with darker interior, a smaller AO, shorter BD and SPM. For comparison with S. javanica refer to comparative remarks under that species. Siphonaria maloensis sp. nov. (sympatric in Vanuatu) is of similar shell size but has a darker, grey-brown shell with slightly stronger edge scalloping, an apex offset to posterior and left, and a golden-brown interior. A further eight species are sympatric with S. viridis in tropical Australia (along with S. alba, S. atra, S. campestra sp. nov. and S. opposita). Siphonaria costellata sp. nov. has a larger, darker shell with a darker interior, a smaller AO, shorter BD, and shorter, wider SPM. Siphonaria gemina sp. nov. has paler shell with stronger edge scalloping and flared siphonal ridge, a smaller AO, and shorter SPM. Siphonaria jiigurruensis sp. nov. has taller shell with more posteriorly offset apex, and paler whitish brown spatula, and a broader ED. Siphonaria oblia has a far smaller shell, weakly ribbed, with strongly posteriorly offset apex, and a smaller to indistinct AO (Jenkins 2018: 278, figs 3 C – D). Siphonaria restis sp. nov. has a shell with stronger edge scalloping, more strongly raised ribbing, a smaller AO, and larger BC. Siphonaria zelandica has lower, paler shell with finer ribbing and a less prominent siphonal ridge, smaller AO and BC, and a smaller, drop-like SPM. Two species are sympatric with S. viridis in Singapore (including S. alba and S. costellata sp. nov.). Siphonaria sirius has lower shell with single rib siphonal ridge, stronger edge scalloping and a paler spatula, a smaller AO, longer ED, shorter BD, larger BC, and a longer SPM. Siphonaria normalis has a smaller, lower shell with a more posteriorly offset apex and less raised ribbing, a smaller AO, larger BC and shorter F 1. Topotypic specimens of L. mirificus (AM C. 585869), L. optivus (Fig. 22 I) and P. densatus (Fig. 22 E) are conspecific based on mitochondrial phylogenetics and comparative morphology. The record of ‘ S. sipho’ in Adam & Leloup (1939: 7, pl. 2, fig. 1 a – d) from Java, Bali, Mansinan, PNG and of ‘ S. funiculata ’ (9, pl. 2, figs 2 a, b) from Iles Psang, PNG are here attributed to S. viridis and are well within its distributional range (Fig. 25). Hubendick (1946: 30 – 32, 63) considered the validity of ‘ Parellsiphon promptus’ (= S. viridis) doubtful, revealing considerable resemblance in shell characters and possible synonymy with S. acmaeoides. Hubendick (1946: 53) further indicated that the mollusc is ‘ Entirely yellowish white’ (i. e., likely discolouration in preserved material; external foot dark grey in live S. viridis). The specimen from Queensland figured in Hubendick (1946: 47, pl. 3, fig. 18) as ‘ S. laciniosa v. sipho’ is attributed here to S. viridis. References to ‘ S. laciniosa ’ (in part) in Hubendick (1946: 47, from New Guinea and Huon Is) and ‘ S. laciniosa ’ in Hubendick (1955: 7, from Cairns (MV F 15031, MV F 13925, MV F 13923) are also most likely specimens of S. viridis. Hubendick’s (1946: pl. 3, fig. 22) specimen of ‘ S. stellata ’ from Broome and Thursday Island (pl. 3, fig. 20) are likely specimens of S. viridis, too. A specimen of ‘ S. zanda ’ from ‘ Java Sea’ figured in Hubendick (1946: 91, pl. 4, fig. 8 – 11) is likely a specimen of S. alba and a specimen from Madagascar (figs 10 – 11) is likely S. madagascariensis. Specimens of ‘ S. australis ’ from Masthead and Heron Islands (MV F. 13924, 15032, 15034, 15038) mentioned by Hubendick (1955: 7) are likely S. viridis. Reference to ‘ S. laciniosa ’ from New Guinea in Hubendick (1947 b: 3) is likely of S. viridis. The records of ‘ S. laciniosa ’ from East coast Malaysia in Way & Purchon (1981: 321) are most likely of S. viridis. Figured specimens of ‘ laciniosa group, unit 25 ’ in Dayrat et al. (2014) from Timor (fig. 4 J), Singapore (fig. 4 L), Magnetic Is (fig. 4 P), Broome, WA (fig. 4 N), PNG (fig. 4 Q) and Vanuatu (fig. 4 R) correspond well with typical features of S. viridis.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFA48223FCCAFD22FBCAFBF6.taxon	distribution	Distribution and habitat. Tropical IWP, including Palau, Singapore (St Johns Island), Vanuatu, and northern Australia (Qld through to Kimberley, WA) as well as Mauritius, Indian Ocean (Fig. 25). Records from Sulawesi, Bali, and PNG arebased on specimens sequenced by Dayrat et al. (2014) and found to be members of unit 25. In this study found in protected positions (e. g., hollows) on moderately exposed rocky boulder shores, mid littoral level (upper oyster zone).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFA2825EFCCAF922FF4BFD96.taxon	description	(Figs 26 A – E, 27 A – B)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFA2825EFCCAF922FF4BFD96.taxon	materials_examined	Material examined. Type material. Lectotype of Siphonaria guamensis Quoy & Gaimard, 1833, present designation, from ‘ Est tres-commune dans l’ile dont elle porte le nom, surtout au port d’Humata’ [is very common in the island of which it is named, especially at the port of Humata, Iles Mariannes [Umatac Bay, Guam] (MNHN IM 2000 - 35953, Fig. 26 A). Ten paralectotypes, same data as lectotype (MNHN IM 2000 - 5054). Other, non-type material. Guam: Umatac Bay Nth point, 13 ° 17.917 ’ N, 144 ° 39.494 ’ E GM 01 - 1 (AM C. 585553 10 + p, C. 584911 p [SK 142], C. 584929 p [SK 378], C. 584928 p [SK 379], C. 585031 p [SK 426]); Umatac Bay, Sth point, 13 ° 32.549 ’ N, 144 ° 48.443 ’ E GM 01 - 3 (AM C. 585392 10 p, C. 584870 p [M 344], C. 584871 p [M 345], C. 584872 p [M 346]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFA2825EFCCAF922FF4BFD96.taxon	discussion	Taxonomic remarks. The largest syntype with clearest external sculpture is herein designated as the lectotype of Siphonaria guamensis for the stabilisation of the name (Art. 74.1 of the Code). Labels associated with type specimens just give ‘ Iles Mariannes’. Our delineation of this species is based on comparative analyses of the morpho-anatomy and mitochondrial genetics of freshly collected topotypes specimens (Figs 26 B – C) and a series of additional specimens (Table S 1). The topotypes studied here match characteristics mentioned in the original description, such as ‘ raised and distinctive ribbing, prominent dark radial banding and the slightly hooked apex anteriorly [sic! posteriorly] ’.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFA2825EFCCAF922FF4BFD96.taxon	description	External morphology. Foot sole, foot wall, pneumostome, cephalic folds and mantle evenly cream / light grey; genital pore inconspicuous, located on foot wall posterior to right cephalic fold; mantle narrower than foot wall, thin translucent, edge weakly lobed with light black bands aligning to rib interstices extends to shell edge, pneumostomal lobe thin and within mantle between the right ADMs, closes the pneumostomal and anal openings at the mantle edge; two small black epithelial eye spots centralised on two thick centrally touching cephalic folds; light black pigmentation over posterior foot wall and centre of cephalic folds. Shell (Figs 26 A – C, E; Table S 9). Small sized (max sl mean = 10.96 mm, SD = 1.6 mm, n = 6), height medium; ovate; apex offset weakly to posterior and left; apical sides convex, posterior concave to straight; protoconch hooks to posterior, area black colouration, protoconch direction weakly homostrophic (n = 2; Fig. 26 E), shell whorl dextral; growth striae weak; shell thickness thin; colouration uneven with some radial banding; rib count (mean = 38, SD = 4.9, n = 6); slightly raised, pale white, fairly straight, faintly protrude beyond shell lip; predominantly primary ribs, in parts interspersed with 0 – 2 finer secondary ribs, rib interstices darker; siphonal ridge indistinct. Interior evenly dark brown to black from margin to spatula, paler on shell lip aligning under rib ends, siphonal groove clear; ADM scar indistinct, CMS weakly convex; No evidence of growth variations in shell thickness or shell margin colouration. Reproductive system (Fig. 27 A; n = 2). Positioned withincoelomundertherespiratorycavity, hermaphroditic glands positioned to posterior against right foot wall and over foot sole, epiphallic parts positioned over and looped / folded in front of BM and to side of RAM; AO large, elongated, bluntly pointed (embeds into MG), centrally bent with MA, merges with indistinct GA; ED relatively short, slightly twisted, thick with side appendage; EG relatively large, folded, elongated and pointed; single short narrow bent flagellum F 1 on EG; AO, GA and ED all muscular white tissue; BD and CD with opposing connections (bulbous at CD) into GA between ED, AO and GP; BD longer and narrower than CD with a prominent distal loop, top of loop attached via a long MA to inner foot wall in front of BM, both ducts smooth and pass together through RAM connecting into folds of MG (BD above CD), BC translucent test, midsized and bulbous; HD brownish long coiled links AG to a small elongated narrow brownish finely granulated HG; MG and AG small folded soft white tissue; dark SV embedded within AG, AG larger than HG, sides match curvature of inner foot wall at right posterior quarter of coelom. Spermatophore (Fig. 27 B). Thread-like, test thin, smooth, featureless, translucent (length = 9.38 mm, n = 1), head bulbous, tip bluntly rounded, containing a white gelatinous mass; taper region into the filamentous transparent flagellum is extended; both sections smooth, featureless; head longer and much thicker than flagellum (head length = 7.00 mm, head ~ 75 % of SPM length; head width = 80 μm; flagellum width = 13 μm); three SPM tightly coiled in a brown gelatinous mass found in one BC [SK 241]. Radula. Dentition formula 26: 1: 26 (Hubendick 1946: 42).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFA2825EFCCAF922FF4BFD96.taxon	diagnosis	Comparative remarks. Based on our molecular phylogeny (Figs 1, 3), S. guamensis (plicata group, unit 70) is the sister species of an unidentified Siphonaria species from Rarotonga (unit 17). The next most closely related species is S. nusalikensis sp. nov. (unit 89). The minimum distance between S. guamensis and these species is 9.6 % (unit 17) and 11.1 % (unit 89) in COI. The minimum distance to any other species is at least 25 % (Table S 7). Three congeners occur in sympatry with S. guamensis in Guam: Siphonaria lirata is very similar, but has finer ribbing, paler interior colouration, a non-hooked protoconch, and a shorter, wider ED, and longer F 1. Siphonaria normalis is extremely variable, but has less raised ribbing, a non-hooked apex, paler interior, a shorter, wider ED and F 1, larger BC, and shorter SPM. Siphonaria tanguissonensis has more raised and prominent ribbing, an unhooked apex, paler golden-brown interior with more prominent dark rays on shell lip, a smaller AO, longer BD, F 1 looped, and a longer SPM. None of these species are closely related to S. guamensis (Figs 1, 3). RS figures of ‘ S. guamensis ’ in Hubendick (1945: 26, fig. 33, 37) correspond well with S. costellata sp. nov., but not S. guamensis. Correspondingly, specimens from ‘ Billiton, Paudua, The Sunda Islands’ depicted as ‘ S. guamensis ’ by Hubendick (1946: 93, pl. 6, fig. 30 – 32) are likely S. costellata sp. nov. for resembling shell characteristics of this species, but not of S. guamensis. ‘ Siphonaria guamensis ’ recorded from Singapore (Tan & Chou, 2000: 115, fig 115) is also likely S. costellata sp. nov. based on shell characteristics, such as interior colouration, protoconch shape, which differs in S. guamensis. Unfigured records of ‘ S. guamensis ’ from Hong Kong (Christiaens, 1980 a: 81), Anambas and Natuna Islands (Tan & Kastoro, 2004: 50; Chim & Tan, 2009: 269) are mis-identified specimens of S. sipho. The identity of ‘ S. guamensis ’ reported from Visakhapatnam, India by Murty et al. (2013: 104, fig. 1) is erroneous and outside of the range of this species.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFA2825EFCCAF922FF4BFD96.taxon	distribution	Distribution and habitat. Endemic to Guam (Fig. 28), found in crevices on rock platforms and boulders on exposed rocky shores at mid to upper littoral levels (Fig. 26 D).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFDF825BFF68FD22FED7FB16.taxon	description	(Figs 26 F – O, 27 C – E)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFDF825BFF68FD22FED7FB16.taxon	materials_examined	Material examined. Type material. Lectotype of Siphonaria zelandica Quoy & Gaimard, 1833, from Nouvelle Zealandie [New Zealand (in error for Australia, see Jenkins, 1983: 5)] (MNHN IM 2000 - 5055, Fig. 26 F). Two paralectotypes, same data as lectotype (MNHN IM 2000 - 5135). Lectotype of Siphonaria baconi Reeve, 1856 from Swan River [Perth, WA] (NHMUK 1979164.1, Fig. 26 J). Two paralectotypes of Siphonaria baconi Reeve, 1856. Same data as lectotype (NHMUK 1979164.2 and 1979164.3). Lectotype of Siphonaria zebra Reeve, 1856 from Port Jackson [on type label; ‘ Philippine Islands’ in Conch. Icon. incorrect — see Taxonomic remarks below] [Sydney, Australia], (NHMUK 1979168 / 1, Fig. 26 H). Three paralectotypes of Siphonaria zebra Reeve, 1856. Same data as holotype (NHMUK 1979168 / 2 - 4). Syntype of Planesiphon elegans Iredale, 1940 from Keppel Bay, Qld; coll. H. Bernhard, Sept 1935 and 1936 (AM C. 103710, Fig. 26 L). Syntypes of P. elegans Iredale 1940. Expanded syntype data (AM C. 108528, 6 d, locations on label of North Keppel Island and Emu Park; AM C. 126707, 5 d, location on label of Emu Park; MV F 13845, 1 d, location on label of Emu Park). Other, non-type material. Australia, Qld: Keppel Bay, 23 ° 25 ’ S, 150 ° 55 ’ E (AM C. 103710 d); Emu Park, 23 ° 15 ’ S, 150 ° 49 ’ E (AM C. 126707 5 d); North Keppel Is & Emu Park 2, 3 ° 25 ’ S, 150 ° 55 ’ E (AM C. 108528 6 d). Caloundra Shelly Beach, 26 ° 48 ’ S, 153 ° 9 ’ E (AM C. 391924 d). NSW: Fingal Bay nr Port Stephens, 32 ° 44.990 ’ S, 152 ° 10.481 ’ E, NSW 09 - 1 (AM C. 585559 3 p); Broken Head Terrigal, 33 ° 26.796 ’ S, 151 ° 27.030 ’ E, NSW 08 - 1 (AM C. 585380 10 p, C. 585052 p [M 098]); Terrigal The Skillion, 33 ° 27.008 ’ S, 151 ° 27.122 ’ E, NSW 08 - 2 (AM C. 585591 4 p, C. 585057 p [M 222]); Wy-ar-gine Point Balmoral, 33 ° 49.159 ’ S, 151 ° 15.195 ’ E, NSW 06 - 5 (AM C. 585445 11 p, C. 585047 p [M 155], Fig. 26 I, C. 585048 p [M 156], C. 585049 p [M 157]); Bombo Kiama, 34 ° 39.232 ’ S, 150 ° 51.649 ’ E, NSW 03 - 1 (AM C. 585557 3 p); Murunna Point Camel Head, 36 ° 22.720 ’ S, 150 ° 04.766 ’ E, NSW 02 - 1 (AM C. 595960 2 p). Vic: Bear Gully, 38 ° 53.519 ’ S, 145 ° 59.029 ’ E, V 07 - 3 (AM C. 585572 3 p); Caves, Inverloch 38 ° 39.777 ’ S, 145 ° 40 ’ 871 ’ E, V 07 - 2 (AM C. 585359 p); San Remo, headland, 38 ° 31.913 ’ S, 145 ° 22.209 ’ E, V 07 - 4 (AM C. 585582 34 p); San Remo, 38 ° 31.489 ’ S, 145 ° 21.858 ’ E, V 07 - 1 (AM C. 585358 p); Frankston, 38 ° 09.236 ’ S, 145 ° 06.457 ’ E, V 06 - 1 (AM C. 585651 5 p); West Head Flinders, 38 ° 28.883 ’ S, 145 ° 01.727 ’ E, V 06 - 3 0 (AM C. 585717 8 p); Point Lonsdale (nr Queenscliff), 38 ° 17.276 ’ S, 144 ° 36.977 ’ E, V 05 - 1 (AM C. 585608, 4 p); Roadknight Point, 38 ° 25.707 ’ S, 144 ° 11.102 ’ E, V 04 - 1 (AM C. 585474 16 p); Marengo Rocks Apollo Bay, 38 ° 46.772 ’ S, 143 ° 39.997 ’ E, V 03 - 1 (AM C. 585650 5 p); Crofts Bay, 38 ° 35.363 ’ S, 142 ° 50.633 ’ E, V 01 - 3 (AM C. 585607 4 p, C. 585284 p [M 177]); Armstrong Bay, 38 ° 21.012 ’ S, 142 ° 21.633 ’ E, V 01 - 2 (AM C. 585726 9 p). SA: Cape Northumberland, 38 ° 03.503 ’ S, 140 ° 40.378 ’ E, SA 15 - 1 (AM C. 585351 p); Cape Northumberland Port Macdonnell, 38 ° 03.308 ’ S, 140 ° 39.398 ’ E, SA 15 - 2 (AM C. 585723 9 p, C. 585223 p [M 200]); Cape Thomas, 37 ° 04.461 ’ S, 139 ° 44.659 ’ E, SA 14 - 1 (AM C. 585766 7 p, C. 584893 d [R 3032], C. 584935 p [SK 038], C. 585218 p [M 199]); Fisheries Bay Lands End, 35 ° 37.999 ’ S, 138 ° 06.921 ’ E, SA 13 - 2 (AM C. 585713 8 p, C. 584900 p [SK 509], C. 585216 p [SK 024]); Groper Bay nr West Cape, 35 ° 14.108 ’ S, 136 ° 49.883 ’ E, SA 10 - 1 (AM C. 585601 4 p); Pondalowie Bay, 35 ° 13.989 ’ S, 136 ° 49.892 ’ E, SA 10 - 2 (AM C. 585473 16 p, C. 585492 p); Port Neill, 34 ° 07.102 ’ S, 136 ° 21.271 ’ E, SA 06 - 1 (AM C. 585710 8 p); Port Moonta, 34 ° 03.273 ’ S, 137 ° 33.592 ’ E, SA 09 - 1 (AM C. 585647 5 p); Salmon Point, 33 ° 38.547 ’ S, 134 ° 51.916 ’ E, SA 04 - 2 (AM C. 585688 7 p); Whyalla, 33 ° 02.539 ’ S, 137 ° 35.511 ’ E, SA 07 - 1 (AM C. 585675 6 p); Rocky Point, 32 ° 12.250 ’ S, 133 ° 14.861 ’ E, SA 02 - 4 (AM C. 5856877 p); Ceduna, 32 ° 07.438 ’ S, 133 ° 40.260 ’ E, SA 03 - 2 (AM C. 585707 8 p); Denial Bay Ceduna, 32 ° 05.886 ’ S, 133 ° 34.817 ’ E, SA 03 - 1 (AM C. 585762 10 p); Port Le Hunte Point Sinclair, 32 ° 05.554 ’ S, 132 ° 59.476 ’ E, SA 02 - 2 (AM C. 585706 8 p); Wandrilla Beach, nr Cape Nuyts 32 ° 01.894 ’ S, 132 ° 16.052 ’ E, SA 01 - 1 (AM C. 585522, 20 p). WA: Cave Point, 35 ° 06.965 ’ S, 117 ° 54.080 ’ E, WA 62 - 1 (AM C. 584768 5 p, WAM S 74186 5 p); Waterbay Point, 35 ° 05.540 ’ S, 117 ° 56.963 ’ E, WA 62 - 2 (AM C. 584684 2 p, WAM S 74187 2 p); Peaceful Bay, 35 ° 02.989 ’ S, 116 ° 55.769 ’ E, WA 60 - 7 (AM C. 584744 4 p, WAM S 74183 4 p); 35 ° 02.865 ’ S, 116 ° 55.722 ’ E, WA 60 - 8 (AM C. 585579 8 p); Wilson Head Ocean Beach, 35 ° 02.250 ’ S, 117 ° 19.894 ’ E, WA 61 - 1 (AM C. 584730 10 p, WAM S 74185 5 p); Cape Riche 34 ° 36.213 ’ S, 118 ° 45.401 ’ E, WA 62 - 5 (AM C. 584714 5 p, WAM S 74188 5 p); Bremer Bay Boat Harbour, 34 ° 25.613 ’ S, 119 ° 23.818 ’ E, WA 63 - 2 (AM C. 584773 5 p, WAM S 74189 5 p); Sarge Bay Cape Leeuwin, 34 ° 22.091 ’ S, 115 ° 08.820 ’ E, WA 60 - 4 (AM C. 585371 p); Augusta, 34 ° 20.451 ’ S, 115 ° 10.069 ’ E, WA 60 - 5 (AM C. 584699 3 p, WAM S 74182 3 p); Alexander Bay 2, 33 ° 53.467 ’ S, 122 ° 44.995 ’ E, WA 64 - 4 (AM C. 584776 5 p, 74191 5 p); Alexander Bay, 33 ° 53.374 ’ S, 122 ° 44.922 ’ E, WA 64 - 3 (AM C. 584774 5 p, 74190 5 p); Salmon Beach Esperance, 33 ° 53.254 ’ S, 121 ° 50.381 ’ E, WA 64 - 2 (AM C. 585580 3 p); Blue Haven Beach Esperance, 33 ° 53.105 ’ S, 121 ° 51.687 ’ E, WA 64 - 1 (AM C. 585314 p [M 097]); Cowaramup Point, 33 ° 51.934 ’ S, 114 ° 58.904 ’ E, WA 60 - 3 (AM C. 585521 2 p); Yallingup, 33 ° 38.358 ’ S, 115 ° 01.481 ’ E, WA 60 - 9 (AM C. 584696 3 p, WAM S 74184 2 p); Point Dalling Dunsborough, 33 ° 35.955 ’ S, 115 ° 06.315 ’ E, WA 59 - 4 (AM C. 584750 5 p, C. 595950 p [M 129], WAM S 74180 5 p); Point Casuarina Bunbury, 33 ° 18.544 ’ S, 115 ° 38.201 ’ E, WA 59 - 3 (AM C. 585578 3 p); groyne nr Robert Point Mandurah, 32 ° 31.270 ’ S, 115 ° 42.409 ’ E, WA 59 - 1 (AM C. 585437 5 p, WAM S 74179 5 p); Fremantle Hbr breakwater 32 ° 03.342 ’ S, 115 ° 43.987 ’ E, WA 58 - 5 (AM C. 585612 4 p); Longreach Bay point, Rottnest Is, 31 ° 59.333 ’ S, 115 ° 32.063 ’ E RI 01 (AM C. 584783 4 p, WAM S 74164 4 p); Point Brown Swan River, 32 ° 02.344 ’ S, 115 ° 45.471 ’ E, WA 59 - 5 (AM C. 584709 4 p, C. 584899 p [SK 039], C. 585935 p [M 127, SK 243], WAM S 74181 4 p); Grey, 30 ° 39.968 ’ S, 115 ° 08.072 ’ E, WA 58 - 2 (AM C. 585367 p); Jurien Bay 30 ° 17.244 ’ S, 115 ° 02.482 ’ E, WA 58 - 1 (AM C. 585576 3 p); Cambawarra Head Green Head, 30 ° 04.136 ’ S, 114 ° 57.830 ’ E, WA 57 - 6 (AM C. 585518 2 p); Illawong, 29 ° 42.254 ’ S, 114 ° 57.542 ’ E, WA 57 - 5 (AM C. 584708 4 p, WAM S 74178 4 p); Illawong bch rocks, 29 ° 42.198 ’ S, 114 ° 57.551 ’ E, WA 57 - 4 (AM C. 584705 4 p, WAM S 74177 3 p); S end Leander Point Port Denison, 29 ° 16.725 ’ S, 114 ° 54.918 ’ E, WA 57 - 2 (AM C. 584682 2 p, WAM S 74176 2 p); Leander Point Port Denison, 29 ° 16.568 ’ S, 114 ° 54.858 ’ E, WA 57 - 1 (AM C. 584692 3 p, WAM S 74175 2 p); Cape Burney Geraldton, 28 ° 52.084 ’ S, 114 ° 38.056 ’ E, WA 54 - 1 (AM C. 584748 3 p, WAM S 74174 3 p); Turtle Bay East Wallabi Is, 28 ° 25.804 ’ S, 113 ° 44.538 ’ E, WA 55 - 1 (AM C. 585658 5 p); Horrocks, 28 ° 21.469 ’ S, 114 ° 24.751 ’ E, WA 53 - 1 (AM C. 584681 2 p, WAM S 74173 2 p); Red Bluff, 27 ° 44.627 ’ S, 114 ° 08.576 ’ E, WA 52 - 2 (AM C. 584741 10 p, C. 595943 p [M 124], WAM S 74172 10 p); Chinamans Rock Kalbarri, 27 ° 42.776 ’ S, 114 ° 09.361 ’ E, WA 52 - 1 (AM C. 584740 10 p, WAM S 74171 10 p); Pepper Point (Zuytdorp), 26 ° 23.826 ’ S, 113 ° 18.268 ’ E, WA 51 - 1 (AM C. 584756 5 p, WAM S 74170 5 p); Bottle Bay Cape Peron, 25 ° 32.566 ’ S, 113 ° 29.467 ’ E, WA 49 - 1 (AM C. 585364 p); Point Quobba, 24 ° 29.124 ’ S, 113 ° 24.501 ’ E, WA 45 - 1 (AM C. 584707 4 p, WAM S 74169 4 p); Point Maud, 23 ° 08.322 ’ S, 113 ° 46.294 ’ E, WA 44 - 1 (AM C. 584723 5 p, WAM S 74168 4 p); Pt S of Bruboodjoo Pt Bateman Bay, 23 ° 02.991 ’ S, 113 ° 49.371 ’ E, WA 43 - 1 (AM C. 584733 12 p, WAM S 74167 6 p); Tantabiddi, 21 ° 54.739 ’ S, 113 ° 58.706 ’ E, WA 42 - 1 (AM C. 584691 3 p, C. 585121 p [SK 188], C. 585310 p [M 207], C. 585311 p [M 208], WAM S 74166 2 p); NW Cape Exmouth, 21 ° 48.360 ’ S, 114 ° 07.665 ’ E, WA 41 - 1 (AM C. 585363 p); Cape Keraudren, 19 ° 57.393 ’ S, 119 ° 46.358 ’ E, WA 29 - 2 (AM C. 584706 4 p, WAM S 74165 4 p); Cape Latouche Treville nr Gourdon Bay, 18 ° 27.457 ’ S, 121 ° 48.725 ’ E, WA 27 - 1 (AM C. 584342 p [SK 067], C. 584927 p [SK 066]); Catamaran Bay formal, 16 ° 27.622 ’ S, 123 ° 00.242 ’ E, WA 22 - 3 (AM C. 585304 p [M 045]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFDF825BFF68FD22FED7FB16.taxon	discussion	Taxonomic remarks. The original description of P. elegans contains no type designation. Hence, we consider all types as syntypes. Jenkins (1983) designated the lectotypes of S. zelandica (5, pl. 1, fig. a), S. baconi (10, pl. 1, fig. b) and S. zebra (11, pl. 1, fig. e). Anatomical and molecular analyses of freshly preserved topotypic specimens matching type specimens of each nominal species and geographical series validate the identity and occurrence of S. zelandica and confirm that S. zebra, Planesiphon elegans and S. baconi are junior synonyms. The descriptions of S. zebra and S. bifurcata were mixed up in Reeve’s monograph: The types of ‘ S. zebra’ match the description of S. bifurcata (species 22, figure 21) while the types of ‘ S. bifurcata’ match the description of S. zebra (species 21, figure 22) (see Jenkins, 1983: 28; White & Dayrat, 2012: 60). As a result, the correct type locality of ‘ S. zebra’ is Port Jackson instead of the Philippines Islands. The labelling errors in Reeve (1856) confused many subsequent authors. Iredale (1924: 276) also noted the transposition and correctly considered that S. zebra referred to the same species as S. baconi. Siphonaria zebra has been treated as a synonym of S. sipho by Reeve (1856), repeated in Schrenck (1867: 306). Hutton (1883: 143) refrained from accepting Reeve’s (1856) synonymy. The record of ‘ S. luzonica ’ in Adcock (1893, 11, SA) and listing of S. stellata, S. exigua, S. baconi and S. kurracheensis as its synonyms are incorrect. Herein we demonstrate that ‘ S. luzonica ’ (= S. sipho) and S. zelandica are distinct species with different distributions. Paetel (1883: 178, 1889: 430) and Galindo (1977: 416) incorrectly recorded S. zebra from the Philippines. Verco (1907: 104) incorrectly considered Trimusculus albida (Angas, 1878) as a possible white form of S. baconi. Hubendick (1946: 37) pointed out that Adcock’s (1893: 11) synonymisation of S. baconi with S. luzonica was questionable. The record of ‘ S. zebra ’ in Kuroda (1941: 137, pl. 3, figs 49 – 50) is a misidentification of S. rucuana. Hubendick (1946: 49) treated S. zelandica Quoy & Gaimard, 1833 as a synonym of S. australis Quoy & Gaimard, 1833. McAlpine (1952: 42) compared the conchology and anatomy of P. elegans, S. diemenensis and S. ‘ bifurcata’ and species interpretations by Iredale and Hubendick, from which Hubendick (1955: 4 – 5) subsequently indicated P. elegans was a synonym of S. zelandica. Confirmed in Jenkins (1983: 4 – 5, 28) reviewing the type specimens and redescribing the anatomy and distributions. Hubendick (1946: 69) uncertain of the identity of S. zebra, suggested it may be identical with S. plana. Morrison (1972: 56 – 58) assigned S. elegans, along with 29 other nominal species (S. zelandica not mentioned), to S. laciniosa without specific explanation other than based on similarity to a ‘ laciniosa ’ shell form and ‘ common reproductive development’. These synonymies are not supported by examination of type specimens and morpho-anatomy. ‘ S. zelandica’ in Berry (1977: 204 – 5, pl. 1), based on Hutton (1882: 343, correctly as S. australis) refers to S. australis. Trew (1983: 7) considered S. zebra a synonym of S. kurracheensis, probably following Hubendick (1946: 54), however, is presently indeterminable as location of the material in the collection is not indicated. Jenkins (1983: 5, 10 – 11, 28) and Wells (1984: 52), treated S. baconi as a synonym of S. zelandica. The taxon was redescribed by Jenkins (1983: 6, fig. 1 C). Here we utilise and expanded the information provided there.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFDF825BFF68FD22FED7FB16.taxon	description	External morphology (Fig. 26 M). Foot sole evenly dark grey, foot wall, foot edge mantle, cephalic folds and pneumostome lobe evenly yellow; cephalic folds large; mantle wider than foot wall, thin, translucent; mantle edge unthickened, black edge bands absent; faint irregular black blotches on foot wall and centre of cephalic folds, touch and extend over the mouth region; two small black subepithelial eye spots are centrally located cephalic folds. Shell (Figs 26 F – O; Table S 9). small to medium sized, ovate, (max sl mean = 17.2 mm, SD = 1.6 mm, n = 12), height low; apex offset posterior and left, apical sides straight to concave; shell whorl dextral, protoconch direction homostrophic (n = 2; Fig. 26 O); growth striae fine to inconspicuous; thickness fine; outer lip is uneven, often broken and finely scalloped with primary ribs extending past the outer lip; rib count (mean 48, SD = 5.1, n = 12), white primary ribs slightly raised, mainly broad, apically continuous, radiate from the apex with black to light brown interstices finely striated with four to six apically discontinuous secondary ribs; exteriorly the siphonal ridge is indistinct; pneumostomal margin of the shell slightly raised and the outer edge extends weakly out to the right side; interior smooth with brown to white spatula, slightly swollen margin; brown apical rays are often presented extending from the spatula to slightly swollen lip margin corresponding with exterior rib interstices; siphonal groove (fig. 1 C, sg) is shallow, smooth, of a similar colouration to the spatula; CMS (fig. 1 C, cms) is shallow and weakly concave to straight; ADM scars are similarly shallow, may be darker or lighter brown than the spatula. Juvenile shells are miniature adults displaying similar ribbing but with paler interiors and exteriors. Reproductive system (Figs 27 C – D; n = 5). HG (ovotestis), large yellow, granular and AG both located in posterior region of coelom, against inner foot wall and under the respiratory system; HG joined to anterior of soft folds of AG by a short pink lobed HD, passes and between the translucent soft white semicircular folds of the MG and the white AG; thin white CD emerges from MG. SV small ovate pinkish, connected via a short thin duct to the junction of the HD and CD. BD long thin white, passes alongside but dorsal to the much thicker CD through the adductor muscle and both enter adjacently into the GA; both CD and BD smooth, non-looped, featureless; BC small, brown, appears spherical when holding SPM but deflated when empty; ED long, thick, white and may be centrally looped (Jenkins, 1984: fig 1 b), enters the GA almost opposite the BD and CD juxtaposed points of entry, no AO present; EG, cream, soft, lobed, longer than ED; single F 1, short, broad, white, branching from the junction of the EG and ED. GP small, opens from GA through the foot wall, under the mantle and posterior to the right cephalic fold. Spermatophore (Fig. 27 E). Drop-like with short, hooked flagellum (length = 1.12 mm, n = 1); head section bulbous, round tip (head length = 1.0 mm, head ~ 86 % of SPM length; head width = 150 μm; flagellum width = 34 μm), body and flagellum test opaque, thick, tapering to a pointed end; both sections featureless; 1 SPM in brown gelatinous mass in one BC. Radula (in Jenkins, 1983: pl. 6, fig. a – c). Wide intraspecific variation exists in the number of transverse rows and the number of inner, mid and outer lateral teeth in longitudinal rows. The mean dentition formula is 28: 1: 28 (SD = 3.8, n = 7) with 122 parallel and slightly curved (anteriorly convex) transverse rows (SD = 6.5, n = 8). Of the 28 half row laterals, 10 (SD = 2.4) are inner, 2 (SD = 0.8) mid and 16 (SD = 1.5) outer teeth means respectively (n = 8). In each longitudinal row a gradual reduction in tooth size and an increase in distance between transverse rows occurs from the central tooth to the outer laterals. All teeth are bluntly concave posteriorly. The central tooth has a lower profile than the flanking laterals and a short pointed mesocone less than half the base length. The anterior is forked and wider than the notched posterior. The narrow base interlocks and articulates with the adjacent central teeth. Lateral teeth have broad bases. The mesocone of the inner laterals is either pointed or bicuspidate. The inner cusp is always longer and usually overlapping the posterior of the inner lateral in front. Ecto and endocones are absent. The mid laterals have an ectocone and a bluntly bicuspidate mesocone with the inner cusp being longer. Outer laterals have a square shaped base with a broad blunt mesocone flanked by short pointed ecto and endocones. The clefts between the mesocones and the side denticles of the outer laterals are widely variable in width and angle of separation. Around the third or fourth outer longitudinal row, often aberrant outer lateral teeth appear as fused teeth with double mesocones.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFDF825BFF68FD22FED7FB16.taxon	diagnosis	Comparative remarks. Siphonaria acmaeoides from Japan is the closely related sister species of S. zelandica (atra group, unit 26) (Figs 1, 2). Both species differ from each other by a genetic distance of at least of 5.8 % (Table S 4). The next closely related species are S. restis sp. nov. (distance = 23 %) and S. stowae (distance = 17.7 %) (Table S 4). Examination of the type specimens and topotypic specimens of S. zelandica and S. acmaeoides revealed differences in shell geometry, the spacing of ribs, secondary ribbing, colouration, and reproductive features. Both taxa occur in splendid isolation from each other and are maintained as distinct species. Throughout the range of S. zelandica, we found seventeen congeners with partly sympatric distributions. Five species are sympatric in southern Qld. For comparisons with S. atra, S. denticulata, and S. viridis refer to comparative remarks under these species. Siphonaria hienghenensis sp. nov. has a larger, darker shell with more raised ribbing, a more prominent siphonal ridge, darker interstices, and interior colouration, a larger AO and BC, BD with distal loop, and a thread-like SPM. Siphonaria opposita has a larger shell with a more prominent and flared siphonal ridge, and stronger edge scalloping, a larger AO, ED and BC, and a thread-like SPM. Siphonaria scabra has a larger, taller, darker shell with more raised ribbing, rougher exterior, stronger edge scalloping and a darker interior, larger AO and BC, longer ED, and a long thread-like SPM. Seven congeners are sympatric in NSW (along with S. denticulata and S. scabra). For comparison with S. diemenensis refer to comparative remarks there. Siphonaria emergens has a much smaller, orange-brown shell with less raised ribbing and an apex more posteriorly offset. Siphonaria funiculata has a shell with paler ribs and with dark brown interstices, a larger AO and BC, shorter BD, and a longer ED. Siphonaria pravitas sp. nov. has a taller shell with more raised prominent ribbing and siphonal ridge, stronger edge scalloping, a larger BC, and a thread-like SPM. Siphonaria stowae has a taller and much smaller, yellowish-cream shell, a larger BC, and a thread-like SPM. Six congeners are sympatric in Vic and SA Australia (along with S. denticulata, S. diemenensis, S. funiculata and S. stowae). Siphonaria jeanae has a smaller, darker, grey blue shell with brown ribs and purplish spatula, a very small to absent AO, broader and twisted ED, and a bulbous SPM. Siphonaria tasmanica has a taller, blue grey shell with a more central apex and less distinct siphonal ridge, a shorter BD, and smaller BC. Six congeners are sympatric in WA (along with S. atra, S. jeanae and S. stowae). Siphonaria alba has a larger shell with central apex, raised ribbing and stronger edge scalloping, a larger AO and BC, longer ED, and a longer, thread-like SPM. Siphonaria gemina sp. nov. has taller darker shell with more prominent siphonal ridge and stronger edge scalloping, a larger ED and BC, shorter BD, and a thread-like SPM. Siphonaria restis sp. nov. has a taller shell with more prominent and raised ribbing and siphonal ridge, stronger edge scalloping, a larger AO, BC and ED, long narrow F 1, and a thread-like SPM. Siphonaria zelandica displays a shell morphology like other species in the plicata group, but the structure of RS in each of these species differs (mainly size of BC; epiphallus parts of ED and F 1). Hubendick (1945: 29, 72, figs 44, 47; 1946: 56 – 57, pl. 4, figs 32 – 34) treated S. elegans as an accepted species. However, he mis-identified specimens of either S. sipho or S. bifurcate S. zelandica. The RS of ‘ S. elegans’ figured in Hubendick (1945: 30 – 31, figs 44, 47) corresponds to S. zelandica depicted herein (Fig. 27 C – D). The RS of ‘ D. bifurcata ’ figured by McAlpine (1952: 43, fig. 3) does also closely resemble that of S. zelandica. The records of S. zelandica by Lamarck (1836: 558), Hutton (1873: 55, 1878: 41; 1880: 36; 1883: 143, pl. 17, figs H – M; 1904: 68), Suter (1904: 68; 1913: 600; 1915), Iredale (1915), Oliver (1923: 498), Odhner (1924: 55), Bucknill (1924: 84), Powell (1933: 186; 1937: 86; 1939: 217; 1946: 91), Borland (1950: 386), Knox (1955: 86), Powell (1955: 120; 1957 a: 114), Dell (1960: 147, 1963: 227), Morton & Miller (1968: 83, pl. 19, fig. 8, 8 a), Galindo (1977: 416), Powell (1979: 292, pl. 54, fig. 10), Trew (1983: 7) and Wells & Wong (1978: 417) are all misidentifications of S. australis (refer Jenkins, 1983: 1). Hubendick (1946) was uncertain about the identities of S baconi and S. zebra; probably because of the mix-up of the figures in Reeve’s descriptions of both species). Hubendick (1945: 80; 1946: 37) treated ‘ S. baconi’ as accepted, but Hubendick (1946: 37, pl. 2, figs 9 – 13) tentatively placed it under ‘ S. bifurcata ’. The figures of ‘ S. baconi ’ in Hubendick (1946: 92, pl. 4, figs 33 – 34) are specimens of S. zelandica. Hubendick (1946: 54) also tentatively placed S. zebra in synonymy of ‘ S. kurracheensis ’ and listed the variant of S. kurracheensis var. zebra (pl. 2, fig. 37). This figure is attributed here to S. belcheri. Figures of ‘ S. elegans ’ in Hubendick (1946: pl. 4, fig. 33 – 34) from ‘ Kupang, Timor and Java Sea’ are likely specimens of S. viridis. Specimens figured as ‘ S. baconi’ in Hubendick (1946: 91, pl. 6, fig. 10 – 12) are specimens of S. zelandica. The SPM depicted herein (Fig. 27 E) resembles that figured in Hubendick (1955: 10, fig. 8) of ‘ S. zealandica’ apart from a shorter flagellum (possibly broken). The record of S. zelandica in Davey (1998: 117) is a specimen of S. denticulata. Figured specimens ‘ unit 26 ’ in Dayrat et al. 2014: fig. 5 B, C) correspond well with S. zelandica.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFDF825BFF68FD22FED7FB16.taxon	distribution	Distribution and habitat. Endemic toAustralia, from Keppel Bay, Qld, south along NSW, Victorian and SA coasts to Broome, WA (Fig. 28). Reasonably common in sheltered positions, often within runoff areas and shallow platform pools (Fig. 26 D), on exposed to moderately exposed rocky intertidal marine shores across upper and mid littoral levels.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFDA8254FF68FAA2FB24FB56.taxon	description	(Figs 29 A – D, M, 30 A – B)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFDA8254FF68FAA2FB24FB56.taxon	materials_examined	Material examined. Type material. Lectotype of Siphonaria plana Quoy & Gaimard, 1833, present designation, from Île-de-France [Mauritius] (MNHN IM 2000 - 35955, Fig. 29 A). Two paralectotypes, same data as lectotype (MNHN IM 2000 - 5056). Three syntypes of Siphonaria ferruginea Reeve, 1856 (NHMUK 1981001, Fig. 29 B). Other, non-type material. Mauritius: Souillac, 20 ° 31.467 ’ S, 57 ° 31.582 ’ E, MRU 01 - 2 (AM C. 584968 p [M 254]); Nth Albion, MRU 02 - 1 (AM C. 585734 21 p, AM C. 584969 p [M 251]; C. 584970 p [M 252]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFDA8254FF68FAA2FB24FB56.taxon	discussion	Taxonomic remarks. The largest syntype of S. plana is herein designated as the lectotype for the stabilisation of the name (Art. 74.1 of the Code). Our delineation of this species is based on comparative analyses of the morpho-anatomy and mitochondrial genetics of freshly collected topotypes (Fig. 29 C) and geographic series of additional specimens (Table S 1). We establish S. ferruginea (Fig. 29 B) as a new synonym. Reeve (1856) treated S. plana as a synonym of S. lineolata Sowerby I, 1835 based on specimens from Chile and Central America; which were misidentified (see Hubendick 1946: 68). Later, Carpenter (1864 a 545) incorrectly considered S. ferruginea as a variety of the New World species S. lecanium Philippi, 1846, intermediate between S. maura and S. palmata. Hubendick (1946: pl. 4, figs 16 – 19) depicted specimens from Mauritius that closely resemble the type of S. ferruginea (Fig. 29 B). However, his interpretation of this species is confused. He stated that S. plana was a ‘ nondefined or insufficiently defined’ species that may be identical with ‘ S. kurracheensis var. luzonica or zebra ’. He also assigned specimens from Queensland figured by Iredale (1940: pl. 34, figs 26 – 27) as ‘ L. optivus’ under S. ferruginea. However, we treat L. optivus as a junior synonym of S. viridis. Morrison (1963: 8) erroneously considered S. ferruginea as a synonym of the New World species S. alternata (Say, 1826). Subsequently, Morrison (1972: 56 – 58) treated S. plana as a junior synonym of S. laciniosa based on similarity in the shell and a ‘ common reproductive development’. This synonymy is not accepted herein based on examination of types and supplementary material.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFDA8254FF68FAA2FB24FB56.taxon	description	External morphology. Foot sole, foot wall, mantle, cephalic folds and pneumostomal lobe evenly pale grey / cream, paler at edge foot / wall; blotches of black pigmentation on centre of cephalic folds, faintly on foot wall; mantle narrower than width of foot wall, non-translucent, covers exposed inner shell lip, edge thickened, lobed, vertical bands of black pigmentation aligned with shell rib interstices; genital pore indistinct, located on foot wall to right anterior of right cephalic fold; two small black epithelial eye spots centralised on two centrally touching cephalic folds; pneumostomal lobe long under the mantle between the right ADMs. Shell (Figs 29 A – D; Table S 9). Small to medium sized (max sl mean = 16.8 mm, SD = 2.1 mm, n = 3), circular ovate; height low; apex offset left central and slightly to posterior, apical sides weakly convex, straight to concave at posterior; shell edge uneven; protoconch direction heterostrophic (n = 3), shell whorl dextral; growth striae prominent, exterior brown, radial colour banding often present, edge dark, mid pale, protoconch area darker; shell thickness medium; rib count (mean = 39.3, SD = 1.7, n = 3), ~ 20 distinct primary ribs, white to pale, fairly straight, rib growth uneven, ridges rounded, broaden to scallop and protrude beyond shell edge (<1 mm); 2 interspersed pale white finer secondary ribs, rib interstices darker; paired primary ribs on siphonal ridge, more prominent and extend further than other primary ribs. Interior shell dark chocolate brown, maybe mottled whitish; white rays extend from the shell lip to over the shell margin fading to the spatula, align under primary / secondary ribs, spatula dark chocolate brown; siphonal groove distinct, similar colour to margin and spatula; ADM scar distinct, CMS concave, variable colouration of shell interior common (e. g., Fig. 29 D). Reproductive system (Fig. 30 A; n = 1). Positioned within coelom under the respiratory cavity, hermaphroditic glands positioned to posterior against right foot wall and over foot sole, epiphallic parts positioned to anterior over back of BM and between RAM; size of RS small to animal size; GP singular, positioned through foot wall behind right cephalic fold, GA very small; AO short, narrow, bluntly pointed, joins at top underside of GA in conjunction with ED; ED elongated, broad, joins to back side of GA alongside AO; single short broad blunt centrally bent flagellum (F 1), same width as ED, appears as extension of ED, marked by connection of very small white folded EG; AO, GA and ED all muscular white tissue; BD and CD connect closely side-by-side into upper GA, CD connection bulbous, BD twisted with distal loop and MA over ED, both ducts narrow smooth featureless, pass together through RAM connecting into MG (BD above CD), BD often looped immediately in front of BC and longer than increasingly broader CD; BC large, elongated, embedded in folds of AG / MG; SV embedded on left side of AG; HD short, thick uncoiled, links AG to smaller yellowish granular HG; MG and AG folded, soft white tissue; both curved reflecting the close positioning to curvature of inner foot wall on right posterior of coelom; outer edge of small MG lobed. Spermatophore (Fig. 30 B). Thread-like, test thin, translucent (length = 10.17 mm, n = 1, AL = 14 mm); head section cylindrical, bulbous, centrally bent, rounded tip; test thin, smooth, featureless, translucent, tapers into short flagellum; head slightly shorter, wider than translucent flagellum (head length = 4.31 mm, ~ 42 % of total length, head width = 103 μm, flagellum width = 17 μm, n = 1); six tightly coiled SPMs found in a single BC of a topotypic specimen (AM C. 584970).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFDA8254FF68FAA2FB24FB56.taxon	diagnosis	Comparative remarks. In our mitochondrial phylogeny (Figs 1, 2), Siphonaria plana (atra group, unit 35) is the sister taxon of unit 34 (= S. opposita sp. nov.). Both species are separated by COI distances of ≥ 8.1 % (Table S 3). Siphonaria plana differs from S. denticulata by COI distances of ≥ 14.8 % and from other species by distances of ≥ 20 % (Table S 3). Throughout the range of S. plana we found six congeners to occur in partial sympatry. Three are sympatric in Mauritius: For comparison with S. viridis refer to comparative remarks under that species. Siphonaria griffithsorum sp. nov. has a smaller, taller shell with weaker edge scalloping, larger AO, longer BC and F 1. Siphonaria gemina sp. nov. has a smaller shell with more raised ribbing and weaker edge scalloping, a smaller BC, shorter BD, and shorter SPM. Siphonaria incerta has a smaller, taller shell with less raised ribbing and weaker edge scalloping, a larger AO, smaller BC, and longer F 1. Two species are sympatric in Mozambique: For comparison with S. capensis refer to comparative remarks under that species. Siphonaria carbo has a smaller, darker shell with finer ribbing and weaker edge scalloping, an indistinct AO, larger, elongate BC, BD without bursal or distal loops, and a smaller, bulbous SPM. Figured specimens ‘ atra group, unit 35 ’ in Dayrat, Goulding & White (2014: 263, fig. 5 M, Q) as well as Ossenbrügger et al. 2023 (figs 3 – 4) correspond well with S. plana and fall into the same genetic cluster, unit 35, as topotypic specimens examined herein.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFDA8254FF68FAA2FB24FB56.taxon	distribution	Distribution and habitat. Indian Ocean: recorded from Mauritius, Réunion, the Seychelles (Ossenbrügger et al. 2023), and eastern Africa (Pemba Bay, Zanzibar) (Fig. 28). Dayrat et al. (2014: 255) listed a single sequenced but unfigured specimen from Rangong Kampuan, Thailand in ‘ unit 35 ’. The occurrence of S. plana in Thailand requires confirmation. Common in sheltered positions on moderately exposed inner-lagoon rocky intertidal marine shores across upper littoral levels.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFD58255FCCAFB62FBF5F7F6.taxon	description	(Figs 29 E – G, N – O, Q – R, 30 C – D)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFD58255FCCAFB62FBF5F7F6.taxon	materials_examined	Material examined. Type material. Lectotype of Siphonaria plicata Quoy & Gaimard, 1833, present designation, from ‘ Île de Tonga-Tabou au village de Hifo’ [Hihifo, Tongatapu, Tonga] (NMHN IM 2000 - 35956, Fig. 29 E). Two paralectotypes, same data as lectotype (NMHN IM 2000 - 5057). Holotype of Siphonaria tongensis Hubendick, 1943 from ‘ Foua, Tonga-Inseln’ [Foua Island, Tonga]; coll. Eugenie Expedition, 1851 – 1853 (UUZM 1576, Fig. 29 N). Paratype, same data as holotype (UUZM 1576). Other, non-type material. Tonga: Tongatapu, Ha’atafu Beach Hihifo, 21 ° 04.140 ’ S, 175 ° 20.048 ’ W, TO 02 - 1 (AM C. 585539 20 + p, C. 585275 p [M 418, SK 104], C. 585276 p [M 419, SK 106], C. 585277 p [SK 105], C. 585278 p [SK 225]); Mau’i Rock, W coast Tongatapu, 21 ° 08.154 ’ S, 175 ° 20.706 ’ W, TO 01 - 1 (AM C. 585431 20 + p); Halafuoleva Beach, S coast Tongatapu, 21 ° 08.358 ’ S, 175 ° 02.443 ’ W, TO 03 - 1 (AM C. 585947 20 p, C. 585283 p [SK 223 [); nr Kolonga, N coast Tongatapu, 21 ° 12.021 ’ S, 175 ° 14.680 ’ W TO 04 - 2 (AM C. 585432 10 + p, C. 585432 p [SK 380]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFD58255FCCAFB62FBF5F7F6.taxon	discussion	Taxonomic remarks. The largest syntype with best preserved external sculpture (Fig. 29 E) is herein designated as the lectotype of S. plicata for the stabilisation of the name (Art. 74.1 of the Code). Our delineation of this species is based on comparative analyses of the morpho-anatomy and mitochondrial genetics of freshly collected topotypes (Fig. 29 F – G) and geographic series of additional specimens (Table S 1). We establish S. tongensis as a junior synonym based on the examination of the types (Fig. 29 N). Reeve (1856) erroneously considered S. plicata as a synonym of S. sipho, a view adopted by Hanley (1858 b: 152). Hubendick (1946) and Trew (1983: 6) placed S. plicata in the synonymy of S. laciniosa without providing an explanation or having examined the types. Morrison (1972) treated S. tongensis as a junior synonym of S. laciniosa based on similarity in shell form and for a ‘ common reproductive development’.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFD58255FCCAFB62FBF5F7F6.taxon	description	External morphology (Fig. 29 R). Foot sole centrally dark grey fading to yellowish at foot / wall edge; intensity of tissue colouration (i. e. of foot edge, foot wall and mantle) varies between paler and darker shell forms; external morphology is otherwise consistent; paler form (LIF 01 and FI 03 - 2) — foot wall and mantle intense yellow; darker form (TO 03 - 1 and LIF 02 - 1) — foot wall and mantle more greenish; both forms possess black pigmentation spots on foot wall, paler to foot edge; fringing mantle weakly lobed, translucent and reflects foot wall colouration, covers exposed inner shell lip; genital pore inconspicuous, located on foot wall to right anterior of right cephalic fold; two small black epithelial eye spots centralised on two centrally touching cephalic folds; pneumostomal lobe thin, part of the mantle, between the right ADMs, closes the pneumostome at the mantle edge. Shell (Figs 29 E – G, N, Q; Table S 9). Small to medium sized (max sl mean = 14.9 mm, SD = 0.4 mm, n = 6), ovate; height medium to tall; apex offset weakly posterior and to left, apical sides strongly convex, posterior side concave close to protoconch convex to margin; protoconch direction homostrophic to central (n = 3; Fig. 29 Q); shell whorl dextral; growth striae prominent in bands, shell thickness thick; ribs fairly even in width, rib count (mean = 56.3, SD = 1.7, n = 6), exterior uneven without prominent radial colour bands; primary ribs pale grey, fairly straight to wavy, slightly broaden increasingly raised to shell edge, ridges rounded narrow; edge finely scalloped and unevenly corrugated; 6 – 8 prominent spread radial ridges, one being siphonal ridge, each formed by 3 – 5 primary ribs; few finer secondary ribs, rib interstices darker grey; interior shell margin white, dark brown rays from shell lip over shell margin to spatula, align under rib interstices, siphonal groove distinct, same colour as shell margin; spatula golden brown to white; ADM scar distinct, darker brown, CMS convex; thickening and whitening of shell lip occurs (Fig. 29 E). Reproductive system (Fig. 30 C; n = 3). Positioned within coelom under the respiratory cavity, hermaphroditic glands positioned to posterior against right foot wall and over foot sole, epiphallic parts positioned over back of BM, close to BC and to side of RAM; AO medium, broad, blunt, merges with MA, joins to inner side of a small GA; ED relatively short, centrally twisted, bent; EG folded, bulbous with pointed tip; single long narrow looped or bent flagellum F 1 on EG; AO, GA and ED all muscular white tissue; BD and CD with opposing connections (bulbous at CD) into GA between ED, AO and GP; BD longer and narrower than CD with a prominent distal loop, top of loop attached via a long MA to inner foot wall in front of BM; both BD and CD smooth and pass together through RAM connecting into folds of MG (BD above CD), BC small, translucent test and bulbous drop shaped; HD brownish, broad coils, links AG to a small elongated narrow brownish granulated HG; MG and AG small folded soft white tissue; dark SV embedded within AG, AG larger than HG, sides match curvature of inner foot wall at right posterior quarter of coelom. Spermatophore (Fig. 30 D). Test thin, translucent (length = 10.89 ± 1.66 mm, n = 2); head bulbous, tip bluntly rounded, containing a white gelatinous mass; taper region into the filamentous transparent flagellum is extended; both sections smooth, featureless; head longer and much thicker than flagellum (head length = 6.09 mm, head ~ 57 % of SPM length; head width = 64 μm, SD = 1 μm; flagellum width = 1 μm); Six SPM tightly coiled in brown gelatinous mass in BC (AM C. 585275).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFD58255FCCAFB62FBF5F7F6.taxon	diagnosis	Comparative remarks. In our mitochondrial phylogeny, S. plicata (plicata group, unit 56) is genetically well-differentiated. It is related to several species from Pacific Islands, such as Guam, New Caledonia and Hawaii (Figs 1, 3). The species differs from other species by COI distances of ≥ 25 % (Table S 7). In Tonga we found one sympatric congener, S. tongatapuensis sp. nov. which differs by having a more prominent dual ribbed siphonal ridge, a more central apex, lower height, paler interior colouration, a shorter wider ED, a larger pointed AO and longer BD. with bursal loop. Generally, the shell of S. plicata is similar to that of other species in the plicata group.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFD58255FCCAFB62FBF5F7F6.taxon	distribution	Distribution and habitat. Known only from Tonga (Fig. 28). In this study found in sheltered positions (i. e., hollows on beach rock platforms, small pools, hollows of rocky cliff bases) on exposed rocky shores or landside of lagoons at mid and lower littoral levels. Two distinct forms appear to occupy different substrates: a smaller, taller shell morph (‘ monticulus ’ - morph) occupies pits or hollows in vertical limestone cliffs, the larger, flatter shell morph was (‘ plicata ’ - morph) is found on rock platforms, in hollows or on flat surfaces.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFD68252FF68FF02FDB3FDF6.taxon	description	(Figs 29 H – L, P, S – T, 30 E – G)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFD68252FF68FF02FDB3FDF6.taxon	materials_examined	Material examined. Type material. Holotype of Siphonaria lateralis Gould, 1846 from ‘ Burnt Island, Orange Harbor’ [Tierra del Fuego, Argentina]; coll. Carpenter (USNM 5853, figured in Güller et al. 2015: 88, figs. 6 A, E, I). Eight syntypes of S. macgillivrayi Reeve, 1856 from Island of St. Paul’s [Indian Ocean] (NHMUK 1981003). Lectotype of S. redimiculum Reeve, 1856, designated by Güller et al. (2015: 92, fig. 8 O, T), without type locality (NHMUK 1981004 / 1). Four paralectotypes of S. redimiculum, same data as lectotype (NHMUK 1981004 / 2 - 5; figured by Güller et al. 2015: fig. 8 P – S). Lectotype of Kerguelenella innominata (Iredale, 1915), designated by Boreham (1959: 71), from Disappointment Island, Auckland Islands [New Zealand] (GNS TM 1203, Fig. 29 H); two paralectotypes, from Auckland Is; coll. Capt Bollons (AM C. 43852). Holotype of Kerguelenia stewartiana Powell, 1939 from Akers Point, Stewart Island, NZ; coll. R. H. Harrison (AWMM MA. 70377); paratypes of K. stewartiana; same data as holotype, one (MA 71589), two (GNS TM 1969, TM 1970), 10 (AM C. 585966); one from Akers Island, Stewart Island, New Zealand; coll. R. H. Harrison (USNM 880297). Holotype of Kerguelenia macquariensis Powell, 1939: 238 from Macquarie Island, Australia; coll. H. J. Finlay (AWMM MA. 70376; Fig. 29 I). Two paratypes of K. macquariensis, same data as holotype (GNS TM 1969 – 1970). Holotype of Kerguelenella flemingi Powell, 1955 from Bay S of Crozier Pt, Auckland Island, NZ (GNS TM 1204). Two paratypes of K. flemingi, same data as holotype (GNS TM 1205, TM 1208). Other, non-type material. South Georgia Islands: Grytviken Bay, 54 ° 16 ′ 53.4 ” S, 36 ° 30 ′ 28.8 ″ W SGI 01 (WAM S 101182 p [M 475, SK 282]); Husivk 54 ° 10.76 ’ S, 36 ° 42.68 ’ W (NHMUK 1994092 p [SK 204]). NZ: Auckland Island, 50 ° 34 ‘ 38.4 “ S 166 ° 10 ‘ 01.5 “ E (AM C. 43852 2 d; C. 586004, 2 p). MI: (AM C. 46754 p [SK 550 K 9200 protoconch F 12]). Garden Cove, 54 ° 30 ’ S, 158 ° 57 ’ E (AM C. 265944 4 d, C. 220182 d [R. 211181], C. 91954 10 + p, C. 91973 p, C. 91974 p, C. 46727 d, C. 46701 3 d); North Garden Cove, 54 ° 29.91 ’ S, 158 ° 56.45 ’ E (AM C. 91965 10 + p, C. 91964 10 + p, C. 91963 10 + p, C. 91966 10 + p); Hasselborough Bay, 54 ° 29.91 ’ S, 158 ° 56.08 ’ E (AM C. 91970 10 + p, C. 91971 10 + p, C. 91969, C. 91967 10 + p), Hasselborough Corner (AM C. 91968 10 + p); Buckles Bay, 54 ° 30.04 ’ S, 158 ° 56.18 ’ E (AM C. 91962 10 + p, C. 585698 MI 02 8 p, C. 584960 p [M 468], C. 584961 p [M 469], C. 584962 p [SK 272]); Opposite Tern Rock (AM C. 91959 10 + p, p [SK 202], C. 91958 10 + p, C. 99957 6 p); Cosray Rocks (AM C. 91955 10 + p); Mawson Point, (AM C. 91952 10 + p); Sandy Bay, 54 ° 34 ’ S 158 ° 56 ’ E (AM C. 91953 10 + p, C. 586003 p).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFD68252FF68FF02FDB3FDF6.taxon	discussion	Taxonomic remarks. The type of S. lateralis was subsequently figured in Gould (1856: 12, pl. 30, fig. 462, 462 a – b). The holotype matches shell profile, dimensions and colouration of the figured specimen well. In an integrated morphological and molecular study Güller et al. (2015) established the identity of S. lateralis and its distinction from S. lessonii and S. fuegiensis. Hubendick (1946: 27) pointed out that various nominal species with shells resembling S. lateralis have been reported from the south Pacific between Australia, NZ and the Kerguelen. Building on the findings of Güller et al. (2015), our anatomical and molecular analyses of freshly preserved topotypic specimens matching the types of each nominal species validate the identity and occurrence of S. lateralis and establish K. innominata (TS, Fig. 29 K) and K. macquariensis (TS, Fig. 29 J) as new synonyms. The identity of other nominal species described from this region, such as S. flemingi, S. stewartiana, S. redimiculum and S. macgillvrayi remain uncertain and are not reviewed herein. Güller et al. (2015: 88) stated that due to provenance issues and character differences, the name S. redimiculum is likely not applicable to Siphonaria species from the South American region, while assigning the Magellanic records of S. macgillivrayi in Hubendick (1945; 1946) to S. fuegiensis. Iredale (1915: 478) introduced the new name Kerguelenia innominata for ‘ Siphonaria lateralis from the subantarctic islands of New Zealand’ as described by Suter (1913: 601, pl. 49, fig. 10). Subsequently, Powell (1955: 122) considered Kerguelenia innominata Iredale 1915 as a nomen nudum. However, it is a replacement name based on a detailed description (of S. lateralis) by Suter (1913) and therefore an available name (Art. 12.2. of The Code). Suter (1913: 601) also included S. redimiculum Reeve, 1856 and S. tristensis Leach (not Sowerby I, 1823) in the synonymy of S. lateralis. However, Tasmania is considered as an unlikely occurrence for a species otherwise distributed through the sub-Antarctic. The genus name Kerguelenella was introduced by Powell (1946: 91) as a replacement name for Kerguelenia with S. redimiculum Reeve, 1856 as the type species by original designation. Powell (1955: 120 – 122; 1979: 293 – 294), Dell (1964: 290 – 292), and Forcelli (2000: 132) treated this taxon as an accepted genus. Powell (1955: 122) treated K. macquariensis as a subspecies of K. lateralis stating that K. macquariensis is ‘ closer to K. lateralis than any other NZ form’. This was followed by Powell (1960: 168) and Dell (1964: 291). However, subsequently (Powell, 1979: 293) synonymised K. macquariensis with K. lateralis. Boreham (1959: 71) subsequently designated the lectotype of Kerguelena innominata restricting the type locality to Disappointment Island, Auckland Islands. A note on the lectotype label (GNS TM 1203) indicates ‘ nom. nov. for Siphonaria lateralis Suter (not S. lateralis Gould, 1846) (in part) Ex Suter colln 4015 ’. Powell (1979: 293, pl. 54, figs 16 – 17) considered Kerguelenella innominata as an accepted species. Dell (1964: 290) considered S. magellanica as a synonym of S. lateralis; however, Güller et al. (2015: 83) showed it to be a synonym of S. lessonii instead.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFD68252FF68FF02FDB3FDF6.taxon	description	External morphology (Fig. 29 P). Foot sole orange grey, paler to foot edge; foot wall grey to yellow green, cephalic folds and pneumostomal lobe paler, fold small; irregular blotches of black pigmentation around foot wall and cephalic folds. Shell (Figs 29 H – L; Table S 9). Medium to large sized (max sl mean = 15.8 mm, SD = 2.3 mm, n = 8, MI specimens), elongate ovate; height medium; exterior uneven, smooth to undulating often wrinkled, radially ribbed, reddish to greenish brown, apex offset strongly posterior and left, anterior and right apical sides strongly convex, posterior and left apical sides shorter straight to concave; protoconch hooked, below apex, close to posterior edge; protoconch direction weakly heterostrophic (n = 1; Fig. 29 T); growth striae prominent in bands, shell thin, lip even fragile, periostracum freely extending; rib count (mean = 31, SD = 2.5, n = 8), primary ribs flatly rounded, not protruding beyond shell lip; often interspersed finer secondary ribs, rib interstices darker; siphonal ridge indistinct but may be bulged, formed by paired primary ribs. Interior glossy, shell margin dark brown to tan, lip paler, paler markings weakly highlight under ribs, siphonal groove distinct, shallow, same colour as margin; spatula mottled tan, uneven darker markings; ADM scar distinct, CMS straight; thickening of shell lip not observed. Reproductive system (Figs 30 E, G; n = 3). RS positioned within right side of coelom under the respiratory cavity, hermaphroditic glands positioned to posterior against right foot wall and over foot sole, epiphallic parts positioned between over rear of BM and side RAM, tip of AO embedded in EG; ED large wide short centrally curved, EG indistinct and curved continuation at top of ED, no flagellum (F 1); AO and GA merged in base of ED as protruding bulb with MA; epiphallic parts all muscular tissue; CD and BD both short wide muscular wall featureless slightly curved, jointly connect to AO on inner side, BD over CD; pass just within outer side of RAM, BC large rounded, positioned between footwall and AG, test thick opaque, filled with orange-brown gelatinous mass; CD enters AG close to BD; single puckered GP at end of GA; HD small, brownish coils, links AG to brownish finely granulated HG; MG and larger AG small folded soft white tissue, MG at anterior of AG; dark SV embedded within AG, sides match curvature of inner foot wall at right posterior quarter of coelom. Spermatophore (Fig. 30 F). Elongated drop shaped, test thin, translucent (length = 4.4 ± 1.5 mm, n = 2), head bulbous, broad, tip bluntly rounded, containing a white gelatinous mass; taper region to flagellum reduced; both sections smooth, featureless; head much larger than flagellum (head length = 3.9 ± 1.3 mm, head length of SPM length ~ 88 %; head width = 1.06 ± 0.7 mm, flagellum width = 0.14 ± 0.03 mm, n = 2). Radula and jaw. Described and figured in Güller et al. (2015: 87, figs 2 D – F, 3 G – H). Dentition formula 46: 1: 46 (Hubendick 1946: 27).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFD68252FF68FF02FDB3FDF6.taxon	diagnosis	Comparative remarks. Siphonaria lateralis (lateralis group, unit 6) is the sister species of S. fuegiensis; both species together form a well differentiated subclade in the mitochondrial tree (Figs 1, 4). They differ by COI distances of ≥ 6.9 % (Table S 8). Siphonaria lateralis differs from other species by COI distances of ≥ 18 % (Table S 8). Within the Australian and New Zealand subantarctic distribution of S. lateralis, we found one sympatric congener on Auckland Island, S. obliquata (refer to comparative remarks under that species). No congeners were found on MI. Topotypic specimens of K. innominata (Fig. 29 H, lectotype) and K. macquariensis (Fig. 29 K) reveal typical features of S. laterialis, such as a relatively smooth (Fig. 29 I) or rough shell (Fig. 29 J), and corresponding RS structure, size and shape of epiphallic parts. They fall well within a very tight genetic cluster with sequences from South Georgia (unit 6; Dayrat et al. 2014: 266). Güller et al. (2015: 93) appeared to have mixed-up the species identifications when differentiating ‘ units 5 and 6 ’ delineated in Dayrat et al. (2014: 259). Figured specimens in Hubendick (1946: pl. 1, figs 22 – 25, from South Georgia) match shells of this species figured here (Figs 29 H – L). The RS depicted herein (Fig. 30 E, G) corresponds well with that depicted by Hubendick (1945: fig. 15) and Simpson (1977: 126, fig. 7). We found no noteworthy anatomical differences between specimens from South Georgia and MI (Figs 30 E, G).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFD68252FF68FF02FDB3FDF6.taxon	distribution	Distribution and habitat. Recorded from MI, south Pacific, southern Argentina and New Georgia, south Atlantic Ocean. In this study found at sheltered positions, clustered in crevices and hollows, on rocky shores at upper to mid littoral levels (Fig. 29 S).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFD3824DFF68FDC2FB8CF935.taxon	description	(Figs 31 A – I, O, S – U, 32 A – D)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFD3824DFF68FDC2FB8CF935.taxon	materials_examined	Material examined. Type material. Syntype of Siphonaria normalis Gould, 1846 from Hawaii, United States, North Pacific Ocean; coll. Carpenter (USNM 15346, Fig. 31 A). Lectotype of P. soranus Iredale, 1940, present designation, from Townsville Qld, Australia; coll. 1929 (AM C. 103709, Fig. 31 D). Seven paralectotypes, same data as lectotype (AM C. 124989). Two paralectotypes, same data as lectotype (MV F 13840). Other, non-type material. Marquesas: Baie des Vierges, Fatu Hiva, 10 ° 27,84 ’ S, 138 ° 39,97 ’ W (MNHN IM- 2013 - 74897 p [M 570, SK 508], IM- 2013 - 74898 p [M 569], IM- 2013 - 74899 p [M 565]). Hawaii, Oahu: N end Waikiki Beach, 21 ° 15.743 ’ N, 157 ° 49.307 ’ W (AM C. 584887 p [M 293]); N end Waikiki, 21 ° 16.646 ’ N, 157 ° 50.048 ’ W (AM C. 585622 5 p, C. 584886 p [SK 210], C. 585930 p [SK 209]). Maui: Hanakao’o Beach, 20 ° 54.564 ’ N, 156 ° 41.310 ’ W (AM C. 585929 p [M 296]); Makalua-puna Point, 21 ° 54.586 ’ N, 156 ° 41.338 ’ W (AM C. 585555 3 p); Ho’okipa Beach, 20 ° 56.029 ’ N, 156 ° 21.411 ’ W (AM C. 584892 p [SK 212]). Guam: Umatac Bay, N end, 13 ° 17.917 ’ N, 144 ° 39.494 ’ E (AM C. 585325 p C. 584869 p [SK 142], C. 585493 p [M 347], C. 585494 p [M 348]); Pago Bay, below UoG Marine Lab, 13 ° 25.645 ’ N, 144 ° 47.927 ’ E (AM C. 585996 10 + p); Tanguisson Beach, S end, 13 ° 32.549 ’ N, 144 ° 48.443 ’ E (AM C. 585994 2 p, C. 584873 p [M 343], C. 584874 p [M 439]). Fiji, Viti Levu: Vuda Point Marina seawall, 17 ° 40.878 ’ S, 177 ° 23.009 ’ E (AM C. 585970 20 + p, C. 584866 p [SK 115 protoconch C 9], C. 584867 p [M 286], C. 584868 p [M 287]). PNG: Sek Is., 05 ° 04,7 ’ S, 145 ° 48,9 ’ E (MNHN IM- 2013 - 13133 p [M 558], IM- 2013 - 13135 p [M 562]); Wonad I., 05 ° 08,1 ’ S, 145 ° 49,3 ’ E (MNHN IM- 2013 - 15280 p [M 563]); Riwo waters, 05 ° 08,9 ’ S, 145 ° 48,2 ’ E (MNHN IM- 2013 - 15250 p [M 555] IM- 2013 - 15251 p [M 554]). NC, Lifou: We Baie de Chateaubriand E coast (AM C. 585396 10 p, C. 584946 p [M 388], C. 584947 p [M 389]). S of Pouebo 20 ° 25.950 ’ S, 164 ° 39.251 ’ E NC 04 - 2 (AM C. 585006 p [M 381], C. 585007 p [M 382]); Bonhomme de Bourail, La Roche Percee, 21 ° 36.487 ’ S, 165 ° 27.423 ’ E (AM C. 585013 p [M 370], C. 585014 p [M 371]); Presqu’ile de Ouano La Foa, 20 ° 51.434 ’ S, 165 ° 48.479 ’ E (AM C. 595911 6 p). Australia, Qld: Umagico, 10 ° 53.125 ’ S, 142 ° 20.799 ’ E (AM C. 585178 p [SK 195]); Capt Billy Landing, 11 ° 38.019 ’ S, 142 ° 51.472 ’ E (AM C. 585415 10 + p, C. 584792 p [M 180], C. 584793 p [M 181], C. 584794 p [M 402]); S of Bathurst Head, 14 ° 17.583 ’ S, 144 ° 11.845 ’ E (AM C. 585348 10 p); Lizard Is, bch rock, 14 ° 40.730 ’ S, 145 ° 26.838 ’ E (AM C. 585643 5 p); Lizard Is, 14 ° 40.908 ’ S, 145 ° 27.007 ’ E (AM C. 585566 4 p, C. 585175 p [M 030]); Cape Kimberley, 16 ° 16.535 ’ S, 145 ° 28.737 ’ E (AM C. 585720 9 p, C. 585168 p [M 041], C. 585169 p [M 394], C. 585170 p [M 395], C. 585171 p [M 397]); Pebbly Beach Yule Reef Trinity Bay, 16 ° 35.031 ’ S, 145 ° 30.823 ’ E (AM C. 585703 8 p); Gribble Pt Mission Bay Yarrabah, 16 ° 53.781 ’ S, 145 ° 51.852 ’ E (AM C. 585347 p); Mourilyan Harbour, 17 ° 35.951 ’ S, 146 ° 07.583 ’ E (AM C. 585411 10 + p, C. 585155 p [M 012], C. 585156 p [SK 123]); W side Kissing Pt Townsville, 19 ° 14.332 ’ S, 146 ° 48.040 ’ E (AM C. 585672 6 p, C. 585146 p [M 083], C. 585147 p [M 186], C. 585148 p [M 187], C. 585149 p [M 188], C. 585931 p [SK 197], C. 585932 p [SK 075]); Slade Pt Mackay, 21 ° 03.813 ’ S, 149 ° 13.527 ’ E (AM C. 585501 2 p). Gulf of Carpentaria: Mutee Head, 10 ° 54.682 ’ S, 142 ° 15.204 ’ E (AM C. 585416 10 p); Weipa, 12 ° 37.795 ’ S, 141 ° 51.853 ’ E, Q 52 - 1 (AM C. 585450 12 p); Sweers Is: 17 ° 07.029 ’ S, 139 ° 35.805 ’ E (AM C. 585645 5 p), 17 ° 07.413 ’ S, 139 ° 35.816 ’ E, Q 56 - 2 (AM C. 585350 p), Inspection Pt, 17 ° 08.471 ’ S, 139 ° 36.868 ’ E (AM C. 585417 15 + p). NT: Cape Wirawawoi Nhulunbuy, 12 ° 09.513 ’ S, 136 ° 46.904 ’ E (AM C. 585530 20 + p); Sandy Is Pt 11 ° 07.862 ’ S, 132 ° 11.187 ’ E (AM C. 585406 10 + p); Smith Pt 2, 11 ° 07.466 ’ S, 132 ° 08.538 ’ E (AM C. 585636 5 p); Luxmore Hd Melville Is, 11 ° 20.639 ’ S, 130 ° 23.149 ’ E (AM C. 585349 7 p); Nightcliff Darwin, 12 ° 22.836 ’ S, 130 ° 50.402 ’ E (AM C. 585977 10 + p); Cox Peninsula, 12 ° 24.824 ’ S, 130 ° 40.921 ’ E (AM C. 585978 10 + p); N of Native Pt Dundee Bch, 12 ° 42.182 ’ S, 130 ° 20.881 ’ E (AM C. 585668 6 p); Native Pt oyster reef Dundee Bch, 12 ° 42.906 ’ S, 130 ° 20.653 ’ E (AM C. 585413 8 p); Native Pt reef Dundee Bch, 12 ° 42.981 ’ S, 130 ° 20.807 ’ E (AM C. 585381 10 p).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFD3824DFF68FDC2FB8CF935.taxon	discussion	Taxonomic remarks. Siphonaria normalis was subsequently figured by Gould (1856: 13, pl. 30, figs 468, 468 a – b). The syntype corresponds well with this figure in shell profile, dimensions and colouration. The description of P. soranus does not contain an original type designation. The figured specimen of P. soranus (Iredale, 1940: pl. 34, figs 20 – 21) is herein designated as the lectotype for the stabilisation of the name (AM C. 103709). Our delineation of this species is based on comparative analyses of the morpho-anatomy and mitochondrial genetics of freshly collected topotypes of S. normalis and P. soranus and geographic series of additional specimens (Table S 1). We stablish P. soranus as a junior synonym of S. normalis. Pilsbry (1920 b: 379) erroneously considered S. amara, S. nuttallii, S. lirata, S. crebricostata and S. normalis f. chirura as synonyms of S. normalis. This was followed by Cernohorsky (1972: 210). Dayrat et al. (2014: 267) recognised several independent molecular units within the normalis group (i. e., unit 12 from Thailand, unit 13 from Singapore, and unit 14 from Hawaii). These units are herein recognised as three distinct species, S. radiata (unit 12), S. costellata sp. nov. (unit 13), and S. normalis (unit 14).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFD3824DFF68FDC2FB8CF935.taxon	description	External morphology. Foot sole, foot wall, mantle, cephalic folds and pneumostomal lobe evenly pale grey / cream, paler at edge foot / wall; blotches of black pigmentation on centre of cephalic folds, faintly on foot wall; mantle narrower than width of foot wall, non-translucent, covers exposed inner shell lip, edge thickened, lobed, vertical bands of black pigmentation aligned with shell rib interstices; genital pore indistinct, located on foot wall to right anterior of right cephalic fold; two small black epithelial eye spots centralised on two centrally touching cephalic folds; pneumostomal lobe long under the mantle between the right ADMs. Shell (Figs 31 A – I, O, S – U; Table S 9). Small sized (max sl mean = 12.56 mm, SD = 2.02 mm, n = 13), circular ovate; height medium to tall; apex offset posterior and weakly left, apical sides straight to weakly convex, shell edge; protoconch direction heterostrophic, initially hooked (n = 4; Fig. 31 S – T), shell whorl dextral; growth striae prominent, radial colour banding often present, shell thickness thick; rib count (mean = 33, SD = 0.87, n = 13), primary ribs distinct, white to pale, usually radially evenly spread, fairly straight, broaden to and align with fairly flat faintly scalloped shell edge; ribs may be raised or flat,; 1 – 2 interspersed pale white finer secondary ribs, rib interstices darker; paired primary ribs on siphonal ridge, no more prominent than other primary ribs. Interior shell margin varies from pale tan to dark brown; white rays extend from the shell lip to over the shell margin fading to the spatula, align under primary / secondary ribs, spatula varies from pale yellow (Fig. 31 E), pale tan (Fig. 31 B) to dark chocolate brown (Fig. 31 F); siphonal groove distinct, paler than shell margin or spatula; ADM scar distinct, CMS straight, paler than shell lip; thickening of shell lip common, margin becomes whitened (Fig. 31 E). Reproductive system (Figs 32 A, C; n = 9). Positioned within coelom under the respiratory cavity, hermaphroditic glands positioned to posterior against right foot wall and over foot sole, epiphallic parts positioned to anterior between BM and RAM; GP small, singular, positioned through foot wall behind right cephalic fold, GA small, prominent; AO very small, short, narrow, blunt, slightly bent centrally, joins to top of GA in conjunction with ED; ED short, broad, joins to back side of GA alongside AO; single short broad blunt flagellum (F 1), longer than and same width as ED, join of F 1 to ED indistinct, marked by connection of very small white folded EG; AO, GA and ED all muscular white tissue; BD and CD connect closely side-by-side into GA between connections of ED and AO, both ducts narrow smooth featureless, pass together through RAM connecting into MG (BD above CD), BD longer than CD, often loop immediately in front of BC; BC large to medium, spherical, embedded in folds of MG, test translucent; SV embedded on left side of AG; HD short, thick coils, links AG to smaller yellowish granular HG; MG and AG folded, soft white tissue; sides match curvature of inner foot wall on right posterior of coelom; outer edge of MG lobed. Spermatophore (Figs 32 B, D). Broad head with short flagellum (length = 2.47 ± 0.195 mm, n = 3); head section cylindrical, bulbous, centrally bent, rounded tip; test thin, smooth, featureless, translucent encasing a white opaque central core; short tapering section (often looped) merges head to filamentous flagellum; head slightly shorter, wider than translucent flagellum (head length = 1.23 ± 0.45 mm, flagellum length = 1.24 ± 0.34 mm, SPM head ~ 49 % of total lenght; head width = 110 ± 13 μm, flagellum width = 11 ± 0 μm, n = 3); up to 9 SPMs tightly coiled in BC of topotypic specimens (AM C. 585930).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFD3824DFF68FDC2FB8CF935.taxon	diagnosis	Comparative remarks. Siphonaria normalis (normalis group, unit 14) is the sister species of a clade containing S. fuliginata and S. madangensis sp. nov. (Figs 1, 4). These three species are closely related and differ from each other by COI distances of ≥ 5.9 % (S. fuliginata, unit 80) and ≥ 5.6 % (S. madangensis sp. nov., unit 88) (Table S 8). Two other closely related species are S. campestra sp. nov. (unit 86, COI distance ≥ 18.6 %) and S. costellata sp. nov. (unit 13, COI distance ≥ 7.8 %) (Table S 8). Siphonaria campestra sp. nov. differs in having a shell with slightly broader ribs, predominantly primary and few secondary ribs, slightly more scalloped shell edge, darker interior, a smaller AO, shorter ED, and a thicker SPM. Siphonaria fuliginata has a much paler, thicker shell, less scalloped edge, a thicker BD, smaller BC, shorter and narrower ED, and a shorter SPM. Throughout the range of S. normalis we found twentyseven congeners to occur in partial sympatry. Siphonaria gemina sp. nov. has a smaller shell with stronger raised ribbing and edge scalloping, more prominent siphonal ridge, a larger AO and GA, a shorter, wider BD, and a wider SPM. Siphonaria mauiensis sp. nov. (sympatric in Hawaii) has a smaller shell with more raised ribbing and weaker edge scalloping, a smaller BC, shorter BD, and a shorter SPM. Siphonaria nuttallii (sympatric in Hawaii) has a taller shell with a more central apex, slightly greater raised ribbing, stronger edge scalloping, more prominent and multi ribbed siphonal ridge, a larger AO, smaller BC, broader BD, and a longer SPM. Eight congeners are sympatric in Fiji and NC. For comparison with S. atra refer to comparative remarks under that species. Siphonaria hienghenensis sp. nov. has a larger, taller, paler shell with a more prominent siphonal ridge and centralised apex, a larger AO, shorter ED and a smaller BC. Siphonaria monticulus has taller shell with slightly more raised and even ribbing, paler interior, a larger AO and a longer ED. Siphonaria namukaensis sp. nov. has a paler shell with more central apex, more prominent siphonal ridge, paler golden spatula, a larger AO and a smaller BC. Siphonaria caledonica sp. nov. has a taller shell with interstice markings, a darker interior, a larger AO and ED, and a smaller BC. Siphonaria bourailensis sp. nov. has a taller, paler shell with more prominent raised ribbing, stronger edge scalloping, a larger AO, and a smaller BC. Siphonaria poindimiensis sp. nov. has a taller shell with more prominent raised ribbing, stronger edge scalloping, a larger AO and ED, and a smaller BC. Siphonaria vudaensis sp. nov. has a larger shell with narrower ribbing, a more prominent and flared siphonal ridge, stronger edge scalloping, a larger AO, and a smaller BC. Three congeners are sympatric in Guam. Siphonaria guamensis has a slightly darker shell with finer and raised ribbing, paler margin, and darker spatula, a longer AO and ED, a smaller BC, and a longer SPM. Siphonaria lirata has a shell with finer raised ribbing, a larger AO, broader ED, and a smaller BC. Siphonaria tanguissonensis sp. nov. has a slightly darker shell with finer and raised ribbing, paler margin, and darker spatula, a smaller BC, shorter BD, and longer SPM, Four congeners are sympatric in PNG. For comparison with S. javanica refer to comparative remarks under that species. Siphonaria madangensis sp. nov. has a smaller shell with more raised ribbing and stronger edge scalloping, a smaller, less prominent AO, smaller ED, larger BC, longer narrower BD, and a very similar but shorter SPM. Siphonaria recurva sp. nov. has a darker shell with more prominent white ribbing and weaker edge scalloping, a larger AO and BC, and longer ED and F 1. Siphonaria viridis has a taller shell with dark patterning and distinct dual siphonal ridge, a larger AO, smaller BC, and longer SPM. Seven congeners occur sympatrically throughout Australia. Siphonaria costellata has a browner shell with more raised ribbing, prominent siphonal ridge, a longer BD, and longer SPM. Siphonaria gemina sp. nov. has a slightly taller darker shell with more raised ribbing, stronger edge scalloping, a smaller BC, and shorter BD. Siphonaria jiigurruensis sp. nov. has a taller darker shell with more prominent primary ribbing, red-brown patterning, a larger AO, smaller BC, longer and broader ED, and a longer SPM. Siphonaria oblia has a far smaller, darker, browner, and more fragile shell with unraised ribbing, apex strongly offset, a smaller AO, BD without distal loop, and larger ED and BC (Jenkins 2018: 278, fig. 3 C – D). Siphonaria opposita has a larger, lower shell with more prominent and flared siphonal ridge, central apex, stronger edge scalloping, a larger AO and ED. Siphonaria scabra has a larger, darker shell, greater raised ribbing, more prominent siphonal ridge, a larger AO, and a longer SPM. Siphonaria alba is sympatric in Singapore. It has a larger, darker shell with more prominent siphonal ridge, stronger edge scalloping, a larger AO, longer ED, and a longer SPM. Siphonaria normalis has a very wide distribution spanning from the northern to the southern Tropical Pacific (Fig. 25). We have not found any discontinuities in the ranges of anatomical, morphological, or mitochondrial variation that may suggest that this species as currently delineated may represent a species complex. The SPM of S. normalis resembles that of S. radiata and S. gemina sp. nov. Hubendick (1946: 30 – 32, 63) considered Parellsiphon soranus from Townsville, N Qld (= S. normalis) as of questionable status and possible synonym of S. acmaeoides for the ‘ considerable resemblance’ in shell characters. Figured specimens of S. normalis in Hubendick (1947 b: 2, fig 3 b – i, 4 b – i) appear to be a mixture of S. normalis (figs 3 a – d, i) and S. waikoloaensis sp. nov. (figs 3 f – h). The RS figured by Hubendick (1947 b: 2, fig 5) closely matches that of S. waikoloaensis sp. nov. Figured specimens of ‘ S. normalis ’ from Oahu, Hawaii in Hubendick (1947 b: 2, fig 3 a – I, 4 a – i) closely match S. normalis rather than S. mauiensis sp. nov. (finer ribbing, dark interior), S. nuttallii (usually paler interior, ribbing different), S. undans sp. nov. (distinct ribbing) or S. waikoloaensis sp. nov. (distinct ribbing). While Hubendick (1947 b: 2) stated that the specimen depicted in figs 3 a, 4 a ‘ except for its size, agrees with S. nuttalli [sic nuttallii] ’. However, it differs from types of S. nuttallii. The figured specimen of ‘ S. cf normalis ’ in Maes (1967: 154, pl. 14, fig. L, from CKI) is a specimen of S. gemina sp. nov. The figured specimen of ‘ S. normalis ’ from Lomalagi, Fiji in Cernohorsky (1972: 210, pl. 60, fig. 2) matches S. normalis depicted herein. The figured specimen of ‘ S. normalis ’ in Kay (1979: 493, figs 157 I – J) is a misidentification of S. nuttallii. The figured specimens of ‘ unit 14 ’ in Dayrat et al. (2014: figs 3 P, 5 Q, 3 R, 3 S) closely resemble S. normalis as delineated herein.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFD3824DFF68FDC2FB8CF935.taxon	distribution	Distribution and habitat. Widespread through tropical Pacific Ocean, including Society Islands, Gambier Islands, Marquesas, Hawaii, Nauru to Guam, American Samoa, Santa Cruz Islands, Solomon Islands, NC, PNG, northern Australia (Cape York through to Broome, Kimberley, WA), and Timor-Leste. In the Indian Ocean recorded from Praslin Island, Seychelles (Ossenbrügger et al. 2023). In this study found to be rather common in sheltered positions on exposed rocky shores at upper littoral levels in Australia, Fiji, New Caledonia, Hawaii, and Guam (Fig. 25).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFCC824EFCCAF882FD11F876.taxon	description	(Figs 14 J – L, O)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFCC824EFCCAF882FD11F876.taxon	materials_examined	Material examined. Type material. Lectotype of S. savignyi Krauss, 1848, present designation, type locality unknown, likely Red Sea; collector and date unknown (Savigny collection) (IM 2000 - 35936, Fig. 14 J). Paralectotypes, same data as lectotype (MNHN IM 2000 - 35935 d, Fig. 14 K; MNHN IM 2000 - 35934 d, Fig. 14 L).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFCC824EFCCAF882FD11F876.taxon	discussion	Taxonomic remarks. In the original description, Krauss (1848) attributed this name to Philippi. However, Krauss is author of the name, and not Philippi in accordance with Art. 12.2.6 of the Code (see also Bouchet & Danrigal, 1982: 15; Kabat & Coan, 2017: 205). The new name S. savignyi has been made available by Krauss (1848) through providing a bibliographic reference to pl. 1, fig. 1 in the work of Savigny (1817). Hubendick (1946: 55) incorrectly attributed the name to an alleged publication by Philippi (1826) entitled “ Gasteropodes et Qoquille in Savignyi: Descriptions de l’Egypte ”. However, this title is that of Savigny’s (1817) work and no published work was authored by Philippi under this title (Kabat & Coan, 2017).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFCC824EFCCAF882FD11F876.taxon	description	Shell (Figs 14 J – L; Table S 9). Small to medium sized (max sl mean = 41 mm, SD = 2.2 mm, n = 3), ovate; height medium to tall; apex offset weakly posterior and to left, apical sides strongly convex, protoconch direction undetermined; shell whorl dextral; exterior whitish, uneven without prominent radial colour bands, growth striae prominent in bands, shell thickness thick; rib count (mean = 49, SD = 2.9, n = 3); ribs fairly even in width, straight to wavy, ridges rounded narrow, slightly broaden and increasingly raised to shell edge, 8 – 10 spread prominent whitish primary ribs whitish; siphonal ridge formed by 3 – 5 primary ribs, flared up at shell edge; few finer secondary ribs, rib interstices darker, edge weakly scalloped and unevenly corrugated; interior shell margin white, short dark brown rays on shell lip, align under rib interstices, margin thickened white, siphonal groove distinct; same colour as mottled reddish to dark brown spatula; ADM scar distinct, darker brown, CMS slightly convex; thickening and whitening of shell lip occurs (Fig. 14 J). Reproductive system and Spermatophore. unknown.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFCC824EFCCAF882FD11F876.taxon	diagnosis	Comparative remarks. Figured shells from Port Fenfick and Gulf of Suez, Red Sea, and the description in Hubendick (1948: 55, pl. 4, 26 – 29) and figured shells from Israel in Morrison (1972: 60, figs 3 – 4) exhibit a morphology consistent with the types (Figs 14 J – L). No material has been available to study the anatomy and / or mitochondrial phylogenetics of this species.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFCC824EFCCAF882FD11F876.taxon	distribution	Distribution and habitat. So far recorded from the Red Sea and the Mediterranean coast of Israel.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFCF824FFCCAFF02FCF1FDF6.taxon	description	(Figs 30 H – I, 31 J – K)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFCF824FFCCAFF02FCF1FDF6.taxon	materials_examined	Material examined. Type material. Neotype of Siphonaria radians, present designation, from Natuna Regency, Riau Islands, NE coast of Palau Panjang (NW of Pulau Natuna Besar), Indonesia, South China Sea. 04 ° 15.9 ’ N, 108 ° 12.27 ’ E, Expedition Anambas, ZRC EA-ZJ 09; coll. 17 March 2002 (ZRC. MOL. 24912 [M 519, SK 435], Fig. 31 J). Other, non-type material. Indonesia: Pulau Panjang, Riau Islands 1 ° 10.215 ’ N, 104 ° 18.905 ’ E ZRC EA ZJ 09 (AM C. 595976 p [M 590, SK 512], C. 595977 p [M 591, SK 513]). Malaysia: Bak Bak Beach, Kudat, Sabah 07 ° 00 ′ N, 116 ° 46 ′ E (AM C. 585940 p [SK 521]); Tangung Bidara, Malacca 2 ° 17.568 ’ N, 102 ° 5.207 ’ E (ZRC. MOL. 24893 p [M 595, SK 527], ZRC. MOL. 24894 p [M 594, SK 526], Fig. 31 K, Pulau Langkawi 6 ° 18 ′ N, 99 ° 52 ′ E (AM C. 585697 p [SK 524]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFCF824FFCCAFF02FCF1FDF6.taxon	discussion	Taxonomic remarks. The name S. radiata A. Adams & Reeve, 1850 is a junior secondary homonym of S. radiata (Blainville, 1826) and a junior objective homonym of S. radiata Sowerby I, 1835. It is not invalidated by S. radiata Gray (1824), however, which is an unnecessary replacement name (see under S. sipho). H. Adams & A. Adams (1855) introduced S. radians as a replacement name (Art. 12.2 of the Code). No type material of S. radiata A. Adams & Reeve, 1850 could be found in the NHMUK and the types are therefore considered lost. The neotype of Siphonaria radians is designated herein to clarify the taxonomic status and the type locality of this taxon (Art. 75.3 of the Code).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFCF824FFCCAFF02FCF1FDF6.taxon	description	External morphology (preserved). Foot sole grey, lighter to foot edge; mantle, footwall, pneumostome and cephalic lobes cream with irregular darker pigmentation markings concentrated over cephalic lobes, mantle thin translucent lobed with white edge band and black marking aligning under rib interstices. Shell (Figs 31 J, K; Table S 9). External sculpture and height variable, lower fine ribbed form (Fig. 31 J), taller prominent ribbed form (Fig. 31 K); small to medium sized (max sl mean = 16.8 mm, SD = 2.1 mm, n = 11), circular ovate, low; apex slightly curved, apex offset slightly to posterior and left, apical sides weakly convex, posterior straight to concave, protoconch direction weakly homostrophic to central (n = 1), shell whorl dextral, shell thin, growth striae prominent, even, unraised; radial colour bands indistinct; rib count (mean = 48, SD = 12.4, n = 11), primary ribs indistinct from secondary ribs; ~ 16 pale brown to off white indistinct primary ribs, ridges weakly raised, rounded; siphonal ridge formed by 3 primary ribs, align with shell lip; shell edge uneven, weakly scalloped and corrugated; 2 – 3 finer secondary ribs between primary ribs, rib interstices darker. Interior shell margin and spatula dark chocolate brown, centre of spatula and distinct siphonal groove paler, off white to cream rays on shell margin, align under primary / secondary ribs; ADM scar distinct, similar to margin and spatula; CMS convex; whitening or thickening of shell lip not observed. The neotype (Fig. 31 J). Shell (sl = 17.9, sw = 14.2, sh = 5.2 mm) circular ovate, low; apex offset weakly to posterior and left, exterior grey, 68 fine weakly curved ribs; siphonal ridge formed by three adjacent ribs. Interior shell lip chocolate brown and cream under rib ends, dark brown rays on shell lip aligning under rib interstices. RS (Fig. 30 H). Reproductive system (Fig. 30 H; n = 5). Positioned within right side of coelom, against foot wall on foot muscle, under the respiratory cavity, epiphallic parts positioned between inner side RAM and behind BM; GA medium, with singular GP through foot wall; AO medium sized, broad, bluntly pointed, central bend, joined to upper GA; ED short wide, centrally twisted, joins to outer side of GA; GA, AO, ED all white muscular fibrous tissue; EG large sized, soft whitish tissue, folded, joins at junction of end of ED and extension of single broad long twisted flagellum (F 1); BD and CD connect closely in opposite directions into GA between ED junction and GP, CD junction into GA bulbous, both ducts long, smooth, narrow, whitish, featureless, pass closely together outside RAM (BD over CD) and broaden at junction into soft white folded tissues of MG; BD with distal loop and MA; CD connects to large MG / AG complex; BC embedded in MG folds, close to embedded SV; BC small, thin whitish translucent test; HD short, wide, coiled, links ducts in soft white folds of AG to yellowish granulated HG; outer edge of MG lobbed; AG slightly larger than HG. Spermatophore (Fig. 30 I). Thread-like, test thin, comprises a translucent cylindrical body section containing a white gelatinous core, tapers rapidly into a filamentous transparent flagellum (head length = 8.34 ± 1.5 mm, n = 4, ~ 76 % of SPM length, head width = 168 μm, ± 33 μm; flagellum width = 19 μm ± 2 μm), head section much thicker than flagellum, head tip bluntly rounded; both sections smooth, featureless; 3 and 4 SPM coiled, embedded in whitish gelatinous mass within 2 BC’s.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFCF824FFCCAFF02FCF1FDF6.taxon	diagnosis	Comparative remarks. Specimens identified as S. radians herein match well the original description and figures of S. radiata (A. Adams & Reeve, 1850). Siphonaria radians (atra group, unit 95) is the sister species of S. umbra sp. nov. (unit 46). Both species together form a well-differentiated and well-supported subclade (Figs 1, 2). Both species differ from each other by COI genetic of ≥ 21.8 % (Table S 4). Throughout its range, S. radians has been found in sympatry with three congeners. On the Riau Islands it occurs in sympatry with Siphonaria sipho; for comparative remarks refer to that species. Two congeners are sympatric at Kudat, Sabah, E. Malaysia. For comparisons with these species refer to comparative remarks under Siphonaria radiata and S. kudatensis sp. nov., respectively. Specimens from Malaysia and China Sea match the shell sculpture (wide to fine and wavy ribbing), external (grey / brown) and internal colouration (black and white rays on shell margin, typical for this species. A specimen figured as ‘ Siphonaria radians ’ in Habe & Kosuge (1966: pl. 42, figs 24, 25) is attributed to S. rucuana.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFCF824FFCCAFF02FCF1FDF6.taxon	distribution	Distribution and habitat. Currently known only from Riau Islands, Indonesia and Sabah, E Malaysia (Fig. 25). Found on rocky headlands, at upper and lower littoral levels.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFCE8249FCCAFDC2FEA1F896.taxon	description	(Figs 31 L – N, P – Q, V – W, 32 E – F)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFCE8249FCCAFDC2FEA1F896.taxon	materials_examined	Material examined. Type material. Lectotype of Siphonaria scabra Reeve, 1856, present designation, from Port Jackson, [Sydney], Australia (NHMUK 1981011 / 1, Fig. 31 L). Four paralectotypes, same data as lectotype (NHMUK 1981011 / 2 - 4, Figs 31 P – Q). Other, non-type material. Australia, Qld: Slade Point, Mackay, 21 ° 03.813 ’ S, 149 ° 13.527 ’ E, Q 14 - 1 (AM C. 585702 8 p, C. 585139 d [M 214], C. 585140 p [M 215], C. 585141 p [M 216]); Wreck Pt, Yeppoon, 23 ° 08.736 ’ S, 150 ° 45.865 ’ E, Q 08 - 4 (AM C. 585138 p [M 426]); Northwest Island, 23 ° 17.683 ’ S, 151 ° 42.997 ’ E, Q 07 - 3 (AM C. 585130 p [M 078]); Urangan Hervey Bay, 25 ° 17.504 ’ S, 152 ° 54.664 ’ E, Q 05 - 1 (AM C. 585342 p, C. 585127 p [M 403], C. 585128 p [M 182]); Drury Point Scarborough, 27 ° 12.168 ’ S, 153 ° 06.980 ’ E, Q 03 - 8 (AM C. 585748 1 p); NSW: Bolton Point Lake Macquarie, 33 ° 0.389 ’ S, 151 ° 36.889 ’ E, NSW 08 - 3 (AM C. 585448 12 p, AM C. 585060 p [SK 168], AM C. 585062 p [SK 169]); Laings Point Sydney Harbour, 33 ° 50.419 ’ S, 151 ° 16.638 ’ E, NSW 06 - 3 (AM C. 585898 7 p, C. 585061 p [M 152], C. 585114 p [M 154]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFCE8249FCCAFDC2FEA1F896.taxon	discussion	Taxonomic remarks. The syntype marked as ‘ type’ (Fig. 31 L) is herein designated as the lectotype of S. scabra for the stabilisation of the name (Art. 74.1 of the Code). Our delineation of this species is based on comparative analyses of the morpho-anatomy and mitochondrial genetics of freshly collected topotypes (Fig. 31 J) and geographic series of additional specimens (Table S 1). Several authors treated S. scabra as a junior synonym or variety of S. diemenensis, which is rejected herein. The record of ‘ S. scabra ’ from SA in Adcock (1893: 11) is probably a misidentification of S. zelandica and incorrectly synonymised with S. luzonica.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFCE8249FCCAFDC2FEA1F896.taxon	description	External morphology. Foot sole, foot wall, foot edge, mantle and cephalic folds all evenly cream, paler at foot edge; regular vertical black pigmentation bands / stripes on foot wall, short of foot edge; mantle narrow, strongly finely lobed, thickened at edge; pneumostomal lobe narrow, faint black pigmentation, under mantle behind right cephalic fold; closes the pneumostomal and anal openings at the mantle edge; two small black epithelial eye spots centralised on two centrally touching cephalic folds; genital pore inconspicuous, located on foot wall to right anterior of right cephalic fold. Shell (Figs 31 L – N, P – Q, V – W; Table S 9). Medium sized (max sl mean = 16.1 mm, SD = 3.4 mm, n = 7), elongate ovate; height medium; apex offset sightly posterior and to left, apical sides convex, protoconch direction homostrophic (n = 3; Fig. 31 V), shell whorl dextral; exterior sculpture variable, uneven, rough, growth striae layered leaving rib protrusions, uneven; radial colour bands, protoconch dark, central pale, edge with dark rib interstices; shell thick; rib count (mean = 37, SD = 6.8, n = 7), with ten primary ribs (Fig. 31 N), pale white to cream, wavy, broad, ridges flat to rounded, shell lip uneven ribs weakly protrude at shell edge; few (1 – 2) secondary ribs between and of similar width to primary ribs; rib interstices darker, width slightly less than rib width; siphonal ridge not prominent, formed by paired primary ribs. Interior shell margin dark chocolate brown with white rays under primary / secondary ribs extending to spatula, shell edge corrugated by rib ends, siphonal groove distinct; spatula dark chocolate brown, sometimes reddish, mottled to lighter tan; ADM scar distinct, deep, CMS straight.; thickening of shell lip occurs without whitening, infills and reduces lip scalloping; coarse and fine rib as well as prominent primary / secondary rib change (Fig. 31 N). Reproductive system (Fig. 32 E; n = 2): Positioned within coelom under the respiratory cavity, hermaphroditic glands positioned to posterior against right foot wall and over foot sole, epiphallic parts positioned to anterior between BM and RAM; merge of AO and GA indistinct, AO relatively large, elongated, bluntly pointed, twisted and centrally bent (without a prominent MA), slightly larger than GA, thicker than ED; ED relatively long, centrally bent, thick; EG white, folded, elongated; single flagellum F 1 on EG, long, twisted, thickish; AO, GA and ED all muscular white tissue; BD and CD with opposing connections to GA between ED, AO and singular GP; BD long, heavily looped on anterior side with prominent MA, much longer than CD of similar thickness, both ducts smooth and pass together through RAM connecting into MG (BD above CD), BC translucent, clear test, relatively large, bulbous, (0 to 1 SPM in BC’s of three specimens); HD with prominent brown markings, short, thickened, coiled, links AG to a small, elongated, narrow, brownish / yellow spotted, finely granulated HG, inner edge firmly moulded; MG and AG small, folded, soft white tissue; purple SV embedded on left side of AG, AG larger than HG, sides match curvature of inner foot wall at right posterior quarter of coelom. Spermatophore (Fig. 32 F): Body cylindrical, thread-like (length = 10.88 mm, n = 1), test thin, translucent; head section, bluntly rounded, containing a white gelatinous core, tapers to a thin flagellum and tip; both sections smooth, featureless; head shorter, thicker than flagellum (head length = 10.03 mm; 92 % of SPM length; flagellum length = 0.85 mm; head width = 93 μm; flagellum width = 13 μm). Single SPM tightly coiled in brown gelatinous mass in BC of topotypic specimen.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFCE8249FCCAFDC2FEA1F896.taxon	diagnosis	Comparative remarks. Siphonaria scabra (atra group, unit 50) is the sister species of S. pravitas sp. nov. (unit 51); both species together form a well-differentiated and well-supported sub-clade (Figs 1, 2). They differ from each other by COI distances of ≥ 8.2 % (Table S 4). Siphonaria scabra differs from any other species by COI distances of ≥ 24 % (Table S 4). Throughout its range, S. scabra has been found in sympatry with nine congeners. Five congeners are sympatric in eastern Australia: For comparisons with S. normalis, S. atra, S. denticulata refer to comparative remarks under these species. Siphonaria opposita has a lower shell with a dual-ribbed siphonal ridge, a larger pointed AO, larger BC, and a shorter ED. Four species occur in sympatry with S. scabra in Sydney Harbour: For comparison with S. diemenensis and S. zelandica refer to comparative remarks under these species. Siphonaria funiculata has a taller shell with a dual ribbed but indistinct siphonal ridge, less raised ribbing and weaker edge scalloping, a shorter ED and AO, larger BC, and a shorter, drop-like SPM. Siphonaria pravitas sp. nov. has a lower shell with stronger raised ribs and edge scalloping, a smaller AO, shorter ED, larger BC, a shorter wider BD with no distal loop, and a shorter SPM. Specimens figured as ‘ S. diemenensis var. scabra ’ from ‘ Port Jackson [Sydney Harbour]) in Hubendick (1946: pl. 2, figs 14 – 15) are likely specimens of S. denticulata based on patterning on shell lip and margin and only possibly of S. scabra based on the presence of few secondary and even primary ribs, yet clearly not of S. diemenensis.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFCE8249FCCAFDC2FEA1F896.taxon	distribution	Distribution and habitat. Endemic to subtropical to temperate east coast of Australia (Fig. 25). In this study found on sheltered rocky shores across upper to mid littoral levels.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFC88245FF68F822FBCEF936.taxon	description	(Figs 33 A – H, O – P, S – U, 34 A – D)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFC88245FF68F822FBCEF936.taxon	materials_examined	Material examined. Type material. Lectotype of Siphonaria funiculata Reeve, 1856, present designation, from Tasmania [Australia] (NHMUK MC. 197927 / 1, Fig. 33 A). Three paralectotypes, same data as lectotype (NHMUK MC. 197927 / 2 - 4). Holotype of Siphonaria blainvillei Hanley, 1858 (NHMUK 1907.10.28.90, Fig. 33 D). Holotype of Siphonaria virgulata Hedley, 1915, from Terrigal [E of Gosford, NSW], Sydney; coll. C. Hedley, 1915 (AM C. 39858, Fig. 33 C). Twenty paratypes, same data as holotype (AM C. 337311). Holotype of Siphonaria oblivirgulata Hubendick, 1945, from Port Jackson [Sydney, Australia]; coll. Eugenie Exp., 1851 – 1853 (UUZM 1575, Fig. 33 E). Other, non-type material. Australia: NSW: Brunswick Heads, 28 ° 32.297 ’ S, 153 ° 33.444 ’ E, NSW 12 - 1 (AM C. 585592 4 p, C. 585068 p [M 185]); Cape Byron, 28 ° 38 ’ S, 153 ° 38 ’ E (AM C. 311682 5 p); Sand Point, N of Ballina, 28 ° 50.66 ’ S, 153 ° 36.45 ’ E (AM C. 343608 p); MinniE, WAter, E of Grafton 29 ° 46.6 ’ S, 153 ° 18 ’ E (AM C. 343604 p); N of Coffs Harbour, 30 ° 14 ’ S, 153 ° 9 ’ E (AM C. 311683 7 p); Nambucca Heads, 30 ° 38.5 ’ S, 153 ° 1 ’ E AM (AM C. 311681 7 p); Fingal Bay nr Port Stephens, 32 ° 44.990 ’ S, 152 ° 10.481 ’, E, NSW 09 - 1 (AM C. 585335 p); Catherine Hill Bay, 33 ° 9.3 ’ S, 151 ° 38 ’ E (AM C. 343605 3 p); S end Catherine Hill Bay, 33 ° 9.5 ’ S, 151 ° 38 ’ E (AM C. 311679 6 p); Broken Head Terrigal, 33 ° 26.796 ’ S, 151 ° 27.030 ’ E, NSW 08 - 1 (AM C. 585665 6 p, C. 585051 p [M 044]); Terrigal The Skillion, 33 ° 27.008 ’ S, 151 ° 27.122 ’ E, NSW 08 - 2 (AM C. 585405 10 + p, C. 585055 p [M 223], C. 585056 p [M 224]). Sydney, Long Reef Collaroy, 33 ° 44.6 ’ S, 151 ° 18.6 ’ E (AM C. 343674 5 p); S side Long Reef Collaroy, 33 ° 44.7 ’ S, 151 ° 19 ’ E (AM C. 343678 3 p; AM C. 446108 6 p); Fairlight, North Harbour, 33 ° 47.986 ’ S, 151 ° 16.837 ’ E, NSW 06 - 1 (AM C. 585470 16 p); Shelly Beach Headland Manly, 33 ° 48 ’ S, 151 ° 17.5 ’ E (AM C. 311680 5 p); North Harbour, SE side Reef Bay, 33 ° 48.5 ’ S, 151 ° 16.38 ’ E (AM C. 311687 7 p). Middle Harbour, Wy-ar-gine Point, 33 ° 49 ’ S, 151 ° 15 ’ E (AM C. 311678 10 + p); Wy-ar-gine Point, 33 ° 49.159 ’ S, 151 ° 15.195 ’ E, NSW 06 - 5 (AM C. 585664 6 p, 3 d); Edwards Beach Balmoral, 33 ° 49.38 ’ S, 151 ° 15 ’ E (AM C. 311890 10 + p); Balmoral, 33 ° 49.7 ’ S, 151 ° 15.02 ’ E (AM C. 343610 2 p), Laings Point Sydney Harbour, 33 ° 50.419 ’ S, 151 ° 16.638 ’ E, NSW 06 - 3 (AM C. 585475 17 p, C. 585035 p [M 162, SK 035], C. 585036 p [M 163], C. 585037 d [M 164]); Tamarama Point, 33 ° 54 ’ S, 151 ° 16 ’ E (AM C. 311684 7 p; AM C. 343615 2 p). Bombo Kiama, 34 ° 39.232 ’ S, 150 ° 51.649 ’ E, NSW 03 - 1 (AM C. 585455 13 p, C. 584884 p [SK 384], C. 585280 p [SK 048]); Ulladulla, Wardens Head, 35 ° 21 ’ S, 150 ° 29 ’ E (AM C. 311677 7 p); Batemans Bay, Batehaven, 35 ° 44 ’ S, 150 ° 12.5 ’ E (AM C. 311685 5 p); Wimbie Beach, 35 ° 47 ’ S, 150 ° 14 ’ E (AM C. 343613 2 p); Eurobodalla Shire, Pretty Point SE facing, 35 ° 48.28 ’ S, 150 ° 14 ’ E (AM C. 343611 p); Burrewarra Point, 35 ° 50 ’ S, 150 ° 13.5 ’ E (AM C. 343606 3 p); Mullimburra Point SE facing, 35 ° 59.75 ’ S, 150 ° 9.58 ’ E (AM C. 343607 5 p); Murunna Point Camel Head, 36 ° 22.720 ’ S, 150 ° 04.766 ’ E, NSW 02 - 1 (AM C. 585718 9 p, C. 585030 p [SK 025], C. 585332, d [SK 026]); Bermagui, 36 ° 25.18 ’ S, 150 ° 3.78 ’ E (AM C. 343612 2 p); Wapengo Lagoon estuary S side near Bithry Inlet, 36 ° 37.7 ’ S, 150 ° 59 ’ E (AM C. 395918 8 p); Aslings Beach, N end Twofold Bay, 37 ° 3.1 ’ S, 149 ° 55 ’ E (AM C. 148856 p); Oman Point Eden, 37 ° 04.634 ’ S, 149 ° 53.445 ’ E, NSW 01 - 1 (AM C. 585629 5 p). Twofold Bay, Murrumbulga Pt, 37 ° 4.75 ’ S, 149 ° 53.06 ’ E (AM C. 343622 2 p, C. 343623 p, C. 343624 p, AM C. 311688 4 p); Red Point, 37 ° 6.083 ’ S, 149 ° 57.1 ’ E (AM C. 343625 p); Munganno Point, 37 ° 6.2 ’ S, 149 ° 55.48 ’ E (AM C. 343616 p, C. 311689 4 p, C. 343617 3 p, C. 343618 2 p, C. 343619 3 p, C. 343621 3 p, C. 150599 p); Fisheries Beach, 37 ° 6.78 ’ S, 149 ° 55.6 ’ E (AM C. 343620 p). Green Cape, 37 ° 15.8 ’ S, 150 ° 3 ’ E (AM C. 343614 p, C. 311676 5 p); Wonboyn Beach, Disaster Bay, 37 ° 16 ’ S, 149 ° 57 ’ E (AM C. 343733 2 p); Nadgee Fauna Reserve, N of Little River, 37 ° 24 ’ S, 149 ° 57 ’ E (AM C. 343735 3 p). Vic: Bastion Head Mallacoota, 37 ° 34.429 ’ S, 149 ° 45.927 ’ E, V 09 - 1 (AM C. 585459 13 p); Cape Conran, 37 ° 48.798 ’ S, 148 ° 43.608 ’ E, V 08 - 2 (AM C. 585610 4 p); Wilsons Promontory, 39 ° S, 146 ° 22 ’ E (AM C. 311674, 3 p); Bear Gully, 38 ° 53.519 ’ S, 145 ° 59.029 ’ E, V 07 - 3 (AM C. 585652 5 p); Flat Rock, Inverloch, 38 ° 38.877 ’ S, 145 ° 41.638 ’ E, V 07 - 6 (AM C. 585573 3 p); Blow Hole, Western Port Bay nr. Flinders, 38 ° 29 ’ S, 145 ° 1 ’ E (AM C. 311673 2 p); Cape Schanck, 38 ° 29.951 ’ S, 144 ° 53.369 ’ E, V 06 - 4 (AM C. 585515 p [SK 548]); Port Phillip, Portsea, 38 ° 19 ’ S, 144 ° 43 ’ E (AM C. 311675 2 p); Point Lonsdale (nr Queenscliff), 38 ° 17.276 ’ S, 144 ° 36.977 ’ E, V 05 - 1 (AM C. 585514 2 p); Roadknight Point, 38 ° 25.707 ’ S, 144 ° 11.102 ’ E, V 04 - 1 (AM C. 585457 13 p); Loutit Bay Lorne, 38 ° 31.190 ’ S, 143 ° 59.429 ’ E, V 03 - 2 (AM C. 585465 15 p). Tas: Flinders Island, Northeast River, 39 ° 43.8 ’ S, 147 ° 57.6 ’ E (TMAGE 542122 d); PalanaBeach, 39 ° 45.6 ’ S, 147 ° 52.8 ’ E (TMAG E 54211 d); Port Davies & Cave Beach, 40 ° 0.6 ’ S, 147 ° 52.8 ’ E (TMAG E 27150 d; TMAG E 27142 4 d). Boat Harbour, 40 ° 57 ’ S, 145 ° 38 ’ E (AM C. 311672 3 p); West Point, 40 ° 57 ’ S, 144 ° 36 ’ E AM (AM C. 311668 3 p); Little Peggs Beach, 40 ° 51 ’ S, 145 ° 21.6 ’ E (TMAG E 41987 d); Rocky Cape: Picnic Beach & rocks to S, 40 ° 52.2 ’ S, 145 ° 28.8 ’ E (TMAG E 41989 d); S of Granite Point Bridport, 40 ° 59.739 ’ S, 147 ° 23.468 ’ E, T 01 - 1 (AM C. 585251 p [M 174]); Bridport: beach, 41 ° 24.6 ’ S, 147 ° 23.4 ’ E (TMAG E 41986 d); Bellingham, 41 ° 0.6 ’ S, 147 ° 9.6 ’ E (TMAG E 54215 4 d); Greens Beach, 41 ° 4.8 ’ S, 146 ° 45 ’ E (TMAG E 54216 d). The Gardens, Seatons Cove, 41 ° 12.6 ’ S, 148 ° 16.8 ’ E (TMAG E 41988 3 d); Swimcart Beach, 41 ° 13.8 ’ S, 148 ° 17.4 ’ E (TMAG E 27411 4 d, TMAG E 54214 5 d, TMAG E 41975 3 d). Beaumaris, Shelly Point, 41 ° 26.4 ’ S, 148 ° 16.8 ’ E (TMAG E 41981 d); Steels Beach, 41 ° 28.2 ’ S, 148 ° 16.2 ’ E (TMAG E 54217 d); Bicheno, 41 ° 52.837 ’ S, 148 ° 18.525 ’ E, T 02 - 1 (AM C. 585693 7 p); Bicheno, S end Redbill Beach, 41 ° 53 ’ S, 148 ° 18 ’ E (AM C. 311669 2 p); Maria Island, Darlington Bay, 42 ° 34.8 ’ S, 148 ° 3.6 ’ E (TMAG E 41984 3 d); Dodges Ferry, 42 ° 51.083 ’ S, 147 ° 36.981 ’ E, T 03 - 1 (AM C. 585462 14 p); Park Beach Dodges Ferry, 42 ° 51.716 ’ S, 147 ° 36.665 ’ E, T 03 - 4 (AM C. 584883 p [SK 138], C. 585660 p [M 111], C. 585262 p [SK 139]); Lagoon Bch (near Saltwater River), 42 ° 56.903 ’ S, 147 ° 39.962 ’ E, T 03 - 2 (AM C. 585648 5 p, C. 595922 p [SK 551]); Blackmans Bay, 43 ° 0.6 ’ S, 147 ° 19.8 ’ E (TMAG E 15879 10 + d); Tasman Arch, 43 ° 02.033 ’ S, 147 ° 56.963 ’ E, T 03 - 3 (AM C. 585772 12 p, C. 585258 p [M 116]); Huon Point d’Entrecasteaux Channel, 43 ° 17.471 ’ S, 147 ° 05.778 ’ E, T 04 - 1 (AM C. 585678 6 p, C. 585267 p [M 106], C. 585268 p [M 135]); South Bruny Island: Coal Point, 43 ° 19.8 ’ S, 147 ° 19.8 ’ E (TMAG E 35240 p); Moss Glen, 43 ° 31.910 ’ S, 146 ° 53.641 ’ E, T 05 - 1 (AM C. 585569 3 p); Flensing Rock, 43 ° 34.291 ’ S, 146 ° 54.856 ’ E, T 05 - 2 (AM C. 585605 4 p). King Island, Little Porky Beach, 39 ° 51 ’ S, 143 ° 51.6 ’ E (TMAG E 41980 2 d), Naracoopa, foreshore, 39 ° 55.2 ’ S, 144 ° 7.2 ’ E (TMAG E 41978 d), Currie Harbour, 39 ° 55.8 ’ S, 143 ° 50.4 ’ E (TMAG E 41985 10 d), Red Hut Point, 40 ° 6 ’ S, 144 ° 6 ’ E (TMAG E 41983 d).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFC88245FF68F822FBCEF936.taxon	discussion	Taxonomic remarks. The lectotype of S. funiculata has been referred to as the holotype by Jenkins (1981: 2), an act that qualifies as the designation of the lectotype. The lectotype (Fig. 33 A) closely matches the figure in the original description (Reeve, 1856: pl. 2, fig. 6 a – b). Reeve (1856: pl. 7, fig. 35 a – b) erroneously labelled species 35 (= S. lirata) as ‘ S. funiculata ’ (see erratum in the appendix). Our delineation of this species is based on comparative analyses of the morpho-anatomy and mitochondrial genetics of freshly collected topotypes of S. funiculata (Fig. 33 B), S. blainvillei (Fig. 33 P), S. virgulata (Figs 33 H, O), and S. oblivirgulata (Fig. 33 F) and a geographic series of additional specimens (Table S 1). Tenison Woods (1877: 58) incorrectly treated S. funiculata as a variety of S. diemenensis. Hutton (1878: 42) treated S. funiculata as a synonym of S. laeviuscula, but Hubendick (1946: 23) thought that Hutton’s (1878: 42) reference to S. laeviuscula was a misidentification of S. funiculata. Hedley (1915: 751) described S. virgulata indicating that Angas’ (1867: 232) record of ‘ S. funiculata ’ was in fact a misidentification of this new species. This conclusion was subsequently upheld by Hedley (1917 b) and Hubendick (1946: 23), the latter also including a record of ‘ S. funiculata ’ by Adam & Leloup (1939: pl. 2, fig. 2 a – b) from Pisang Island, PNG. Hedley (1915) and Jenkins (1981: pl. 1, fig. a) observed that S. blainvillei (based on one specimen) was a tall and broad ribbed specimen of S. funiculata. Hubendick (1946: 23) also treated S. blainvillei as a synonym of S. funiculata. Several tall specimens of S. funiculata have been collected as part of this work with a shell geometry resembling the type specimen of S. blainvillei and analyses of these specimens confirm this conclusion (e. g., AM C. 585660, AM C. 585251). Similarly tall individuals occur in other Siphonaria species, such as S. radiata, S. plicata, S. radians, S. sipho and S. monticulus. Iredale (1924: 275 – 276) and Jenkins (1981) considered S. virgulata as a geographical variant of S. funiculata. Hubendick (1947 b: 1) incorrectly considered S. funiculata Reeve 1856 as a synonym of S. pisangensis Hubendick 1947 b (type locality Pisang Island, PNG). Grove et al. (2006: 60) correctly listed S. blainvillei, S. virgulata and S. oblivirgulata in the synonymy of S. funiculata. No records of S. funiculata are available from NZ based on our examinations of museum samples. The description below is based on the redescription in Jenkins (1981), which is expanded here for completeness and consistency.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFC88245FF68F822FBCEF936.taxon	description	External morphology (Fig. 33 T). Foot sole smooth, greyish yellow centrally, fading to become paler at foot edge; foot wall dark grey with evenly spread white subepithelial pustules becoming more vivid and dense close to the foot sole; genital pore inconspicuous, located on foot wall posterior to right cephalic fold; two small black epithelial eye spots centralised on two thick centrally touching dark grey to blueish cephalic folds, folds fade to a paler cream to yellowish at outer edge, covered with clustered white mucous cells similar (but smaller) to those of the foot wall tissue; fringing whitish mantle around top of foot wall, extends and increasingly transparent to shell edge, mantle edge thickened, lobed with an outer band of cream and brown pigmentation reflecting corrugations and inner colouration of shell lip and ribs; thin whitish pneumostomal lobe part of the mantle, between the right ADMs, closes the pneumostome and anus at the mantle edge. Shell (Fig. 33 A – H, O, P, U; Table S 9). Small to medium sized (max sl mean = 23.3 mm, SD = 4.45 mm, n = 30); height medium to tall; shell thickness medium; ovate; apical sides weakly convex, apex often eroded creating a white spot; protoconch direction homostrophic (n = 3; Fig. 33 U), shell whorl dextral; apex offset weakly posterior and central; exterior sculpture finely costate with irregularly spaced radial ribs and growth striae; ribbing flat, broad with rib widths at shell lip ranging from 0.39 to 1.89 mm (mean = 0.82 mm, SD = 0.02 mm, n = 30), white axial ribs, often bistriate, chocolate coloured interstices, narrow and curve adapically; number of ribs variable, rib count (range 28 – 94; mean = 63, SD = 10.5, n = 60); siphonal ridge is weakly visible with fine, clustered, brown radial striae above a slight fold in the marginal lip; interior is polished and purplish brown with a white to blue spatula (colouration extends into the shallow siphonal groove) fading to a chocolate brown zone above the brown ADM impression and tan margin; marginal lip is shallowly scalloped with alternating chocolate brown and white radial markings, restricted to the lip margin and reflecting the exterior ribbing; ADM impression is horseshoe-shaped with a thin, lightly convex, anterior attachment area and a bare siphonal groove flanked by broad, ovate muscle impressions. Juvenile specimens have fine radial ribs, a dark brown interior and exterior (often obvious around the apex of adult specimens), with a white spatula. Reproductive system (Figs 34 A, B, D; n = 3). Epiphallic parts fill the region between RAM and BM, ED, GA, AO, EG white, smoothly rounded, possessing thick fibrous layers of tissue, ED larger than EG, often elongated; F 1 very stubby, inconspicuous; GA large, bulbous, opening below the mantle on the side of the foot, behind right cephalic fold, anterior to pneumostome; AO very large, sack-like, joins GA; ED very short thick, joins to side of GA, EG larger than AO or GA; BD and CD enter GA very close together and AO, both pass through RAM (BD above CD) are of similar length (although BD may be shorter); HG at posterior right quarter of coelom over the foot muscle tissue; HG usually yellow, granulated, linked by thin duct to the pinkish white, lobed, coiled HD, which in turn links to CD; SV partly lobed, uncoiled, pink to white, connected via thin duct alongside the AG to CD; AG / MG complex yellow to white, folded, lobed, closely attached to HG; SV embedded in folds; BC ovate, brown, patterned, tissue often expanded and stretched or collapsed and wrinkled; test thin, enclosing granulated, brown gelatinous mass. Spermatophore (Fig. 34 C). Elongated drop shaped, test thin, smooth, featureless, translucent (length = 1.12 mm, n = 1), head spherical; flagellum very short; both sections smooth, featureless; head much larger than flagellum (head length = 0.93 mm, head ~ 83 % length of SPM, head width = 0.79 mm, flagellum width = 0.103 mm, n = 1); single SPM found in one BC (AM C. 584848). SPM matches SPM depicted in Jenkins (1983: fig. 3 f). Radula and jaw (figured in Jenkins 1983: 10, pl. 3 a – h). Radula with a central tooth and longitudinally variable number of inner, mid and outer lateral teeth in longitudinal rows. Mean dentition formula 43: 1: 43 (SD = 7.9, n = 17) with around 120 transverse rows (SD = 14.9). These rows are parallel and slightly curved (anteriorly convex) Jenkins (1981: 9, pl. 3 a – h). Of the 43 half row laterals, 7 (SD = 4.2) are inner, 17 (SD = 7.6) mid and 18 (SD = 3.1) outer lateral teeth, respectively. The total number of lateral teeth appears related to the length of the shell (max. 54: 1: 54, shell length = 22.0 mm, mean 43: 1: 43, shell length = 18.1 mm, min 34: 1: 34, shell length = 15.1 mm). However, the numbers of inner, mid and outer lateral teeth vary independently of animal dimensions and distributions. All teeth are bluntly concave posteriorly. The central tooth is narrow and weakly bicuspidate (often pointed) with a lower profile than the flanking laterals. The base is broad with adjacent central teeth. Mid and inner lateral teeth interlock with posteriorly and anteriorly aligned laterals. Outer laterals do not interlock between transverse rows. The space between rows increases to the ribbon edges associated with a gradual decrease in tooth size (Jenkins 1981: 10, pl. 3 c, g). The space varies between individuals (Jenkins, 1981: 10, pl. 3 d, h) as well as posterior and anterior areas of the ribbon. All lateral teeth are broad based and bicuspidate on the mesocone with a longer inner cusp. Outer lateral teeth are often multicuspidate. Increasing side denticle numbers, less elongated shape and increasingly stunted mesocones are transverse row features less accentuated from the central to the outer lateral teeth. Inner lateral teeth are elongated without flanking endo and ecto cones (inner and outer side denticles respectively). The more numerous outer lateral teeth have both ecto and endo cones while the mid laterals possess only an ectocone. The angle of separation from the mesocone of these side denticles is widely variable. Both side cones curve either towards or away from the mesocone (Jenkins, 1981: 10, pl. 3 g, h). The length and width of the separation cleft is also widely variable, both generally increase towards the ribbon edge (Jenkins, 1981: 10, pl. 3 c, d mid half ribbon area, pl. 3 g, h ribbon edge). Aberrant outer lateral teeth are often present on both sides of the ribbon appearing as fused teeth with double mesocones. Not all individuals have inner lateral teeth, most have increased numbers of mid laterals. The number is independent of the number of lateral teeth, for example, of two radulae with 54 half row laterals, one had no inner laterals while the other had 18. The same variability was noted for radulae with fewer numbers of lateral teeth. Inner laterals do not possess endo or ectocones.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFC88245FF68F822FBCEF936.taxon	diagnosis	Comparative remarks. Siphonaria funiculata (lateralis group, unit 8) is the sister species of S. lessonii, both together representing the sister lineage of S. tasmanica (Figs 1, 4). Siphonaria funiculata differs from S. lessonii by COI distances of ≥ 12 % and from S. tasmanica by ≥ 8.5 % (Table S 8). Throughout its range, S. funiculata has been found in sympatry with nine congeners. For comparisons with S. diemenensis, S. denticulata, S. scabra, and S. zelandica refer to comparative remarks under these species. Siphonaria emergens has a much smaller, paler, orange-brown shell with less prominent ribbing, stronger edge scalloping and a strongly offset apex. Siphonaria pravitas sp. nov. has much paler brown shell with a more prominent siphonal ridge, raised ribbing, a darker spatula, a smaller AO, and a narrower, thread-like SPM. Siphonaria stowae has a much smaller, paler, yellowish cream shell with less prominent ribbing, stronger edge scalloping, a strongly offset apex, a smaller AO and BC, a shorter ED, and a narrower, thread-like SPM. Siphonaria jeanae has a smaller, grey-blue shell with slightly more raised brown ribbing, purplish spatula, an indistinct AO, narrower BD, and a more bulbous SPM. Siphonaria tasmanica has a grey-blue shell with a less distinct siphonal ridge, fainter ribbing, a smaller AO and BC. A record of S. funiculata from NZ (Hutton, 1873: 55) has been attributed subsequently to S. australis (Jenkins, 1983: 13). Based on similarity in reproductive anatomy (i. e., closeness of the duct joint, smallness of the genital atrium and greatly swollen epiphallus duct), Hubendick (1946: 23) assigned S. funiculata to Pachysiphonaria. The RS shown herein (Figs 34 A, B, D) correspond well with illustrations of the RS figures of S. funiculata elsewhere (e. g., Hubendick 1945: figs 3, 12; 1946: fig. 5, as ‘ S. virgulata ’; 1955: figs 1 – 2, as ‘ S. virgulata ’; Jenkins, 1983: figs 3 a – c). The mean radula dentition formula observed herein is consistent with the 39: 1: 39 count given by Hubendick (1946: 23). A specimen figured as ‘ S. funiculata ’ from Lakshadweep, India by Ravinesh & Biju Kumar (2015: 38) is a misidentification of an unidentified species. Specimens depicted as ‘ S. funiculata ’ from Kelsi Coast, India in Vakani & Rahul Kundu (2021: 134, figs 2 d, 3 d) are misidentifications and are likely specimens of S. incerta sp. nov.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFC88245FF68F822FBCEF936.taxon	distribution	Distribution and habitat. Endemic to eastern and southeastern coasts of Australia, from Brunswick Heads, northern NSW, south to west of Lorne, Vic, including Tas (Fig. 25). In this study found to be common in sheltered places, such as crevices and vertical faces, on exposed rocky shores, upper to mid littoral levels, often associated with barnacles (Fig. 33 S).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFC48240FCCAF882FE9AF896.taxon	description	(Figs 33 I – N, Q – R, V, 34 E – G)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFC48240FCCAF882FE9AF896.taxon	materials_examined	Material examined. Type material. Lectotype of Siphonaria kurracheensis Reeve, 1856, present designation, from Kurrachee [Karachi], Scinde, [Pakistan] (NHMUK 1979167 / 1, Fig. 33 I). Three paralectotypes, same data as lectotype (NHMUK 1979167 / 2 - 4, Fig. 33 J – L). Other, non-type material. Pakistan: Karachi, Bubiji Beach, 24 ° 53 ’ N, 67 ° 01 ’ E (WAM S 72336 p, WAM S 74134 p [SK 148]); French Beach, 24 ° 50.367 ’ N, 66 ° 49.387 ’ E PA 01 - 1 (AM C. 585741 4 p, C. 585848 p [M 240], C. 585849 p [M 241], C. 585850 p [M 484], C. 585852 p [M 485, SK 304]); Clifton Beach, 24 ° 45.500 ’ N, 67 ° 05.968 ’ E PA 02 - 1 (AM C. 585107 p [SK 008], C. 585744 p [SK 045], C. 585856 p [M 229], C. 585857 p [M 230], C. 585858 p [M 231]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFC48240FCCAF882FE9AF896.taxon	discussion	Taxonomic remarks. The largest syntype with clearest external sculpture (Fig. 33 I) is herein designated as the lectotype of S. kurracheensis for the stabilisation of the name (Art. 74.1 of the Code). The shell figure in Reeve’s (1856) original description provides a ventral view only. Our delineation of this species is based on comparative analyses of the morpho-anatomy and mitochondrial genetics of freshly collected topotypes (Fig. 33 N, M, Q) and geographic series of additional specimens (Table S 1). We do not accept Hubendick’s (1946) extended concept of S. kurracheensis, which included several extralimital nominal taxa (e. g., S. belcheri, S. depressa, S. luzonica, S. savignyi, S. siquijorensis and S. zebra), all of which represent distinct species. Therefore, the accepted distribution of this species is much smaller than previously stated. Morrison (1972: 56 – 58) synonymized S. kurracheensis with S. laciniosa based on similarity in shell form and a ‘ common reproductive development’. This synonymy is rejected herein based on examination of types and topotypes. The species identified as ‘ S. kurracheensis ’ in WA by various authors is shown herein to be morphologically and genetically distinct and is described as a new species, Siphonaria restis sp. nov. (unit 54) below.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFC48240FCCAF882FE9AF896.taxon	description	External morphology. External parts evenly pale cream without black pigmentation apart from faint shading at centre of cephalic folds; mantle weakly lobed, translucent, narrower than foot wall, covers exposed inner shell lip; genital pore inconspicuous, located on foot wall to right anterior of right cephalic fold; single small black epithelial eye spots centralised on two centrally touching cephalic folds; pneumostomal lobe under the mantle, unpigmented, between the right anterior and right posterior ADMs. Shell (Fig. 33 I – N, Q, R; Table S 9). Small sized (max sl mean = 11.5 mm, SD = 1.7 mm, n = 7), ovate, height low to medium; shell thickness thin to medium, apex offset weakly posterior and left, apical sides weakly convex, protoconch direction undetermined, shell whorl dextral; growth striae indistinct; rib count (mean = 42, SD = 4, n = 7), primary ribs pale white, fairly straight, unraised, flat, ridge rounded; secondary ribs even whitish; ribs increasingly widen to shell lip; more primary than secondary ribs; rib interstices very narrow brown, marking on side of primary ribs; siphonal ridge low prominent; shell lip uneven, weakly scalloped, corrugated by primary ribs. Interior mottled red brown blotches on margin; spatula golden to pale brown / whitish; clearly demarked from margin by siphonal groove similar colour paler fairly prominent; irregular narrow brown rays from shell lip to spatula; brown interstices show through shell lip; ADM scar indistinct, CMS convex. Smaller and tall specimens with thicker shells, tend to show dark brown rays on shell margin. Thickening of shell margin and variability in shell height occurs in larger specimens in this species. Reproductive system (Figs 34 E, F; n = 3). Positioned within coelom under the respiratory cavity, hermaphroditic glands positioned to posterior against right foot wall and over foot sole, epiphallic parts positioned between BM and to side of RAM; AO medium, broad, tip bluntly hooked, merges with MA, joins to upper end of small GA; ED very short, centrally twisted, bent, joins to side of GA; EG folded may be large; single wide looped flagellum F 1 joins as an extension of broader ED; AO, GA and ED all muscular white tissue; BD and CD with opposing connections (bulbous at CD) into GA between ED, AO and GP; BD longer and narrower than CD with a prominent distal loop, top of loop attached via a long MA to inner foot wall in front of BM; both BD and CD smooth, similar thickness and pass together through RAM connecting into folds of MG (BD above CD), BC small, translucent test and bulbous drop shaped; HD brownish, narrow coils, links AG to a small elongated narrow brownish granulated HG; dark SV embedded within AG / MG; MG and AG small folded soft white tissue, similar size, sides match curvature of inner foot wall at right posterior quarter of coelom. Spermatophore (Fig. 34 G). Thread-like, test thin, translucent (length = 7.2 mm, n = 1, AL = 9 mm); head section cylindrical, centrally coiled, rounded tip; test thin, smooth, featureless, translucent, contains a white core, tapers into flagellum; head longer, wider than translucent flagellum (head length = 5.5 mm, ~ 76 % of total length, head width = 172 μm, flagellum width = 17 μm, n = 1); SPM tightly coiled in yellowish gel in BC of one specimen.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFC48240FCCAF882FE9AF896.taxon	diagnosis	Comparative remarks. In our mitochondrial phylogeny (Figs 1 – 4), S. kurracheensis (atra group, unit 28) forms a clade with S. alternata from the Caribbean and Gulf of Mexico as well as S. striata sp. nov. from Madagascar (Figs 1, 2). These species differ from S. kurracheensis by COI distances of ≥ 7.1 % (S. alternata, unit 29) and ≥ 8.6 % (S. striata sp. nov., unit 74) (Table S 5). For a comparison with S. striata sp. nov. refer to comparative remarks under that species. Siphonaria kurracheensis occurs in sympatry with four congeners in Karachi, Pakistan. Siphonaria asghar has weaker, finer edge scalloping, less prominent siphonal groove, darker interior, a smaller AO, a shorter, wider DB, and a larger BC. Siphonaria belcheri has a darker exterior and interior, a larger, wider AO, a longer, narrower BD, and a larger F 1. Siphonaria perexigua sp. nov. has a smaller, taller shell with less raised ribbing, a smaller AO, larger BC, and longer, wider ED. For comparison with Siphonaria crenata refer to comparative remarks under that species. Shell sculpture and geometry of S. kurracheensis resembles that of S. zelandica (flat, fine ribs); however, this resemblance is convergent since both species are not closely related with each other. Both species have non-overlapping distributions. A record of ‘ S. kurracheensis ’ from SA in Adcock (1893: 11, in synonymy of S. luzonica) is based on a misidentification of S. zelandica. A record of ‘ S. kurracheensis ’ from the Suez Canal in Moazzo (1939: 280, fig. 2) is a misidentification of S. belcheri (also noticed by Hubendick 1946: 54), the shell closely resembles the weakly ribbed paralectotype of S. belcheri. SPM and RS of ‘ S. kurracheensis ’ and ‘ S. kurracheensis var. siquiorensis ’ depicted in Hubendick (1945: figs 51 – 52) are here attributed to S. japonica and probably S. javanica, respectively. Hubendick (1946: 54) treated five taxa as varieties of S. kurracheensis (i. e., S. savignyi, S. luzonica, S. zebra, S. belcheri, S. depressa and S. siquijorensis). None of these treatments are accepted herein and these taxa are all removed from the concept of S. kurracheensis as delineated herein. Hence, the distribution of S. kurracheensis outlined in Hubendick (1946: 54), which includes locations well beyond Karachi and Persian Gulf (i. e., Java Sea, Philippines, Qld, and Port Jackson), is also disputed. The SPM depicted herein differs from that of ‘ S. kurracheensis ’ figured in Hubendick (1945: 31, fig. 51) and reproduced by Berry (1977: fig. 19) by being longer and without barbs on flagellum. A specimen figured as S. kurracheensis from the Suez Canal (Barash & Danin 1972: fig. 14) is S. savignyi. Dayrat et al. (2014: 261, fig. 5 E) misidentified unit 28 as S. savignyi; it is in fact S. kurracheensis based on the clustering with sequences of that species. Similarly, they misidentified unit 27 as S. kurracheensis, but the correct identification is S. belcheri. Specimens figured as ‘ S. kurracheensis ’ from Mubarak Village, Karachi (Bosch et al. 1995: fig. 863; Ali et al. 2011: fig. 1 B) and from Dwarka Coast, Gujarat (Vakani & Rahul Kundu 2021: figs 2 d, 3 d) are misidentifications and are likely specimens of S. crenata based on size, ribbing and extension of siphonal ridge.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFC48240FCCAF882FE9AF896.taxon	distribution	Distribution and habitat. Recorded from Muscat, Gulf of Oman, Karachi, Pakistan and Gujarat, India (Fig. 25). In the present study found in sheltered positions on exposed rocky intertidal marine shores, upper to mid littoral level.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFC18241FF68F822FB7DFAB6.taxon	description	(Figs 35 A – F, O – P, S, 36 A – C)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFC18241FF68F822FB7DFAB6.taxon	materials_examined	Material examined. Type material. Lectotype of Siphonaria bifurcata Reeve, 1856, present designation, from ‘ Philippine Islands’ (NHMUK 1979169 / 1, Fig. 35 A). Three paralectotypes, same data as lectotype (NHMUK 1979169 / 2 - 4, Figs 35 B – D). Other, non-type material. Philippines: NW Polillo Is, E Quezon, Bolunga District, nr Panukalan, 14 ° 59 ’ N, 121 ° 49 ’ E (WAM S 74096 p [SK 073], WAM S 74098 p [SK 410, protoconch H 8], WAM S 113801 p [SK 411], WAM S 74097 p [SK 412]); Cebu, Mactan Point, 10 ° 20.014 ’ N, 124 ° 02.723 ’ E, PHS 04 - 2 (AM C. 585118 p [M 414, SK 097]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFC18241FF68F822FB7DFAB6.taxon	discussion	Taxonomic remarks. The largest syntype (Fig. 35 A) is herein designated as the lectotype of S. bifurcata for the stabilisation of the name (Art. 74.1 of the Code). This specimen corresponds well with the specimen erroneously figured by Reeve (1856, pl. 3, fig. 21) as ‘ S. zebra ’. Inspection of the types of S. zebra reveal that this is a nominal species from eastern Australia, which is here synonymized with S. zelandica. Correspondingly, the types of S. bifurcata are not of an Australian species but are conspecific with freshly collected specimens from the Philippines. Therefore, we conclude that the original descriptions of these two species have accidentally been mixed up: The text of the description for species 21 (labelled as ‘ S. zebra ’) applies to the shell depicted in figure 22 and is based on the type material originally labelled as ‘ S. bifurcata ’ from the Philippines. In turn, the description of species 22 (labelled as ‘ S. bifurcata ’) applies to figure 21, which corresponds well with the type material of S. zebra from eastern Australia (= S. zelandica; Jenkins, 1983: 28; White & Dayrat, 2012: 61). Hence, the description of ‘ S. zebra ’ (including the type locality but excluding the figure) is herein attributed to S. bifurcata based on the types. Correspondingly, the description of ‘ S. bifurcata ’ (including type locality but excluding the figure) is herein attributed to S. zebra also based on the types. The concept of S. bifurcata in Hubendick (1946: 46) is confused. He appears not to have examined the types and must have been misled by the above-mentioned mix-up of descriptions in Reeve (1856). Hubendick (1946) mentioned ‘ specimens examined’ from Port Jackson [Sydney], which is outside the known distribution of S. bifurcata. The shells figured by Hubendick (1946: pl. 2, figs 9 – 13) and the description of Hubendick (1946: 14, fig. 13) are herein attributed to S. denticulata and S. zelandica, respectively. Our delineation of S. bifurcata is based on comparative analyses of the morpho-anatomy and mitochondrial genetics of freshly collected topotypes (Figs 35 E, F) and geographic series of additional specimens (Table S 1).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFC18241FF68F822FB7DFAB6.taxon	description	External morphology (Fig. 35 O). External parts evenly pale cream without black pigmentation apart from faint shading at centre of cephalic folds; mantle lobed, translucent; single small black epithelial eye spots centralised on two centrally touching cephalic folds; pneumostomal lobe under the mantle, unpigmented, between the right anterior and right posterior ADMs. Shell (Figs 35 A – F, S; Table S 9). Medium sized (max sl mean = 15.8 mm, SD = 4.4 mm, n = 2), elongate ovate; height low to medium; apex offset posterior and left, apical sides convex, protoconch direction weakly heterostrophic (n = 1; Fig. 35 S) shell whorl dextral; growth striae indistinct, irregular growth may create rib irregularities and offsets; shell thick, externally white; rib count (mean = 43, SD = 5, n = 2), ~ 11 primary ribs white, fairly straight but may be bent, raised, rounded ridge, most prominent with paired ribs forming siphonal ridge and prominent protrusion at right quarter; primary ribs broaden to and protrude beyond shell lip to strongly unevenly scallop and corrugate the edge; finer secondary ribs fill gaps between primary ribs, rib interstices narrow. Interior outer shell margin white with pale brown markings aligning under rib interstices, interior pink in one specimen (Fig. 35 E); siphonal groove distinct, appears notched / raised, same colour as shell edge, points to right anterior; spatula mottled brown underlying white; ADM scar distinct, CMS straight, golden / brown; thickening of shell lip apparent, infills and reduces lip scalloping, spatula becomes whitened. Reproductive system (Fig. 36 A, C; n = 3). Positioned within coelom under the respiratory cavity, hermaphroditic glands positioned to posterior against right foot wall and over foot sole, epiphallic parts positioned over and looped / folded in front of BM and to side of RAM; AO large, elongated, bluntly pointed (embeds into MG), centrally bent, often with MA, joins to top of indistinct GA; ED relatively short, broad, slightly twisted; EG small, folded; at join of ED and EG a single long, narrow, twisted flagellum F 1 joins as an extension of ED; AO, GA and ED all muscular white tissue; BD and CD junctions into GA (bulbous at CD) opposing between ED, AO and GP; BD longer than CD with a prominent distal loop, top of loop attached via a long MA to inner foot wall in front of BM, both ducts smooth, similar thickness, pass together through RAM (BD above CD), CD connects to midsized bulbous translucent test BC, CD connecting into folds of MG; HD brownish small coiled, links AG to elongated narrow brownish coarsely granulated HG; MG and AG similar sized, folded soft white tissue, dark SV embedded within AG / MG, AG and HG sides match curvature of inner foot wall at right posterior quarter of coelom. Spermatophore (Fig. 36 B). Thread-like, test thin, translucent (length = 6.78 mm, n = 1, AL = 15 mm); head cylindrical, broad, tip bluntly rounded, tapers into long flagellum; flagellum attached to inside of BC; both sections smooth, featureless; head longer and thicker than flagellum (head length = 4.79 mm, ~ 71 % of total length, head width = 103 μm, flagellum width = 17 μm, n = 1); SPM tightly coiled in light brown gelatinous mass in one BC (WAM S 74097).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFC18241FF68F822FB7DFAB6.taxon	diagnosis	Comparative remarks. Siphonaria bifurcata (atra group, unit 45) forms a well-differentiated subclade within the mitochondrial tree and is the sister clade of a subclade containing S. umbra sp. nov. and S. radians (Figs 1, 2). Siphonaria bifurcata is well-differentiated from any other species by COI distances of ≥ 29 % (Table S 4). This species has been found in Cebu, Philippines in sympatry with four congeners. For comparison with S. sipho refer to comparative remarks under that species. Siphonaria alba has a lower shell with a more scalloped edge, a narrower AO, larger BC, smaller GA, and a shorter F 1. Siphonaria sirius (unit 31), has a more central shell apex and scalloped edge, darker external and internal colour, and a paler spatula. Siphonaria caubianensis sp. nov. has a slightly taller shell with wider primary ribs, a more strongly scalloped edge, darker external and internal colour, paler spatula, a narrower AO, strongly twisted ED, and bursal loop. The RS of S. bifurcata resembles that of several other species, such as S. denticulata, S. javanica, S. poindimiensis sp. nov., S. viridis, and S. tanguissonensis sp. nov. According to the phylogenetic tree (Fig. 1), none of these species is closely related to S. bifurcata. The shell of S. bifurcata resembles that of other species in the laciniosa and atra groups, such as S. sipho, S. alba, and S. atra in that it exhibits variations of a thickened white shell lip. Several authors have been misled by the mix-up of text labels and figures in Reeve’s (1856) description and confused this species with S. zebra. Consequently, S. bifurcata was repeatedly listed as a junior synonym of S. zelandica, which it is not. Figures of ‘ S. bifurcata ’ from ‘ Port Jackson’ [Sydney] in Hubendick (1946: pl. 2, figs 9 – 13) are herein attributed to S. zelandica. The RS figure of ‘ Ductosiphonaria bifurcata ’ in McAlpine (1952: 42) is also of S. zelandica. The SPM of ‘ S. bifurcata ’ from ‘ Port Jackson’ depicted in Hubendick (1946: fig. 19) and reproduced in Berry (1977: fig. 19) closely matches that of S. denticulate but not of S. zelandica.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFC18241FF68F822FB7DFAB6.taxon	distribution	Distribution and habitat. Philippines, recorded from Mactan (Cebu), Quezon, and Siquijor islands (Fig. 37). In the present study found in sheltered places on moderately exposed rocky shores, lower littoral level.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFC08243FCCAFA02FA1EFB35.taxon	description	(Figs 35 G – I, Q, 36 D – E)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFC08243FCCAFA02FA1EFB35.taxon	materials_examined	Other, non-type material. Rodrigues: Rivière Banane, N coast, 19 ° 11.25 ’ S, 63 ° 22.866 ’ E RG 01 - 1 (AM C. 585896 6 p, C. 585190 p [M 428, SK 331], C. 585191 p [M 429, SK 332], C. 585194 p [SK 369]; Anse Quiter, SW coast, 19 ° 46.183 ’ S, 63 ° 22.866 ’ E, RG 02 - 1 (AM C. 585874 p [M 430, SK 134]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFC08243FCCAFA02FA1EFB35.taxon	discussion	Taxonomic remarks. No type locality has been given in the original description of Reeve (1856). For its unknown origins, this taxon has subsequently been considered as a nomen dubium by White & Dayrat (2012). However, examining freshly collected from Rodrigues revealed a shell form that closely resembled the features of the holotype. We therefore infer that these samples and the type of S. fuliginata are conspecific. Here, we provide a detailed redescription of S. fuliginata based on samples from Rodrigues to remove the uncertainty about the identity of this species.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFC08243FCCAFA02FA1EFB35.taxon	description	External morphology. Foot wall, mantle, cephalic folds and pneumostomal lobe all evenly pale cream in colour; dark pigmentation markings absent, foot sole darker cream, foot edge paler; Mantle translucent, narrower than foot wall, lobed with a thickened edge, pneumostome wide between right ADMs and within mantle. Shell (Figs 35 G – I; Table S 9). Small sized (max sl mean = 9.7 mm, SD = 2.9 mm, n = 3), ovate, height medium to low, apex weakly offset central and left, apical sides convex, protoconch direction undetermined, shell whorl dextral growth lines distinct; rib count (mean = 27, SD = 1.7, n = 3), raised, whitish, straight, rib width increases to shell lip; rib interstices paler, may have irregular red / brown markings; ~ 9 primary ribs, extend from apex to shell edge, similar secondary ribs between primary ribs; paired primary ribs over siphonal ridge; shell lip weakly scalloped, corrugated aligning with rib ends; interior colouration of margin rays match underside of white primary ribs and brown rib interstices, inner margin often brown from inner margin to the golden (often mottled blue) coloured spatula (Fig. 35 H); ADM impression and siphonal groove distinct, paler than shell margin colouring, cephalic scar convex; thickening and whitening of shell lip / margin not observed; some shells may be display dark brown exterior and interior (AM C. 585191). Reproductive system (Fig. 36 D; n = 3). Positioned within coelom under the respiratory cavity, epiphallic parts positioned between RAM and over back of BM close. ED joins at underside of small GA, AO short, bluntly pointed, smaller than GA, joins through underside of ED; ED thick, elongated, centrally twisted, broader at EG; single broad curled stubby flagellum F 1, appears as extension of ED at connection with EG; AO, GA and ED all muscular white tissue; EG broad, relatedly large, soft white folded tissue; BD and CD connect in parallel to GA at top and side of GA respectively; BD without distal loop or MA, slightly longer and thinner than CD, both ducts smooth and pass together between RAM and inner foot wall connecting into thick layered folds of MG (BD over CD), bursal loop in BD immediately in front of BC; BC relatively large, spherical, embedded along with part of BD in AG / MG; SV embedded in AG under BC; HD short, narrow, straight, links large AG to a much smaller yellowish granulated HG, AG and MG folded, inner edge MG lobed, both soft white tissue, with outer sides curved reflecting the close positioning to curvature of inner foot wall at right posterior quarter of coelom, MG flattened against inner foot sole. Spermatophore (Fig. 36 E). Bulbous, test thin, translucent (length = 1.43 mm, n = 1, AL = 6.5 mm); head cylindrical, broad, tip bluntly rounded, containing a white gelatinous core, tapers into short flagellum; both sections smooth, featureless; head longer and thicker than flagellum (head length = 1.03 mm, ~ 89 % of total length, head width = 138 μm, flagellum width = 35 μm, n = 1); 2 SPM tightly coiled in cream gelatinous mass in one BC (Fig. 36 E).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFC08243FCCAFA02FA1EFB35.taxon	diagnosis	Comparative remarks. Siphonaria fuliginata (normalis group, unit 80) is closely related to S. normalis (unit 14) and S. madangensis (unit 88) (Figs 1, 4). It differs from these two species by COI distances of ≥ 5.9 % (S. normalis) and ≥ 4.4 % (S. madangensis), respectively (Table S 8). From the next more closely related species, S. costellata, it differs by COI distances of ≥ 6.5 %. The close phylogenetic relationships with S. normalis, S. madangensis, and S. costellata are reflected in shared similarities in shell characters (thickened white shell lip variations) and the RS (broad epiphallic parts). However, S. normalis has a darker, more finely ribbed shell, a longer, narrower BD, larger BC, and a longer SPM. Siphonaria madangensis sp. nov. has a darker shell with a more central apex and prominent siphonal ridge, a smaller to indistinct AO, BD without a bursal loop, and a more bulbous SPM. Siphonaria costellata sp. nov. has a larger, taller, more brownish (less grey) shell, with a more central apex, a smaller AO, and wider BD without a bursal loop. Siphonaria fuliginata has been found in sympatry with Siphonaria rodriguensis sp. nov. on Rodrigues Is, which has a smaller, taller, darker shell with pale parallel-marked primary ribs, a longer wider BD, larger BC and AG, and longer, narrower SPM.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFC08243FCCAFA02FA1EFB35.taxon	distribution	Distribution and habitat. Known only from Rodrigues, Indian Ocean (Fig. 37). In the present study found in sheltered positions on moderately exposed rocky shores, mid littoral level, amongst barnacles (Fig. 35 Q).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFC2827DFCCAFA82FBBCFC35.taxon	description	(Figs 35 J – N, R, T – V, 36 F – H)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFC2827DFCCAFA82FBBCFC35.taxon	materials_examined	Material examined. Type material. Lectotype of Siphonaria lirata Reeve, 1856, present designation, without locality (NHMUK MC. 1979028 / 1, Fig. 35 J). Three paralectotypes, same data as lectotype (NHMUK MC. 1979028 / 2 - 4, Figs 35 K – M). Other, non-type material. Guam: Pago Bay, below UoG Marine Lab, 13 ° 25.645 ’ N, 144 ° 47.927 ’ E, GM 04 - 1 (AM C. 585890 3 p, C. 584885 p [SK 251]); Coast W of Piti Bay, Apra, 13 ° 27.879 ’ N, 144 ° 40.762 ’ E, GM 03 - 1 (AM C. 585889 10 + p, C. 585192 p [SK 252], C. 585832 p [M 448, SK 242], C. 585833 p [M 449, SK 162]). Taxonomic remarks. The largest syntype (Fig. 35 J) is herein designated as the lectotype of S. lirata for the stabilisation of the name (Art. 74.1 of the Code). Reeve (1856: Erratum) clarified a mistake in labelling pl. 7, fig. 35: The correct name is S. lirata and not S. fuliginata. No type locality has been given in the original description of Reeve (1856). However, examining freshly collected from Guam revealed a shell form that closely resembled the features of the syntypes. Our delineation of this species is based on comparative analyses of the morpho-anatomy and mitochondrial genetics of freshly collected specimens from Guam (Fig. 35 N) and geographic series of additional specimens (Table S 1). Pilsbry (1920 b: 379) and Cernohorsky (1972: 210) incorrectly considered S. lirata as a synonym of S. normalis. Both Hubendick (1946: 23) and Jenkins (1981: 2) incorrectly treated S. lirata as a synonym of S. funiculata. External morphology. Exterior evenly grey, paler grey to cream at foot edge and inner foot wall, foot sole darker grey; irregular black blotches of pigmentation on foot wall, pneumostomal lobe and concentrated over front of cephalic folds; mantle translucent, covers shell mantle, wider at anterior, outer edge thickened, weakly lobed with black bands of pigmentation aligning with rib interstices; pneumostomal lobe small, under mantle between the right anterior and right posterior ADMs, closes the pneumostomal and anal openings at the mantle edge; two small black epithelial eye spots centralised on centrally touching cephalic folds; genital pore inconspicuous, located on foot wall to right anterior of right cephalic lobe. Shell (Figs 35 J – N, V; Table S 9). Small sized (max sl mean = 9.6 mm, SD = 0.6 mm, n = 3), ovate, height medium; apex offset posterior and to left; apical sides convex, posterior concave to straight; protoconch direction heterostrophic (n = 2; Fig. 35 V), area black colouration, shell whorl dextral; growth striae weak; shell thickness thin; colouration uneven with some radial banding; rib count (mean = 66.7, SD = 7.3, n = 3); slightly raised, pale white, fairly straight, faintly protrude beyond shell lip; predominantly primary ribs, finer secondary ribs interspersed, develop between primary ribs with shell growth, rib interstices darker; siphonal ridge formed by paired primary ribs, protrudes past shell edge, otherwise indistinct. Interior evenly dark brown to black from margin to spatula, paler on shell lip aligning under rib ends, siphonal groove clear; ADM scar indistinct, CMS weakly convex; No evidence of growth variations in shell thickness or shell margin colouration was observed. Reproductive system (Figs 36 F, G; n = 2). Positioned within coelom under the respiratory cavity, hermaphroditic glands positioned to posterior against right foot wall and over foot sole, epiphallic parts positioned over and looped / folded in front of BM and to side of RAM; AO large, elongated, bluntly pointed (embeds into MG), centrally bent with MA, merges with indistinct GA; ED relatively short, slightly twisted, thick with side appendage; EG relatively large, folded, elongated and pointed; single short narrow bent flagellum F 1 on EG; AO, GA and ED all muscular white tissue; BD and CD opposing connections (bulbous at CD) into GA between ED, AO and GP; BD longer and narrower than CD with a prominent distal loop, top of loop attached via a short MA to inner foot wall in front of BM, both ducts smooth and pass together through RAM connecting into folds of MG (BD above CD), BC translucent test, midsized and bulbous; HD brownish long coiled links AG to a small elongated narrow brownish finely granulated HG; MG and AG small folded soft white tissue; dark SV embedded within AG, AG larger than HG, sides match curvature of inner foot wall at right posterior quarter of coelom. Spermatophore (Fig. 36 H). Thread-like, test thin, translucent; head cylindrical, very narrow, tip bluntly rounded, containing a white gelatinous core; taper region into the filamentous transparent flagellum is extended; both sections smooth, featureless; head longer and much thicker than flagellum (head length = 5.09 mm, n = 1, head width = 66.6 μm, flagellum width = 13 μm, flagellum incomplete); 1 SPM tightly coiled in a white gelatinous mass in one BC (Fig. 36 H). Comparative remarks. Siphonaria lirata (plicata group, unit 58) is the sister species of S. tanguissonensis sp. nov., also from Guam (Figs 1, 3). Both species differ from each other by COI distances of ≥ 11.6 % (Table S 7). Siphonaria lirata differs from S. monticulus by COI distances of ≥ 15 % and from other species by ≥ 21 % (Table S 7). Siphonaria lirata has been found in sympatry with three congeners in Guam. For comparison with S. guamensis and S. normalis refer to comparative remarks under these species. Siphonaria tanguissonensis sp. nov. has a lower, darker shell with a paler, golden brown / tan interior, a smaller AO, and larger ED. Species with similar shell forms are S. mauiensis sp. nov., S. normalis), S. exulum, and S. incerta. Previous records of ‘ S. lirata ’ from southern Australia are based on misidentifications of S. funiculata, which has a somewhat similar shell.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFC2827DFCCAFA82FBBCFC35.taxon	distribution	Distribution and habitat. Endemic to Guam (Fig. 37). In the present study found on rock platforms and boulder areas on exposed rocky shores, mid to upper littoral levels (Fig. 35 T).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFFC827FFCCAF922FEA1FA16.taxon	description	(Figs 38 A – E, 39 A – B)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFFC827FFCCAF922FEA1FA16.taxon	materials_examined	Material examined. Type material. Holotype of Siphonaria carbo Hanley, 1858; coll. Cuming (NHMUK 1981009, Fig. 38 A). Lectotype of Siphonaria nigerrima Smith, 1903, present designation, from Umhlali, KwaZulu-Natal, South Africa; donated J H. Ponsonby (NHMUK 1903.9.9.15, Fig. 38 B). Two paralectotypes, same data as lectotype (NHMUK 1903.9.9.16 – 17). Eight syntypes of Siphonaria tenuicostulata Smith, 1903 from Umhlali, KwaZulu-Natal, South Africa; donated J H. Ponsonby (NHMUK 1903.9.9.7 (Fig. 38 C, largest syntype) – 14). Other, non-type material. Mozambique: Inhaca, Ponta do Farol, 25 ° 58.2 ’ S, 32 ° 59.4 ’ E MM 6 (MNHN IM- 2019 - 1489 29 p, MNHN IM- 2019 - 16164 p [M 586], MNHN IM- 2019 - 16165 p [M 587]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFFC827FFCCAF922FEA1FA16.taxon	discussion	Taxonomic remarks. Hubendick (1946: 35) treated S. nigerrima as a synonym of S. carbo Hanley, 1858. Which was confirmed later in the taxonomic revision of Teske et al. (2007) based on comparative morphology and mitochondrial phylogenetics. They added the species S. anneae and S. tenuicostulata to the synonymy of this species, which is also accepted herein. The largest syntype of S. nigerrima is herein designated as the lectotype (Fig. 38 B) for the stabilisation of the name (Art. 74.1 of the Code). This type specimens matches the typical characteristics of S. carbo (Fig. 38 A) in size, shape, height, colour (Fig. 38 B). Our analysis of topotypical specimens of S. nigerrima confirms its status as a synonym of S. carbo. Previously, Chambers & McQuaid (1994 a: 264) incorrectly stated that S. carbo was described from the Caribbean and ‘ does not occur on South African shores’. This observation is not accepted herein. Based on examinations of topotypic samples matching the morphological characteristics of S. dayi (Fig. 38 D, E), this taxon is also placed in the synonymy of S. carbo.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFFC827FFCCAF922FEA1FA16.taxon	description	External morphology (preserved). Foot sole grey, paler to foot edge; mantle translucent, pigmentation absent on foot wall, mantle edge, pneumostome; faint pigmentation over centre of cephalic folds. Shell (Figs 38 A – E; Table S 9). Small to medium sized (max sl mean = 12.5 mm, SD = 6.0 mm, n = 5), circular ovate, height tall; apex offset central, apical sides convex, protoconch direction weakly homostrophic to central (n = 1), shell whorl dextral; exterior dark metallic brown / grey, often pale brown; primary ribs with 0 – 2 finer in between secondary ribs, interstices similar colour, growth striae prominent; radial colour bands distinct, darker at apex and shell margin; rib count (mean = 48, SD = 7.8, n = 5), majority primary ribs, ridges weakly raised, rounded; siphonal ridge indistinct, formed by paired primary ribs, primary ribs broaden weakly, project weakly beyond shell lip (especially siphonal ridge) to scallop and corrugate shell edge. Interior of shell evenly dark brown to grey (Fig. 38 A – B), may also display off-white rays on shell margin align under primary / secondary ribs, extending to spatula (Fig 38 C); siphonal groove distinct, shallow, often paler than margin; ADM scar clear, similar to margin and spatula; CMS convex; whitening and thickening of shell lip not observed. Reproductive system (Fig. 39 A; n = 1). Positioned within coelom under the respiratory cavity, hermaphroditic glands positioned to posterior against right foot wall and over foot sole, epiphallic parts between RAM and extending over BM; GA large, AO indistinct and appears part of GA, singular prominent GP, ED short, centrally bent; ED enters at upper end of GA, AO, GA and ED all muscular white tissue; EG very large, elongated, folded, soft white tissue; single thick elongated blunt flagellum F 1 appears as extension of ED at EG join; BD and CD connect closely together into GA close to ED junction, both ducts smooth, featureless, wide, slightly bent, pass together between inner foot wall and outer side of RAM (BD over CD) connecting into MG / AG; BC large, bulbous, white opaque test, embedded along with part of BD in soft white folds of MG; HD short, wide, unpigmented, heavily lobed, links soft white folded AG to smaller yellowish granular HG; SV embedded in AG close to BC. Spermatophore (Fig. 39 B). Broad head with short flagellum (length = 1.29 mm, n = 1); head section bulbous but flattened, test thin, smooth, featureless; tapering section merges head to flagellum; head longer, wider than flagellum (head length = 0.77 mm, n = 1; flagellum length; 0.53 mm, ~ 59 % of SMP length; head width = 362 μm, flagellum width = 149 μm); 2 SPMs held in cream gelatinous mass in BC of one specimen [M 587]. Radula. Dentition formula 36: 1: 36 (Hubendick 1947: 162 as ‘ S. nigerrima’), 36: 1: 36 (Hubendick 1946: 35 as ‘ S. carbo ’) and 40: 1: 40 (Hubendick 1946: 35, 1947: 162 as ‘ S. tenuicostulata ’).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFFC827FFCCAF922FEA1FA16.taxon	diagnosis	Comparative remarks. Collectively, the works of Allanson (1958), Chambers & McQuaid, (1994 a: 264) and Teske et al. (2007, 2011: 5025) clarified the taxonomy of South African Siphonaria species, which is followed herein except that S. dayi is treated as a junior synonym of S. carbo. Our study does not include analyses of the South African species S. concinna, S. oculus and S. serrata; refer to Teske et al. (2007, 2011) for comparisons. We found that S. carbo (pectinata group, unit 94) is the sister species of S. itampoloensis sp. nov. in our phylogenetic tree (Figs 1, 4). Both species together form the sister lineage of S. asghar. Siphonaria carbo differs from S. itampoloensis by COI distances of ≥ 5.1 % and from S. asghar by distances of ≥ 11.1 % (Table S 8). Siphonaria carbo differs from S. itampoloensis by having a darker, lower shell with finer ribbing, a wider BD, larger EG, and a larger, elongate BC. Siphonaria carbo has been found in sympatry with two congeners in Mozambique. For comparisons with S. capensis and S. plana refer to comparative remarks under these species. The shape of the SPM of S. carbo closely resembles that of other species, such as S. funiculata, S. zelandica, S. acmaeoides, S. lateralis, and S. tasmanica. However, Siphonaria carbo has a smaller ED / EG and broader CD / BD’s than these species. The RS of ‘ S. tenuicostulata’ (= S. carbo) figured in Hubendick (1945: 21, fig. 22) corresponds well with the typical anatomy of this species except for details of the CD / BD / GP / GA junction, which Hubendick (1946: fig. 20) showed to be wider and with the ducts more separated. Hubendick (1947: 161) considered the RS of ‘ S. tenuicostulata ’ (fig. 1) and ‘ S. nigerrima ’ (fig. 2, epiphallic parts absent), from Umhlali Natal, to be a ‘ closely related’ species ‘ only showing differences in small details’. Hubendick (1947) figured a RS matching that of S. carbo shown here (Fig. 39 A) more closely than any other species. Allanson (1958: 167) stated that the distal genitalia of S. anneae were ‘ Exactly similar to S. teniucostulata ’ referring to ‘ S. tenuicostulata ’ from Oman figured in Bosch (1982: 141; 1991: 75). However, these figures are incorrectly assigned to S. tenuicostulata (= S. carbo) because this location is well outside the known distribution of this species.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFFC827FFCCAF922FEA1FA16.taxon	distribution	Distribution and habitat. Recorded from tropical and subtropical coasts of southeastern Africa (Fig. 25). Found on exposed rocky shores across mid to upper littoral levels.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFFE827AFF68F9A2FA29FC96.taxon	description	(Figs 38 F – T, 39 E – H)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFFE827AFF68F9A2FA29FC96.taxon	materials_examined	Material examined. Type material. Holotype of Siphonaria exulum Hanley, 1858 a from Norfolk Is [Australia] (NHMUK 1900.3.19.27, Fig. 38 F). Holotype of Siphonaria raoulensis Oliver, 1915 from Rocks between tide marks, Sunday Island, Kermadec Islands, [NZ] (CM M. 3666, Fig. 38 I). Four paratypes, Raoul Is; coll. W. R. B. Oliver (AM C. 40293). Holotype of Siphonaria cheesemani Oliver, 1915 from Sunday Island, Kermadec Islands, [NZ]; coll. W. R. B. Oliver (CM M. 3660, Fig. 38 J). Four paratypes, Raoul Is; coll. W. R. B. Oliver (AM C. 40294). Holotype of Siphonaria macauleyensis Oliver, 1915 from Macauley Island, Kermadec Islands, [NZ]; coll. W. R. B. Oliver 1908 (CM M. 3663, Fig. 38 K). Holotype of Siphonaria macauleyensis perplexa Oliver, 1915 from Sunday Island, Kermadec Islands, [NZ]; coll. W. R. B. Oliver, 1908 (CM M. 3661, Fig. 38 L). Two paratypes, Raoul Is; coll. 1908, W. R. B. Oliver (AM C. 40300). Holotype of Siphonaria amphibia Oliver, 1915 from Fleetwood Bluff, Sunday Island, Kermadec Islands, [NZ]; coll. W. R. B. Oliver 1908 (CM M. 3665, Fig. 38 M). Two probable paratypes, Raoul Is; coll. W. R. B. Oliver (AM C. 40299). Syntype of Parellsiphon innocuus Iredale, 1940 from Norfolk Is; coll. 1910 (AM C. 103708, Fig. 38 N). Other, non-type material. NZ, Kermadec Islands: Raoul Island, Fishing Rock landing, 29 ° 14.55 ’ S, 177 ° 54.22 ’ W (AM C. 608196 p [SK 418 protoconch I 5], C. 595914 p [SK 419 protoconch G 12]); Boat Cove, 29 ° 16.783 ’ S, 177 ° 53.65 ’ W K 2011 - 102 (AM C. 475847 p [M 513], 29 ° 16.666 ’ S, 177 ° 53.717 ’ W K 2011 - 29 - 1 (AM C. 475481 d); south side of Te Konui Pt, 29 ° 18.53 ’ S, 177 ° 53.75 ’ W. (AM C. 475477 d, C. 475479 d, C. 475480 d). South Meyer Island, 29 ° 14.817 ’ S, 177 ° 52.817 ’ W (AM C. 475848 p [M 512]). Australia, NI: Anson Bay, NW side of NI, Stn D 3 / 2 rock pools, 29 ° 00 ’ S, 167 ° 55 ’ E (AM C. 595968 p); Anson Bay, 29 ° 00.550 ’ S, 167 ° 55.332 ’ E NFI 04 - 1 (AM C. 585378 10 p, C. 585024 p [M 219], C. 585025 p [SK 041]); Duncombe Bay, N side of NI, 29 ° 00 ’ 02.5 ’ S, 67 ° 55 ’ 48.8 ’ E (AM C. 595965 10 + p, C. 595946 p [SK 005]); Cascade Bay, 29 ° 01.206 ’ S, 167 ° 58.174 ’ E NFI 05 - 1 (AM C. 585551 26 p, C. 585026 p [M 220], C. 585027 d [R 001], C. 585028 d [R 002], C. 585029 p [SK 010]); Ball Bay, 29 ° 02 ’ S, 167 ° 59 ’ E (AM C. 595964 10 + p); Ball Bay 29 ° 02.844 ’ S, 167 ° 59.044 ’ E NFI 01 - 1 (AM C. 585526 20 + p); Creswell Bay 29 ° 03.478 ’ S, 167 ° 56.547 ’ E NFI 03 - 1 (AM C. 585400 10 + p, C. 585023 p [M 221]); Slaughter Bay 29 ° 03.511 ’ S, 167 ° 57.416 ’ E NFI 02 - 1 (AM C. 585581 30 p, C. 585020 p [M 217], C. 585021 p [M 218], C. 585022 p [SK 009]); Emily Bay, S side NI 29 ° 03 ’ 36.0 ’ S 167 ° 57 ’ 40.5 ’ E (AM C. 595967 p); Point Hunter Reserve, E side of cemetery 29 ° 03 ’ S, 167 ° 58 ’ E (AM C. 595966 10 + p, C. 595944 p [SK 042]); Point Hunter 29 ° 03.629 ’ S, 167 ° 58.045 ’ E NFI 02 - 2 (AM C. 585681 7 p, C. 595947 p [SK 006]); Phillip Island 29 ° 06.9 ’ S, 167 ° 56.88 ’ E (AM C. 585944 15 p). LHI: Signal Point 31 ° 31.501 ’ S, 159 ° 03.578 ’ E LHI 2017 Apr 04 - 099 (AM C. 546717 5 p, C. 595945 p [SK 052]); On marine debris Old Settlement, LHI, 31 ° 31.18 ’ S, 159 ° 03.45 ’ E (AM C. 582859 8 p, C. 585945 p [M 467, SK 234]). Taxonomic remarks. The original description of S. exulum gives measurements for a single specimen, the holotype. The holotype is the only type specimen known to exist (J. Ablett, pers. comm. NHM). No type has originally been designated in the description of P. innocuus. The type specimen in Iredale (1940: 441, pl. 34, figs 9 – 10) is therefore considered as a syntype (Fig. 38 N). No other type specimens are known. Our delineation of this species is based on comparative analyses of the morpho-anatomy and mitochondrial genetics of freshly collected topotypes of S. exulum (Figs 38 G), P. innocuus (Fig. 38 Q), S. raoulensis (Fig. 38 O), S. cheesemani (Fig. 38 P), S. macauleyensis and S. macauleyensis perplexa (Fig. 38 O) and geographic series of additional specimens (Table S 1). These analyses confirm the synonymy of P. innocuous, S. raoulensis, S. amphibia, S. cheesemani, S. macauleyensis and S. macauleyensis perplexa. Siphonaria amphibia, S. cheesemani, S. macauleyensis and S. macauleyensis perplexa had previously been synonymised with S. raoulensis by Wood & Gardiner (2007) based on molecular phylogenetic evidence. However, the relevant sequences are not available on Genbank (accessed 28 April 2021) precluding their use in this review. Records of ‘ S. corrugata ’ and ‘ S. lirata ’ from Norfolk Island in Brazier (1888, 1001) are misidentifications attributed herein to S. exulum. Paetel (1889: 428) listed both S. exulum and its emendation S. exulorum as accepted species. Suter (1907: 265) misidentified specimens of this species from the Kermadec Islands as S. diemenensis. Oliver (1915: 545) described several species from the Kermadec, which he considered to be distinct although ‘ difficult to separate’. All these species are synonyms of S. exulum. Iredale (1940, 438) apparently not being aware of Hanley’s description, incorrectly attributed the name exulorum to Suter and tentatively placed it in Ellsiphon. Hubendick (1946: 36, 38, 63) incorrectly treated S. cheesemani as a synonym of ‘ S. cookiana ’ (= S. propria); S. raoulensis, S. macauleyensis and S. perplexa as synonyms or varieties of S. diemenensis; and S. amphibia as a synonym of S. acmaeoides. However, he listed no specimens from the Kermadecs under ‘ specimens examined’. Furthermore, Hubendick (1946: 38) also listed S. exulum and S. exulorum (‘ Error for exulum ’) as varieties of S. diemenensis Quoy & Gaimard, 1833, which is not accepted herein. Morrison (1972: 56 – 58) treated ‘ Parellsiphon innocuous ’ (sic innocuus) as a synonym of S. laciniosa based on similarity in shell form and ‘ common reproductive development’. This synonymy is not supported by examination of type specimens and morpho-anatomy. Trew (1983: 5) treated S. exulorum (= exulum) as a synonym of ‘ Pachysiphonaria diemenensis ’. Grove (2006: 60) also synonymized S. exulum with S. diemenensis, but the specimen figured by Grove (2017) as ‘ S. exulum is actually an individual of S. diemenensis. Brook (1998: 232) stated that ‘ there is only one morphologically variable species of Siphonaria, namely S. raoulensis, at the northern Kermadec Islands’. The correct name for this species is S. exulum, however. External morphology (Fig. 38 S). Foot sole smooth, evenly dark yellow to centrally greyish; foot wall narrow dark yellow with evenly spread white subepithelial pustules becoming more vivid and dense close to the foot sole and around pneumostomal lobe; fringing mantle narrow, yellowish translucent, extends to shell edge, outer edge lobed, strongly banded yellow, reflects the profile shell lip and ribs; pneumostomal lobe paler and within mantle between the right anterior and right posterior ADMs, closes the pneumostomal and anal openings at the mantle edge; two small black epithelial eye spots centralised on two thick centrally touching dark yellow cephalic folds that darken to their outer edge, covered with white mucous cells similar (but smaller) to those of the foot wall tissue; genital pore inconspicuous, located on foot wall to right anterior of right cephalic fold. Shell (Figs 38 K – M, R – T; Table S 9). Small to medium sized (max sl mean = 14.4 mm, SD = 2.2 mm, n = 24); ovate, thin, height tall to flattened (‘ exulum ’, ‘ raoulensis’ and ‘ macauleyensis ’ forms, Figs 38 H, K – O) to flat (‘ cheesemani ’ form, Fig. 38 P); dark and pale shell forms variable, exterior uneven; apex fairly central and weakly offset to posterior; apical sides evenly convex; apex weakly hooked, protoconch direction homostrophic (n = 4; Fig. 38 R), shell whorl dextral; rib count (mean 39, SD = 7, n = 24) slightly raised to strongly raised, primary ribs grey to white, fairly straight, very prominent in some individuals; few secondary ribs, develop between primary ribs to a similar size at shell margin, rib interstices dark brown to black; paired touching primary ribs over indistinct siphonal ridge; growth striae distinct, 2 – 3 discontinuous bands of brown radial shading, protoconch area dark brown; shell edge uneven, scalloped by weakly extending ribs; interior mottled brown to evenly dark brown, apart from short cream markings on shell lip aligning with primary ribs; spatula pale brown to blue / white, pale shell form has golden brown spatula. Siphonal groove shallow, ADM scar variably distinct, CMS convex. Translucent thickening of internal shell occurs, infills and reduces scalloping of lip, spatula becomes whitened (prominent in pale shell forms; ‘ innocuus ’; Fig. 38 Q, P). Reproductive system (Figs 39 E, G, H; n = 3). Single GP located in foot wall on right posterior side of right cephalic fold; opens internally into a relatively small GA positioned between right dorsal of BM and RAM; elongated horn-shaped AO present protrudes as an extension of GA, a short wide whitish muscular tissued ED joins side of GA; a singular short thin blunt hooked cream-coloured flagellum F 1 protrudes from the join between the ED and lobed blunt whitish EG; the long, narrow thickened whitish CD and thin smooth unlooped BD (BD thinner and dorsal to CD) together pass through the RAM to enter the GA close to GP (BD closest), posterior to entry of AO and ED; at the end of BD is a relatively small spherical brownish BC, test thin, loosely embedded in dorsal of AG, SV elongated, deeply embedded; CD (spermoviduct) joins soft white folds of MG and AG; HD thickened distinctly coiled and striated, connects AG to the yellowish granulated crescent shaped HG. The HG is positioned in right posterior of coelom under pallial cavity, to which the AG is distal and positioned under the digestive gland. Spermatophore (Fig. 39 F). Thread-like (length = 8.66 mm, n = 1), translucent, test thin; head section, bluntly rounded, body cylindrical, containing a white gelatinous mass, tapers along the transparent flagellum to a very thin tip; both sections smooth, featureless; head shorter, thinner than flagellum (head length = 3.37 mm; flagellum length = 5.32 mm; ~ 39 % of SPM length; head width = 71 μm; flagellum width = 21 μm). Single SPM coiled embedded in brown gelatinous mass in one BC (LHI, AM C. 546717 [SK 052]). Comparative remarks. Siphonaria exulum (atra group, unit 47) is the sister species of a subclade containing four species, S. scabra, S. pravitas sp. nov., S. bourailensis sp. nov., and S. ouassensis sp. nov. All five species together form Clade B in the siphonariid tree (Figs 1, 2). Siphonaria exulum differs from other species by COI distances of ≥ 18.9 % (Table S 4). Siphonaria exulum has been found in sympatry with S. lentula on LHI, which has a shell with more prominent raised ribs, scalloped edge, and siphonal ridge with a flared end, darker interior, BD with distal loop, and SPM with a bulbous head. Siphonaria pravitas sp. nov. has more prominent and raised ribs on siphonal ridge, greater edge scalloping, darker interior, a shorter wider BD without a distal loop, a larger BC and a shorter SPM. Siphonaria exulum is the only known species in the Kermadec Islands. Siphonaria ouasseensis has a lower, paler, less evenly ribbed shell, a smaller, narrower AO, a more elongate and narrower CD, and a smaller BC, and S. bourailensis has a lower, less evenly and broader ribbed shell, more prominent siphonal ridge, a larger, broader AO, a more elongate and narrower CD, a larger HD, and a shorter SPM. We found that the RS structures of specimens from NI correspond reasonably well with those of specimens from the Kermadec Islands especially in size and shape of the epiphallic parts, AO and HD (hermaphroditic parts may vary) (Figs 39 E, G, H). Conchologically, the most similar species are S. mauiensis sp. nov. (Hawaii), S. griffithsorum sp. nov. (Mauritius), and S. tongatapuensis sp. nov. (Tonga).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFFE827AFF68F9A2FA29FC96.taxon	distribution	Distribution and habitat. Known exclusively from Kermadec Islands, NI and LHI (Fig. 37). In this study, found to be common in sheltered rocky surfaces (e. g., crevices, hollows, pits) on exposed rocky shores, often in vertical positions, upper to mid littoral level; home scars apparent (Fig. 38 T). Also found attached to shells of Scutellastra kermadecensis (Pilsbry, 1894) on Kermadec Islands (see Oliver, 1915: 547) and marine debris at LHI.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFFB827BFCCAFC22FC61FB36.taxon	description	(Figs 39 C – D, 40 A – J, R)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFFB827BFCCAFC22FC61FB36.taxon	materials_examined	Material examined. Type material. Lectotype of Siphonaria belcheri Hanley, 1858, present designation (NHMUK 1900.3.19.29, Fig. 40 A). Six paralectotypes, same data as lectotype (NHMUK 1900.3.19.28, 30 – 34, Figs 40 B – G). Other, non-type material. Pakistan: Karachi, Clifton Beach 24 ° 45.500 ’ N, 67 ° 05.968 ’ E, PA 02 - 1 (AM C. 585449 10 p, C. 585102 p [M 227], C. 585103 p [M 228], C. 585106 p [SK 147], C. 585500 p [SK 146], C. 595930 p [SK 533]); French Beach, 24 ° 50.367 ’ N, 66 ° 49.387 ’ E, PA 01 - 1 (AM C. 585382 10 p, C. 585100 p [M 237], C. 585101 p [M 238]), 24 ° 50.345 ’ N, 66 ° 49.244 ’ E, PA 01 - 2 (AM C. 585669 6 p).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFFB827BFCCAFC22FC61FB36.taxon	discussion	Taxonomic remarks. The largest syntype is herein designated as the lectotype of S. belcheri (Fig. 40 A) for the stabilisation of the name (Art. 74.1 of the Code). Our delineation of this species is based on comparative analyses of the morpho-anatomy and mitochondrial genetics of freshly collected topotypes (‘ Indian Ocean’, from Pakistan; Figs 40 H, J) and geographic series of additional specimens (Table S 1).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFFB827BFCCAFC22FC61FB36.taxon	description	External morphology. Foot wall, mantle and pneumostomal lobe evenly grey, foot sole darker grey; edge of foot sole / wall, cephalic folds and foot wall mantle join fade to pale cream; mantle translucent, wider than foot wall, elongated at anterior, edge thickened lobed whitish with black bands aligning to rib interstices; pneumostome within mantle, between right and posterior ADMs. Shell (Figs 40 A – J; Table S 9). Small sized (max sl mean = 12.08 mm, SD = 1.38 mm, n = 3), circular ovate; height medium; exterior maybe uneven, apex offset weakly to posterior and left, apical sides weakly concave, protoconch direction heterostrophic (n = 1), curls below apex to posterior, shell whorl dextral; growth striae prominent, shell thickness thick; rib count (mean = 33, SD = 3.3, n = 3), primary ribs pale white, fairly straight, ridge raised, rounded, may broaden significantly to shell lip, rib interstices; shell lip to scalloped and weakly corrugated; finer irregular secondary ribs fill gap between primary ribs, rib interstices brown; paired primary ribs form siphonal ridge. Interior shell margin golden brown with irregular mottled dark brown, shell lip white with brown rays under rib interstices variably extending over margin or to spatula, centre of spatula dark brown often orange; siphonal groove distinct, same colour as shell edge; ADM scar indistinct, CMS straight; thickening of shell lip occurs in more mature specimens, in-fills and reduces lip scalloping. Reproductive system (Fig. 39 C; n = 2). Positioned within right side of coelom, against foot wall on foot muscle, under the respiratory cavity; epiphallic parts positioned between BM and RAM. GA small, with singular GP through foot wall; AO large broad bluntly pointed, joined to upper GA; ED elongated narrow thickened, slightly centrally bent, joins to lower side of GA; GA, AO, ED all white muscular fibrous tissue; EG large, soft whitish tissue, folded, joins ED; single wide flagellum (F 1), shorter but similar width to ED, appears as an extension of ED. BD and CD connect in opposing directions into GA between ED join and GP, both ducts narrow long straight smooth whitish, pass together through RAM (BD over thicker CD) into soft white folded tissues of MG, BC embedded in folds close to embedded blackish SV; BD long narrow with prominent distal loop and MA to inner anterior foot wall; CD long, wider than BD; BC relatively small bulbous, thin whitish translucent test (yellowish gelatinous mass in BC); MG / AG complex relatively large; HD short narrow coiled, links ducts in soft white folded tissues of AG to yellowish granulated HG; AG / MG larger than HG, sides match curvature of inner foot wall. Spermatophore (Fig. 39 D). Broad head with short flagellum, coiled (length = 0.57 ± 0.09 mm, n = 2); head section cylindrical, bulbous, elongate, rounded tip; test smooth, featureless, translucent encasing a white opaque central core; tapering section merges head to flagellum; head longer, wider than translucent flagellum (head length = 0.51 ± 0.07 mm, ~ 90 % of SPM length, head width = 47 ± 9 μm; flagellum width = 6 ± 0 μm, n = 2); 2 SPM coiled in one BC (AM C. 595930). Radula. Dentition formula 12: 1: 12 (Hubendick 1946: 38).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFFB827BFCCAFC22FC61FB36.taxon	diagnosis	Comparative remarks. Siphonaria belcheri (atra group, unit 27) is the sister species of S. madagascariensis in our mitochondrial tree (Figs 1, 2). It differs from other species by COI distances of ≥ 20 % (Table S %). We found S. belcheri in sympatry with four congeners in Oman and Karachi, Pakistan. Siphonaria asghar has a lower shell with uneven ribbing, weaker edge scalloping, a larger BC, and a smaller, indistinct AO. Siphonaria perexigua sp. nov. has a lower shell with broader, more raised ribs, weaker edge scalloping, a shorter, bulbous, blunt AO, longer, wider ED and BD, and a larger BC. For comparisons with S. kurracheensis and S. crenata refer to comparative remarks under these species. The RS structure of S. atra and S. bifurcata resemble that of S. belcheri; however, these species differ in shell morphology and are genetically highly distinct. Figures of ‘ S. belcheri ’ in Hubendick (1946: 90, pl. 2, figs 5 – 8, from ‘ Persian Gulf’), are specimens of S. belcheri. However, the RS figured and described in Hubendick (1946: 10, fig. 7) differs from that shown herein in details of epiphallic parts, omitting AO, F 1, distal loop in BD and opposing connections of BD and CD. Hubendick’s RS figure and description matches that of S. carbo, which Hubendick (1946: 35) indicated was ‘ of the same type’. Figured specimen of ‘ S. kurracheensis ’ in Moazzo (1939: 7, 280 fig. 2 closely resembles S. belcheri (in particular the weakly ribbed paralectotype; Fig. 39 G). Figured specimen of ‘ unit 27, S. kurracheensis’ from Masirah Is, Oman in Dayrat et al. (2014: 263, fig. 5 D) matches the coarser ribbed form of S. belcheri rather than the narrower / finer ribbed S. kurracheensis.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFFB827BFCCAFC22FC61FB36.taxon	distribution	Distribution and habitat. Coasts of the Arabian Sea, recorded from Masirah Island, Oman, Karachi, Pakistan, and India (Fig. 37). In the present study found in sheltered positions on exposed rocky shores, upper to mid littoral level (Fig. 40 R).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFFA8276FCCAFA82FA25FA16.taxon	description	(Figs 39 I – J, 40 K – N, S, U – V)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFFA8276FCCAFA82FA25FA16.taxon	materials_examined	Material examined. Type material. Lectotype of Siphonaria nuttallii Hanley, 1858 b, present designation, from Hawaii (NHMUK 20120166 / 1, Fig. 40 K). Four paralectotypes, same data as lectotype (NHMUK 20120166 / 2 - 5, Fig. 40 N). Three syntypes of Siphonaria normalis forma chirura Pilsbry, 1921 from Kahoolawe, Hawaii [USA]; coll. H. A. Pilsbry, 1913 (ANSP 281800 a; ‘ lectotype’, refer Taxonomic remarks). Other, non-type material. Hawaii; Maui: Ho’okipa Beach, 20 ° 56.029 ’ N, 156 ° 21.411 ’ W HA 03 - 4 (AM C. 585444 11 p, C. 585661 3 p form chirura, C. 584894 p [M 470, protoconch G 7], C. 584895 p [M 471, SK 273, form chirura], C. 584896 p [M 472, SK 079], C. 584897 p [M 486, SK 305, form chirura], C. 584898 p [SK 211]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFFA8276FCCAFA82FA25FA16.taxon	discussion	Taxonomic remarks. The largest syntype of S. nuttallii is herein designated as the lectotype (Fig. 40 K) for the stabilisation of the name (Art. 74.1 of the Code). Our delineation of this species is based on comparative analyses of the morpho-anatomy and mitochondrial genetics of freshly collected topotypes (Fig. 40 L) and S. normalis var. chirura (Fig. 40 M) and geographic series of additional specimens (Table S 1). We establish S. normalis var. chirura as a new junior synonym. The designation of lectotype for S. chirura by Baker (1964: 159 ‘ ANSP TSD now: 281800 a’) is invalid as it is not based on a syntype (Art. 74.1 of the Code). The specimen is badly worn. Pilsbry (1920 b: 379) and Cernohorsky (1972: 210) erroneously treated S. nuttallii as a synonym of S. normalis.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFFA8276FCCAFA82FA25FA16.taxon	description	External morphology. Foot sole, foot wall, mantle, cephalic folds, pneumostomal lobe evenly grey / green, paler to foot edge; irregular black blotches of pigmentation on foot wall and concentrated over centre cephalic folds; mantle thin, translucent narrower than foot wall, covers exposed inner shell lip, wider at anterior, mantle edge thickened, dark edge band, weakly lobed with even black pigmentation rays aligned with underside of shell rib interstices; genital pore indistinct; small black epithelial eye spot centralised on each of centrally touching cephalic folds; pneumostomal lobe long broad under mantle between the right anterior and right posterior ADMs. Shell (Figs 40 K – N, S; Table S 9). Small sized (max sl mean = 12.7 mm, SD = 1.9 mm, n = 9), circular ovate; height medium; apex offset weakly posterior and laterally central, apical sides convex to straight; protoconch direction homostrophic to central (n = 3; Fig. 40 S), shell whorl dextral; growth striae prominent in bands, shell thickness thick; rib count (mean = 42, SD = 2, n = 9); exterior uneven with faint radial banding; primary ribs pale tan, fairly straight to wavy, broaden and weakly raised to shell edge, ridges and ends rounded, protrude beyond shell edge; edge finely scalloped and unevenly corrugated; 2 – 5 finer secondary ribs between primary ribs, rib interstices darker brown with white patches; siphonal ridge formed by paired primary ribs; Interior shell margin white rays under primary ribs, dark brown rays under rib interstices, rays extend from shell lip over shell margin fading to spatula; siphonal groove indistinct, paler than shell margin; spatula mottled dark brown to white; ADM scar distinct, dark brown, CMS convex; no thickening or whitening of shell lip observed. Reproductive system (Fig. 39 I; n = 4). Positioned within coelom under the respiratory cavity, hermaphroditic glands positioned to posterior against right foot wall and over foot sole, epiphallic parts positioned to anterior between BM and RAM; GP small, singular, positioned through foot wall behind right cephalic fold, GA small, prominent; AO large, elongate, broad, bluntly pointed, slightly bent centrally, joins to top of GA in conjunction with relatively short, strongly twisted, very broad ED; EG large white folded, single long narrow looped flagellum F 1 appears as an extension of ED at join with EG; AO, GA and ED all muscular white tissue; BD and CD connect closely in opposite directions into GA between connections of ED and AO, both ducts narrow smooth featureless, pass together through RAM connecting into MG, BD above and longer than CD, with distal loop and MA to inner foot wall; BC medium in size, spherical, embedded in folds of MG, test translucent; SV embedded in AG; HD reasonably long, thick coils, brown markings, links AG to smaller yellowish granular HG; MG and AG folded soft white tissue; sides match curvature of inner foot wall on right posterior of coelom; outer edge of MG lobed. Spermatophore (Fig. 39 J). Body cylindrical, thread-like (length = 6.7 mm, n = 1), test thin, translucent; head tip tapered bluntly rounded, section containing a white gelatinous core, tapers to a thin flagellum and tip; both sections smooth, featureless; head longer, thicker than flagellum (head length = 5.1 mm; 76 % of SPM length; flagellum length = 1.6 mm; head width = 95 μm; flagellum width = 16 μm), 2 SPM tightly coiled in white gelatinous mass in one BC [M 471].	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFFA8276FCCAFA82FA25FA16.taxon	diagnosis	Comparative remarks. Siphonaria nuttallii (plicata group, unit 85; Figs 1, 2) differs from other species by COI distances of ≥ 20 % (Table S 7). Siphonaria nuttallii occurs in sympatry with S. normalis at Big Island, Hawaii; refer to comparative remarks under that species. Siphonaria nuttallii has a similar shell morphology like other members of the plicata group, such as S. mauiensis sp. nov. (also occurring on Maui, but not found in immediate sympatry), S. plicata, S. tongatapuensis sp. nov. (both from Tonga), S. monticulus (from Lifou), S. namukaensis sp. nov. (Fiji and NC) and S. poindimiensis sp. nov. However, all these species are genetically well-differentiated. Specimens figured by Hubendick (1946: 91, pl. 4, figs 1 – 4) as S. nuttallii and S. chirura by Edmondson (1946: 188, figs 102 b) are consistent with typical characteristics of S. nuttallii as defined herein and are within the known distribution of this species. Specimens figured as ‘ S. normalis’ in Kay (1979: 493, figs 157 I, J) from Hawaii are likely of S. nuttallii for closely resembling the types (Figs 40 K – N).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFFA8276FCCAFA82FA25FA16.taxon	distribution	Distribution and habitat. Only known from Maui, Hawaii (Fig. 37), found on moderately exposed rocky shores in sheltered positions, mid littoral level (Fig. 40 U).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFF78270FCCAF9A2FE17FB16.taxon	description	(Figs 41 A – C, 42 H – L, O, Q, T – U)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFF78270FCCAF9A2FE17FB16.taxon	materials_examined	Material examined. Type material. Neotype of Siphonaria incerta Deshayes, 1863, present designation, from Cap de la Houssaye, Saint Paul, Réunion (Art. 76.1 of the Code) (NMHN IM- 2000 - 35954 [M 263], Fig. 42 H). Other, non-type material. Réunion. TS (AM C. 585204 [M 260], Fig. 42 I), Saint Denis, 20 ° 53.108 ’ S, 55 ° 28.577 ’ E, RU 01 - 1 (AM C. 585906 p [M 259, SK 276]); Cap de la Houssaye Saint Paul, 21 ° 01.086 ’ S, 55 ° 14.115 ’ E, RU 02 - 1 (AM C. 585204 p [M 260], C. 585205 p [M 261], C. 585206 p [M 262], C. 5859061 p [M 261]). Mauritius: Isle de la Passe, 20 ° 24 ’ S, 57 ° 46.133 ’ E (WAM S 72341 3 p); Souillac, 20 ° 31.467 ’ S, 57 ° 31.582 ’ E, MRU 01 - 2 (AM C. 585975 6 p, C. 584967 p [M 258]); Souillac, 20 ° 31.519 ’ S, 57 ° 31.633 ’ E, MRU 01 - 1 (AM C. 585974 17 p, C. 584964 p [M 253], C. 584965 p [M 255]). Australia, CI: E side Smith Point, Flying Fish Cove, 10 ° 25.749 ’ S, 105 ° 39.957 ’ E, CI 01 - 2 (AM C. 584701 7 p, C. 584840 p [M 320], C. 584841 p [M 321], C. 584842 p [M 322], C. 585203 p [SK 068]; WAM S 74099 3 p); West White Beach, 10 ° 27.748 ’ S, 105 ° 34.934 ’ E, CI 01 - 3 (AM C. 5847021 4 p, C. 584843 [M 309], C. 584844 p [M 310]; WAM S 74100 3 p); Ethel Beach, 10 ° 27.827 ’ S, 105 ° 42.497 ’ E, CI 02 - 1 (AM C. 584847 p [M 302], C. 584848 p [M 303], C. 584889 p [SK 081]; WAM S 74101 5 p).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFF78270FCCAF9A2FE17FB16.taxon	discussion	Taxonomic remarks. The type locality of S. incerta and S. parcicostata can be deduced from the title of article that contains the descriptions (= Réunion). The type specimens of both taxa are missing (V. Hero MNHN, pers comm.). Two original hand-drawn coloured figures of shells in Deshayes (1863: S. incerta (figs 16, 17) and S. parcicostata (figs 18, 19) provide the only indication of species identity. The neotype of S. incerta (Fig. 42 H) is designated herein to clarify the taxonomic status of this taxon (Art. 75.3.1 of the Code). Shell differences between the original figures of S. incerta and S. parcicostata are insignificant and within the observed range of variation of S. incerta (Fig. 42 H, I – L, O; see also Dautzenberg, 1932: 10). Therefore, S. parcicostata is herein treated as a junior synonym of S. incerta by First Reviewer’s Choice. Our delineation of this species is based on comparative analyses of the morpho-anatomy and mitochondrial genetics of freshly preserved specimens from Réunion, Mauritius, and Christmas Islands, including the neotype of S. incerta (Fig. 42 H, Table S 1). Hubendick (1946: 47) considered S. incerta Deshayes 1863 as a possible synonym of S. laciniosa (Linne, 1758) without having examined from Réunion or other Mascarene Islands. Morrison (1972: 56 – 58) treated S. incerta as a synonym of S. laciniosa based on similarity in shell form and ‘ common reproductive development’. This synonymy is not supported by our examinations.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFF78270FCCAF9A2FE17FB16.taxon	description	External morphology (Fig. 42 T). Foot sole, foot edge, foot wall, mantle, cephalic folds and pneumostomal lobe all evenly pale yellow to white in colour without any darker pigmentation markings. Mantle translucent, as wide as foot wall, strongly lobed with a thickened intense yellow to white banded edge; mantle lobes large, align with undulations of primary shell ribs, foot wall and pneumostome pustulose, pneumostome wide, between right anterior and posterior ADMs and within mantle; some individuals with dark brown shell rib interstices and interstice aligned dark pigmentation in mantle. Shell (Figs 42 H – L, O, U; Table S 9). Small to medium sized (max sl mean = 13.5 mm, SD = 2.4 mm, n = 15), ovate, flattened, height medium to tall; apex offset posterior and centre, apical sides convex, posterior side straight to weakly concave; protoconch direction heterostrophic to central (n = 3; Fig. 42 U), shell whorl dextral; growth lines indistinct; rib count (mean = 30.7, SD = 6.9, n = 15), raised, whitish, weakly wavy, width increases strongly to shell lip; rib interstices black to dark brown; 12 – 14 prominent primary ribs, extend up to 1 mm beyond shell edge, rib extremities weakly up turned; 1 – 2 finer secondary ribs may occur between primary ribs; paired primary ribs over siphonal ridge; shell lip uneven, scalloped aligning with protruding ribs; interior colouration matches white primary ribs and dark brown rib interstices, from shell lip to over shell margin to the chocolate to golden coloured spatula; ADM scar impression distinct, same as shell margin colouring, cephalic scar convex; thickening of shell occurs, white layering thickens and covers colouration of shell margin, spatula coated white. The neotype (Fig 42 H). Shell (sl = 11.5, sw = 9.1, sh = 3.9 mm) circular ovate; thin, apex offset weakly to posterior and left, ~ 18 whitish primary ribs, with 0 – 3 in between secondary ribs, rib interstices dark, siphonal ridge formed by adjacent dual primary ribs. Interior evenly dark brown, white rays on shell lip under ribs; RS (Fig. 41 A) and SPM (Fig. 41 B). Neotype specimen grouped within unit 72 (S. incerta). Reproductive system (Figs 41 A, C; n = 4). Positioned within coelom under the respiratory cavity, hermaphroditic glands positioned to posterior against right foot wall and over foot sole, epiphallic parts positioned to anterior between BM and RAM. Join of AO and GA distinct, AO larger than GA, elongated, bluntly pointed, slightly folded, thicker than ED; ED relatively short and narrow, EG elongated, bluntly pointed with a single very long, bent, twisted, thin flagellum F 1, flagellum often lays on top of the BM; AO, GA and ED all muscular white tissue; BD and CD jointly connected to GA between ED, AO and GP; BD slightly longer and thinner than CD with a prominent loop over GA, both ducts smooth and pass together through RAM connecting into MG (BD over CD), BC small and bulbous, HD short thickened coiled, links AG to a small elongated narrow yellowish granulated HG; MG and AG small, folded, soft white tissue; SV embedded on left side of AG, AG larger than HG, sides match curvature of inner foot wall at right posterior quarter of coelom. Spermatophore (Fig. 41 C). Long thread-like; head section (head length = 6.06 mm, head width = 80 μm, n = 1) cylindrical, bulbous, rounded tip, tapers to flagellum; test thin, smooth, featureless, translucent encasing a white opaque central core; a short tapering section merges head to filamentous flagellum; head section wider than translucent flagellum (incomplete); 2 SPMs tightly coiled in brown gelatinous mass in bursa of the neotype.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFF78270FCCAF9A2FE17FB16.taxon	diagnosis	Comparative remarks. Siphonaria incerta (atra group, unit 72) is the sister species of S. recurva (plus an unidentified Siphonaria from Tuituila Island) in our phylogenetic tree (Figs 1, 2). Both species differ from each other by COI distances of ≥ 16.7 %. From other species it differs by distances of ≥ 25 % (Table S 3). Throughout its range, we found seven congeners with partly sympatric occurrences. Five congeners are sympatric on CI: Siphonaria alba has a larger, lower, heavier shell with less raised ribs and a weaker scalloped shell edge, a smaller AO and BC, and a longer ED. Siphonaria christmasensis sp. nov. has a smaller, lower, darker shell with weaker ribbing and scalloped shell edge, dark spatula, and smaller AO and BC. Siphonaria delicata sp. nov. has a taller and brown / grey shell, with weaker ribbing and scalloped shell edge, and a larger BC. Siphonaria tenebrae sp. nov. has a lower shell with a less prominent siphonal ridge, stronger scalloped shell edge, and smaller AO and BC. Siphonaria umbra sp. nov. has a lower, paler shell with less raised ribbing, a weaker scalloped shell edge, darker spatula, a larger BC, and a shorter SPM. Two species are sympatric at Mauritius: For comparison with S. plana refer to comparative remarks under his species. Siphonaria griffithsorum sp. nov. has a smaller shell with less prominent siphonal ridge, even ribbing and weaker scalloped shell edge, and larger AO, ED and BC. Specimens from Mauritius and Port Natal figured as ‘ S. parcicostata’ in Hubendick (1946: 91, pl. 3, fig. 13 – 15) correspond well with features typical of S. incerta (Figs 42 H – L). However, their identity remains uncertain.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFF78270FCCAF9A2FE17FB16.taxon	distribution	Distribution and habitat. Known only from Réunion and CI (Fig. 37); found in sheltered positions on moderately exposed and exposed rocky shores, mid littoral level (Fig. 42 Q).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFF18272FF68FAA2FE81FE76.taxon	description	(Figs 40 O – Q, T, 41 D – G)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFF18272FF68FAA2FE81FE76.taxon	materials_examined	Material examined. Type material. Holotype of S. thersites from ‘ Neeah Bay, Washington, West Coast, North America’; coll. Swan, J. G. (USNM 11852; Fig. 40 O). Paratype, same data as holotype (MCZ 275190). Other, non-type material. USA, Alaska: Cook Inlet, Camel Rock Camel Rock, 59 ° 26.68 ’ N, 151 ° 43.02 ’ W (CBG 11 BIOAK- 0589 p [SK 554], 11 BIOAK- 0592 p [SK 553], 11 BIOAK- 0593 p). Canada, British Columbia: Houston Stewart Channel, Kunghit Island, 53 ° 0 ’ N - 132 ° 0 ’ W (RBCM 80283 d).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFF18272FF68FAA2FE81FE76.taxon	discussion	Taxonomic remarks. Carpenter (1864 b: 423) donated the types to the Smithsonian Institution (i. e., USNM). Palmer’s (1958: 258) statements regarding missing type and potential lectotype specimens are incorrect and explicitly not based on examination of the holotype. Siphonaria thersites is the type species of Liriola Dall, 1870, by original designation.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFF18272FF68FAA2FE81FE76.taxon	description	External morphology. Animal not fully enclosed by shell, foot sole pale grey, foot wall, foot edge, mantle and cephalic folds all darker grey, paler at foot edge; foot wall pustulose, without darker markings; mantle narrow with dark grey edge band, half as wide as foot wall; two small indistinct black epithelial eye spots centralised on two centrally touching cephalic folds, pneumostomal lobe thick, under mantle behind right cephalic fold; closes the pneumostomal and anal openings at the mantle edge. Shell (Figs 40 O – Q; Table S 9). small sized (max sl mean = 10.6 mm, SD = 1.7 mm, n = 9), elongate ovate, apex offset strongly posterior and left, apical sides strongly convex, height low, protoconch below apex, close to posterior edge; protoconch direction homostrophic (n = 2), shell whorl dextral; exterior uneven, radially ribbed, reddish brown, growth striae prominent in shaded radial bands, shell thin, lip even fragile, periostracum freely extending; rib count (mean = 36, SD = 3.2, n = 9), ribs weak to indistinct, primary ribs flatly rounded, not protruding beyond shell lip; often interspersed finer secondary ribs, rib interstices darker; siphonal ridge clear, bulged, extends beyond shell edge, formed by paired primary ribs. Interior glossy, shell margin dark brown to tan, lip paler with white markings aligned under ribs, siphonal groove distinct, shallow, same colour as margin; spatula mottled tan, uneven darker markings; ADM scar distinct, CMS convex; thickening of shell lip not observed. Reproductive system (Figs 41 D, F, G; n = 3). Positioned within coelom under the respiratory cavity, hermaphroditic glands positioned to posterior against right foot wall and over foot sole, epiphallic parts positioned to anterior between BM and elongated RAM; AO indistinct, appears as a bulge under medium sized GA, ED short wide, joins to side of GA, singular prominent GP; EG large bulbous soft, flagellum (F 1) indistinct to absent; BD indistinct, very narrow, enclosed in ventral tissue of wider whitish CD from BC to GA (Fig. 41 F – G); both ducts are of similar length, emerge from folds of MA, together pass between foot wall and outside of RAM, and join into side of GA; GA, ED and BD all muscular white tissue; BC small deflated flattened whitish or expanded (5 SPM in a single BC), positioned under MG / AG; MG and AG small, heavily folded, soft white tissue; yellowish SV embedded on left side of AG, AG larger than HG; HD short, thickened, uncoiled and unlobed, links AG to a small, elongated, brownish / yellow, finely granulated HG. Spermatophore (Fig. 41 E). Body cylindrical, thread-like (length = 14.6 mm, n = 1, 118 % of AL), test thin, translucent; head section long even, bluntly rounded, tapers to a thin flagellum and tip; both sections smooth, featureless; head longer, thicker than flagellum (head length = 11.8 mm; 81 % of SPM length; flagellum length = 2.8 mm; head width = 150 μm; flagellum width = 14 μm). Radula. Dentitionformula 22: 1: 22 or 7.3.3.9: 1: 9.3.3.7 (Dall, 1870: 33), 24: 1: 24 (Hubendick 1946: 20).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFF18272FF68FAA2FE81FE76.taxon	diagnosis	Comparative remarks. Siphonaria thersites (unit 42) is the sister species of all other Siphonaria species included herein (Fig. 1). It differs from other species by COI distances of ≥ 15.6 %. It is not known to occur in sympatry with any other congener. As the only species found in Alaska, it has not been mistaken with other species in previous taxonomic literature. Contrary to Hubendick (1945: 15, fig. 1; 1946: 8, figs 1, 3), a BC is present and the genital pore (GP) is monoaulic (Fig. 41 D, F). Hence, the general layout of the reproductive anatomy of S. thersites corresponds well with that of other Siphonaria species.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFF18272FF68FAA2FE81FE76.taxon	distribution	Distribution and habitat. Endemic to northern hemisphere temperate zone from Kurile Islands, Russia, to west coast of Alaska, USA, British Columbia, Canada and Washington USA (Fig. 45). Found on sheltered rocky shores at lower littoral level often on Fucus rockweed (Fig. 40 T).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFF3826DFF68FE42FEC0FC96.taxon	description	(Figs 41 H – I, 42 A – G, M – N, P, R – S)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFF3826DFF68FE42FEC0FC96.taxon	materials_examined	Materialexamined. Typematerial. Neotypeof Siphonaria tasmanica Tenison Woods, 1877, present designation, from Tasman Arch, Tasmania (Art. 76.1 of the Code); coll. B. W. Jenkins, T 03 - 3, 27 March 2018 (AM C. 585259, Fig. 42 A [M 114], condition Art. 75.3.7 of the code). Seven syntypes of Siphonaria zonata Tenison Woods, 1877 from S Tasmanian coast; coll. J. E. Tenison Woods, 1877 (MV F 686, Figs 42 B – C, M – N; AM C. 103723, 2 d). More than twenty syntypes of Talisiphon tasmanicus nereis Iredale, 1940 from Port Fairy, Vic, [Australia] coll. R. Bell, 1918 – 1919 (AM C. 108499, Figs 42 D, E). Holotype (measurements in text) of Talisiphon tasmanicus turritus Iredale, 1940 from Macquarie Harbour, Tas; coll. A. F. Basset Hull, 1922 (AM C. 595951, Fig. 42 F). Thirteen paratypes, same data as holotype (AM C. 53828; labelled ‘ syntypes’). Other, non-type material. Australia, Vic: Cape Schanck, 38 ° 29.951 ’ S, 144 ° 53.369 ’ E, V 06 - 4 (AM C. 585458 13 p); Point Lonsdale (nr Queenscliff), 38 ° 17.276 ’ S, 144 ° 36.977 ’ E, V 05 - 1 (AM C. 585730 p [M 120]). Tas: Lagoon River: mouth, 41 ° 29.4 ’ S, 144 ° 49.2 ’ E (TMAG E. 41995 2 d); Bicheno, Redbill Beach, 41 ° 51.6 ’ S, 148 ° 17.4 ’ E (TMAG E. 42002 p, d); Maria Island, Howells Point & Painted Cliffs, 42 ° 35.796 ’ S, 148 ° 2.886 ’ E (TMAG E. 42001 p, d); Maria Island, Trigonia Corner, 42 ° 41.196 ’ S, 148 ° 4.404 ’ E (TMAG E. 41993 d, p); Marion Bay, northern beaches 42 ° 45.048 ’ S, 147 ° 53.592 ’ E (TMAG E. 42004 d, p); Park Beach Dodges Ferry, 42 ° 51.716 ’ S, 147 ° 36.665 ’ E, T 03 - 4 (AM C. 585429 10 + p, C. 585266 p [SK 549 protoconch D 6]); Carlton Beach, Spectacle Island, 42 ° 52.044 ’ S, 147 ° 36.024 ’ E (TMAG E. 41994 d, p); Taroona, Dixons Beach, 42 ° 56.358 ’ S, 147 ° 21.414 ’ E (TMAG E. 41999 d, p); Lagoon Bch (near Saltwater River), 42 ° 56.903 ’ S, 147 ° 39.962 ’ E, T 03 - 2 (AM C. 585659 4 p, C. 585876 d); Taroona Beach; 42 ° 57 ’ S, 147 ° 21 ’ E (TMAG E 03651 3 p); 42 ° 57.18 ’ S, 147 ° 21 ’ E (TMAG E. 42006 4 d, 4 p): Kingston Beach, 42 ° 58.8 ’ S, 147 ° 19.2 ’ E (TMAG E. 02013 p); Eaglehawk Neck, eastern side, 43 ° 0.444 ’ S, 147 ° 56.082 ’ E (TMAG E. 42003 d, p), (TMAG E. 01659 2 p); Blackmans Bay 43 ° 0.6 ’ S, 147 ° 19.8 ’ E (TMAG E. 41998, 3 d / 3 p), (TMAG E. 15880 10 p); Calverts Beach & Goats Bluff, 43 ° 1.65 ’ S, 147 ° 29.07 ’ E (TMAG E. 42005 3 d, 3 p); South Arm – Hope Beach, southern end, 43 ° 1.8 ’ S, 147 ° 27.6 ’ E (TMAG E. 01245 3 p); Tasman Arch, 43 ° 02.033 ’ S, 147 ° 56.963 ’ E, T 03 - 3 (AM C. 585728 9 p, C. 585255 p [SK 020], C. 585259 p [M 114]); Tinderbox Beach, 43 ° 3.6 ’ S, 147 ° 19.8 ’ E (TMAG E. 05224 p); North Bruny Island, Dennes Point, 43 ° 3.87 ’ S, 147 ° 21.066 ’ E (TMAG E. 42000 4 p, 4 d); Nubeena, Parsons Bay, 43 ° 6 ’ S, 147 ° 6 ’ E (TMAG E. 05996 2 p); Nubeena, White Beach, 43 ° 7.2 ’ S, 147 ° 43.8 ’ E (TMAG E. 06015 p); Fortescue Bay, 43 ° 8.4 ’ S, 147 ° 57.6 ’ E (TMAG E. 05580 6 p); Port Arthur, 43 ° 9 ’ S, 147 ° 52.2 ’ E (TMAG E. 05550 4 p); Three Hut Point d’Entrecasteaux Channel, 43 ° 16.195 ’ S, 147 ° 14.414 ’ E, T 04 - 3 (AM C. 595913 2 p); South Bruny Island: Simpsons Bay, 43 ° 17.4 ’ S, 147 ° 18.6 ’ E (TMAG E. 04867 4 p), Cloudy Beaches – eastern beach, 43 ° 26.352 ’ S, 147 ° 14.202 ’ E (TMAG E. 41996 20 d, 20 p), (TMAG E. 25061 d); Peaches Point, 43 ° 34.122 ’ S, 146 ° 55.037 ’ E, T 05 - 3 (AM C. 585606 4 p); Flensing Rock, 43 ° 34.291 ’ S, 146 ° 54.856 ’ E, T 05 - 2 (AM C. 585460 13 p, C. 585512 p [SK 080], C. 585877 p [M 119]); Cockle Creek Bay, 43 ° 34.8 ’ S, 146 ° 53.4 ’ E (TMAG E. 32706 d); Pancake Bay, 43 ° 34.673 ’ S, 146 ° 55.293 ’ E, T 05 - 5 (AM C. 585715 8 p); Trial Harbour, 41 ° 55.758 ’ S, 145 ° 10.434 ’ E (TMAG E. 41997 5 d, 5 p); Lucas Point, Pilot Bay, Macquarie Harbour, 42 ° 12.241 ’ S, 145 ° 12.005 ’ E, T 06 - 1 (AM C. 585483 18 p; C. 585538 20 + p, C. 585878 p [M 171], C. 585879 p [M 172]). SA: Fishery Bay Cape Wiles, 34 ° 55.107 ’ S, 135 ° 41.086 ’ E, SA 05 - 1 (AM C. 585689 7 p); Haleys Beach Gibson Peninsula, 32 ° 45.084 ’ S, 134 ° 05.490 ’ E, SA 03 - 4 (AM C. 585467 20 + p); Wandrilla Beach, nr Cape Nuyts, 32 ° 01.894 ’ S, 132 ° 16.052 ’ E, SA 01 - 1 (AM C. 585705 10 + p, C. 585208 p [SK 019]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFF3826DFF68FE42FEC0FC96.taxon	discussion	Taxonomic remarks. The type locality was not explicitly stated in the original designation of S. tasmanica but is evident from the title of the work (“ on Tasmanian Patellidae ”). Originally, Tenison Woods (1877: 45) described S. tasmanica as a variety of S. denticulata. Subsequently, Tenison Woods (1878 b: 99) described the same taxon again as S. zonata without mentioning the earlier introduced name S. diemenensis var. tasmanica. No original types of S. tasmanica are known to exist and we could not locate any in the collection of the AM. The neotype of S. tasmanica (Fig. 42 A) is designated herein to clarify the taxonomic status of this taxon (Art. 75.3.1 of the Code). Our delineation of this species is based on comparative analyses of the morpho-anatomy and mitochondrial genetics of freshly collected topotypes and geographic series of additional specimens (Table S 1). These analyses confirm the synonymy of S. zonata (Fig. 42 B – C, M – N) and establish Talisiphon tasmanicus nereis and T. tasmanicus turritus (Fig. 42 D – F) as new synonyms. The name ‘ Siphonaria zonata (‘ Schub. et Wagn’) ’ in Deshayes (1843: 31, pl. 62, fig. 17 – 18) is misapplication for Patella zonata Schubert & Wagner, 1829. The figured specimens in Schubert & Wagner (1829) and Deshayes (1843) differ from one another, neither showing a species of Siphonaria. Christiaens (1975: 91) listed Patella zonata as a synonym of Scurria scurria (Lesson, 1830), Lottiidae. Iredale (1924: 276) was the first to recognize the synonymy of S. zonata and S. tasmanica transferring the species to Talisiphon. Tate & May (1901: 419) incorrectly considered S. zonata as a synonym of S. funiculata.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFF3826DFF68FE42FEC0FC96.taxon	description	External morphology (Fig. 42 S). Foot sole and foot wall evenly dark grey, paler at foot / wall edge; foot wall and mantle blue-green-grey; fringing mantle narrow, unlobed, translucent, covers exposed inner shell lip; genital pore inconspicuous, located on foot wall to right anterior of right cephalic fold; two small black epithelial eye spots centralised on two centrally touching cephalic folds; pneumostomal lobe small, thin, part of the mantle, between the right anterior and right posterior ADMs, closes the pneumostome at the mantle edge. Shell (Figs 42 A – G, M – N, P, R; Table S 9). small to medium sized (max sl mean = 13.8 mm, SD = 2.4 mm, n = 12), height tall; apex offset weakly to posterior and left, often eroded and appearing as a white spot, apical sides convex; protoconch direction homostrophic (n = 2; Fig. 42 R), shell whorl dextral; growth striae indistinct; rib count (mean = 48.3, SD = 9.6, n = 11), primary ribs fairly straight, unraised, flattened, broad, few secondary ribs; rib interstices distinct dark brown narrow lines extending to shell lip. Exterior shell colouration very distinct and unlike any other siphonariid; 3 prominent colour bands align with shell growth dividing shell height unevenly into thirds; top band upper half of shell dark brown, mid band widest and pale blue, bottom band greenish blue, limited to shell margin (Figs 42 A, G). Internal colouration banded and variable; spatula blueish or cream / white, shell lip markings white aligned under primary ribs, reddish brown aligned under rib interstices; siphonal groove same colour as spatula, ADM scar dark brown, shell margins immediately above and below are paler; CMS straight. Shell thickening not observed. Shell height may be variable; e. g. lower in S. t. turritus (Fig. 42 G) and taller in S. blainvillei (Fig. 42 F) forms. The neotype (Fig. 42 A). Shell medium sized (sl = 17.5, sw = 14.3, sh = 8 mm), circular ovate, tall; medium thickness, apex offset strongly to posterior and weakly to left, ~ 52 mainly primary ribs, few secondary ribs, interior dark brown, spatula and shallow siphonal groove white to bluish; taller and slightly darker interior shell form of S. tasmanica. Neotype specimen grouped within unit 76 (S. tasmanica). Reproductive system (Figs 41 H; n = 2). Positioned against inside of foot wall and over foot sole on the right posterior quarter within coelom, under the respiratory cavity. GA, EG and ED positioned in coelom between BM and RAM. GA very large, smoothly bulbous, with singular GP; AO absent; ED very short, broad, curved, unfolded, joins to side of GA; GA and ED white muscular fibrous tissue; EG soft whitish, slightly folded, smaller than GA; flagellum absent; BD and CD relatively short (BD longer), slightly curved, smooth, featureless, connect to GA together, pass together through RAM (BD over CD), CD connecting into MG / AG; BC small, elongated, bulbous, thin test, embedded in lower folds of MG / AG; HD short, narrow, coiled, links smallish AG to a large elongated yellowish granulated HD; MG / AG complex small, soft white tissue folds, enveloping SV close to embedded BC, AG larger than HG, HG side reflects curvature of inner foot wall. AL = 13.52 mm. Spermatophore (Fig. 41 I). Length very short compared to other congenors; broad head with short flagellum (length = 1.03 mm, n = 1); head section cylindrical, bulbous, rounded tip (head length = 0.64 mm, ~ 62 % of SPM length; head width = 159 μm; flagellum width = 34 μm), contains whitish core, test thin, translucent encasing a white opaque coiled core; flagellum transparent, tapering to a thread-like end; both sections uneven, featureless; 1 SPM in brown gelatinous mass of one BC [SK 080]. Radula. (Fig. 83 M – P) Mean dentition formula 43: 1: 43 (SD = 4.8) with 115 (SD = 26.6), mean transverse rows 115 (SD = 26.6, n = 5); single central rachidian tooth flanked squarely by 43 half row laterals, 0 – 4 are inner (Figs 83 M – N), 14 – 20 mid and 15 outer laterals (Fig. 83 O); central tooth relatively long (half basal length) with narrow unicuspid mesocone; inner laterals (without endo or ectocones) may occur, mesocones of inner and mid laterals single pointed, mid laterals with broad pointed ectocone protruding at an acute angle halfway along the tooth’s length; outer laterals typically with a weakly bicuspidate ‘ chisel’ shaped mesocone flanked by small, pointed single ecto and endocones, angle of separation of each cone from the mesocone varies (Fig. 83 O).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFF3826DFF68FE42FEC0FC96.taxon	diagnosis	Comparative remarks. Siphonaria tasmanica (lateralis group, unit 76) is most closely related to S. lessonii from the Southern Ocean (not revised herein, see Güller et al. (2015: 81), S. funiculata and S. obliquata (Figs 1, 4). It differs from S. lessonii by COI distances of ≥ 9.4 % and from S. funiculata of ≥ 8.5 % (Table S 8). Throughout its range, S. tasmanica has been found in sympatry with five congeners in southeastern Australia. For comparisons with S. diemenensis, S. funiculata, and S. zelandica refer to comparative remarks under these species. Siphonaria jeanae has a lower, grey-blue shell with more prominent unraised brown ribs, a more scalloped edge, a wider ED, larger BC, and more thread-like SPM. Siphonaria stowae has smaller, lower, paler shell with an apex strongly offset to posterior, more prominent ribbing, a larger AO and BC, a smaller ED and a more thread-like SPM. Overall, the combination of shell geometry, size and colouration render S. tasmanica a rather distinctive species. Tasmanian records of ‘ S. tristensis Sowerby I, 1823 ’ in Tate & May (1901) are incorrect and based on misidentified specimens of S. tasmanica. A record of ‘ S. tasmanica ’ from Percy Island, Qld (Singleton, 1937: 396) is likely a misidentification of S. normalis. A specimen figured as S. tasmanica in Davey (1998: 118) is a specimen of S. funiculata.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFF3826DFF68FE42FEC0FC96.taxon	distribution	Distribution and habitat. Endemic to cool temperate coasts of southern Australia, between Mallacoota, Vic, and Gibson Peninsula, SA, Tas (Fig. 37). Found in sheltered positions (e. g., rock hollows, crevices, cracks) on very exposed rocky shores, mid littoral level (Fig. 42 P); frequently associated with black mussels; home scars prominent.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFEC826EFF68FC22FB00FDB6.taxon	description	(Figs 43 A – D, M – N, 44 A – C)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFEC826EFF68FC22FB00FDB6.taxon	materials_examined	Material examined. Type material. Lectotype of Siphonaria acmaeoides Pilsbry, 1894 from Prov. Boshiu [Boso Peninsula], Japan; coll. Frederick Stearns (ANSP 70726 a, Fig. 43 A). Two paralectotypes same data as lectotype (ANSP 70726). Holotype of Siphonaria (Mouretus) acmaeoides paulae Christiaens, 1980 from Ping [Peng] Chau, [Hong Kong, China] (NHMUK 1977171, Fig. 43 D). Other, non-type material. Japan, Honshu: Boso Peninsula, Po int S of Chitose Beach, 34 ° 59.240 ’ N, 139 ° 58.304 ’ E, JP 02 - 2 (AM C. 585393 10 + p, AM C. 584936 p [M 496, SK 315], C. 584937 p [M 500, SK 319], C. 585289 p [SK 356], C. 585513 p [SK 334 protoconch H 4], C. 585918 p [SK 335]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFEC826EFF68FC22FB00FDB6.taxon	discussion	Taxonomic remarks. The description of Pilsbry (1894 b: 16) does not contain an original type designation. Subsequently, Pilsbry (1895: 2, pl. 6, fig. 19 – 22) republished the original description based on three specimens but giving the dimensions for only one specimen. Baker (1964: 159) designated the lectotype (ANSP 70726 a), which matches the original dimensions given by Pilsbry (1894 b: 16). Figures in Pilsbry (1895: pl. 6, figs 19 – 22) correspond reasonably well with the type specimens; fig. 19 (ventral) lectotype, figs 20 (ventral), 21 (dorsal) and fig, 22 (ventral) paralectotypes. The type specimen figured in Higo et al. (2001: fig. G 4976, “ ANSP 70726 ”) differs from the specimens in Pilsbry (1894 b: 16). The type specimens (ANSP 70726) of S. acmaeoides (Fig. 43 A) are of the plicata group. Our delineation of this species is based on comparative analyses of the morpho-anatomy and mitochondrial genetics of freshly collected topotypes (Fig. 43 B, Table S 1). Comparative morpho-anatomy and mitochondrial phylogenetics herein confirm that S. acmaeoides and S. zelandica arecloselyrelated, yetdistinctspecies. Christiaens (1980 a: 79) recorded S. acmaeoides from Hong Kong and described a new subspecies, S. acmaeoides paulae Christiaens (1980 a: 79, pl. 4, figs B, D). The holotype of S. a. paulae (Fig. 43 D) matches specimens of S. acmaeoides from Japan (Fig. 43 B). The emphasised characters of this subspecies, a finer, thinner, more elliptical, lighter coloured shell without a ‘ marked’ central area, are within the range of intraspecific variation. Therefore, we synonymize this taxon with the nominate form. Christiaens (1980 a) record extends the distribution of S. acmaeoides from Honshu, Japan to Hong Kong (Fig. 35).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFEC826EFF68FC22FB00FDB6.taxon	description	External morphology (Fig. 43 N). Foot sole dark grey, paler to foot edge; foot wall, foot edge, mantle, cephalic folds and pneumostomal lobe all evenly pale grey / yellowish in colour; mantle thin, translucent, wider than foot wall, weakly lobed and unbanded edge; no black / dark pigmentation; pneumostome fold large, long between right ADMs and within mantle; vivid yellow or white subepithelial pustules on cephalic lobes and pneumostomal fold; two black ‘ Eye’ spots prominent centrally on thickened cephalic lobes. Shell (Figs 43 A – C, M; Table S 9). Medium sized (max sl mean = 14.5 mm, SD = 1.9 mm, n = 8), circular ovate; height low; apex offset slightly posterior and laterally central, apical sides even, weakly convex; exterior pale brown, with irregular darker / black flecks between primary ribs; rib count (mean = 57.4, SD = 10.3, n = 8), ~ 15 primary ribs, whitish, fairly straight, ridges rounded, broaden to shell edge; 3 – 4 finer secondary ribs between primary ribs, rib interstices dark grey; ribs align with shell edge; growth striae indistinct; siphonal ridge not prominent, formed by paired primary and secondary ribs, more protruding at shell edge; weak radial banding, darker to shell edge; protoconch direction central flat (n = 1; AM C. 585513 [SK 334]), shell whorl dextral. Interior shell lip finely corrugated; shell lip and margin white with dark flecks to dark chocolate brown rays aligning under primary rib interstices; spatula colour variable golden to dark chocolate brown; siphonal groove and ADM scar prominent, paler than margin and spatula; CMS convex to straight, indistinct; thickening and whitening of shell margin occurs. Reproductive system (Figs 44 A – B; n = 4). Positioned within coelom under the respiratory cavity, hermaphroditic glands positioned to posterior against right foot wall and over foot sole, epiphallic parts positioned between BM and RAM; GA large, AO indistinct absent; ED very short, twisted, very broad; EG very large with folds, single long narrow looped flagellum F 1 very short appears as an extension of ED at join with EG; GA and ED all muscular white tissue; BD and CD connect in parallel into GA close to ED joint, both ducts narrow, smooth, featureless, similar length pass together between outer RAM and inner foot wall (BD above CD), slightly bent before connecting into folds of MG; BD without distal loop or MA; BC small, translucent test, bulbous, embedded in MG; coiled brownish HD links white AG to finely granulated HG; MG and AG small folded soft white tissue; SV embedded within AG under BC; AG slightly smaller than HG, sides reflect curvature of inner foot wall at right posterior quarter of coelom. Spermatophore (Fig. 44 C). Body bulbous, elongated (length = 1.03 mm, n = 1, AL = 11 mm), test thin; head tip bluntly rounded, section containing a white gelatinous core, tapers to a thin flagellum and tip; both sections smooth, featureless; head longer, thicker than flagellum (head length = 0.89 mm; 87 % of SPM length; flagellum length = 0.14 mm; head width = 243 μm; flagellum width = 24 μm), 1 SPM in white gelatinous mass in BC of one topotypic specimen. Radula. Dentition formula 26: 1: 26 (Hubendick 1946: 31).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFEC826EFF68FC22FB00FDB6.taxon	diagnosis	Comparative remarks. Siphonaria acmaeoides (atra group, unit 91) is the closely related sister species of S. zelandica (Figs 1, 2). Both species differ by COI distances of ≥ 5.8 % (Table S 4). Both species have a disjunct distribution with one species found in the northern (S. acmaeoides) and one in the southern Pacific (S. zelandica), each. Throughout its range, S. acmaeoides has been found in sympatry with three congeners. For comparisons with S. sirius and S. japonica refer to comparative remarks under these species. Siphonaria camura sp. nov., sympatric in Honshu and Hong Kong, has a smaller, taller shell with more raised ribbing, a darker brown interior, larger BC, and a larger, thread-like and barbed SPM. Siphonaria acmaeoides exhibits a shell morphology similar to other species of the plicata group; however, these are anatomically and genetically distinct: S. zelandica (temperate Australia), S. plicata (Tonga), S. nuttallii (Hawaii), S. tongatapuensis sp. nov. (Tonga), S. namukaensis sp. nov. (Fiji and NC) and S. poindimiensis sp. nov. (NC), S. yagasaensis sp. nov. (Fiji) and S. monticulus (NC, Lifou). Siphonaria acmaeoides resembles S. zelandica in shell sculpture, external morphology, and SPM. Both species occupy similar habitats (upper littoral, shallow rock pools, rarely on rock faces). However, closer examination of the type and topotypic specimens revealed that S. acmaeoides differs in shell geometry, wider rib ridges, external colouration, and secondary ribbing, larger size of GA and ED, smaller HD, longer and narrower BD and CD. Hubendick (1946: 31) correctly pointed out that ‘ S. acmaeoides ’ and ‘ S. bifurcata ’ (= S. zelandica) had ‘ very similar shells’. Dayrat et al. (2015: 268) considered both taxa as possibly conspecific based on similarities in shell morphology.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFEC826EFF68FC22FB00FDB6.taxon	distribution	Distribution and habitat. Recorded from the type locality, Bose Peninsula, and Aichi Prefecture, Honshu, Japan (Fig. 45). In this study found to be common on exposed rocky shores in crevices and small rock pools, upper littoral level (Fig. 43 M).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFEF8269FCCAFD02FD5FFBD6.taxon	description	(Figs 43 E – F, O – P, 44 D – E)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFEF8269FCCAFD02FD5FFBD6.taxon	materials_examined	Material examined. Type material. Lectotype of Siphonaria sirius Pilsbry, 1894 a from Sagami and Kashiurazaki, Boshiu, Japan; coll. Fredrick Stearns (ANSP 70720 a, Fig. 43 E). Six paralectotypes same data as lectotype (ANSP 70720). Seven paralectotypes of Siphonaria sirius Pilsbry, 1904 from Sagami Bay (AM C. 117637). Other, non-type material. Japan, Honshu: Point S of Chitose Beach, Boso Peninsula 34 ° 59.240 ’ N, 139 ° 58.304 ’ E, JP 02 - 2 (AM C. 585395 10 p, C. 584940 p [M 501, SK 320], C. 584941 p [M 502, SK 321], p [SK 336]). China, Hong Kong: Cape D’Aguilar 22 ° 12.28 ’ N, 114 ° 15.38 ’ E (ZRC 2001 - 1768 21 p, ZRC. MOL. 24902 p [SK 176]). Philippines: Mactan, Cebu 10 ° 18.840 ’ N, 124 ° 01.707 ’ E PHS 04 - 1 (AM C. 585339 p). Singapore: Lazarus Island 01 ° 18.643 ’ N, 103 ° 57.077 ’ E SI 04 - 2 (AM C. 585227 p [M 339]); East Coast Park, seawall, 01 ° 18.643 ’ N, 103 ° 57.077 ’ E SI 01 - 2 (ZRC Moll. 9121, p). Indonesia: Pulau Panjang, Riau Islands 1 ° 10.21 ’ N, 104 ° 18.905 ’ E (ZRC EA-ZJ 09, 4 p).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFEF8269FCCAFD02FD5FFBD6.taxon	discussion	Taxonomic remarks. The description of Pilsbry (1894 b: 9) does not contain an original type designation. Pilsbry (1895: 2, pl. 6, fig. 23 – 28) republished the original description with six figures and dimensions for a single specimen. Baker (1964: 159; ANSP 70720 a) subsequently designated the lectotype, which matches the dimensions given in Pilsbry (1894 a: 9) and the figures in Pilsbry (1895: 2, pl. 6, figs 23 – 24; Fig. 43 E herein). A type was also figured in Higo et al. (2001: 142, fig. G 4976). Our delineation of this species is based on comparative analyses of the morpho-anatomy and mitochondrial genetics of freshly collected topotypes (Fig. 43 F) and geographic series of additional specimens (Table S 1). The description of ‘ S. atra ’ in Reeve (1856: pl. 3, species 14) appears to be based on specimens of S. sirius and not S. atra (refer to taxonomic remarks under S. atra). Hubendick (1946: 50) listed ‘ transitional’ forms between ‘ S. sirius <> S. subatra <> S. zanda ’, not explicitly linked to a specimen or locality. The figured specimen (Hubendick 1946: pl. 5, fig. 3, 4) is identified as a specimen of S. sirius. Je (1989: 29) incorrectly listed S. subatra as a synonym of Anthosiphonaria sirius (i. e., S. sirius). Christiaens (1980 a: 79) considered S. sirius as one of three ‘ forms’ of S. laciniosa in Hong Kong. He stated that the shell possesses ‘ heavier ribbing than atra, with six or more solid white ribs’ and that the ‘ siphon is formed by one rib’, which is consistent with features typical of S. sirius (Figs 43 E, F).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFEF8269FCCAFD02FD5FFBD6.taxon	description	External morphology (Fig. 43 O). Foot wall, cephalic folds and pneumostomal lobe all evenly cream in colour, paler to foot edge, foot sole grey; mantle thin, translucent, wider than foot wall, weakly lobed with a thickened edge, white bands on mantle edge align with underside of ribs; irregular black blotches of pigmentation on foot wall and concentrated over cephalic lobes; two black ‘ Eye’ spots prominent centrally on thickened cephalic lobes; pneumostome fold prominent with white subepithelial pustules. Shell (Figs 43 E – F; Table S 9). Medium sized (max sl mean = 18.6 mm, SD = 8.97 mm, n = 9), elongate ovate; height medium; apex offset central sightly left, apical sides straight to weakly convex, protoconch direction homostrophic (n = 1), shell whorl dextral; growth striae prominent in bands, weak shades of radial banding may occur, shell thickness medium; rib count (mean = 41, SD = 3.9, n = 9), 9 – 10 primary ribs pale white, straight, broad, raised and protrude strongly (some> 1 mm) beyond shell lip to prominently scallop and corrugate the edge; 2 – 4 finer brown secondary ribs between primary ribs, interstices narrow, darker, single primary rib forms siphonal ridge. Interior shell margin dark brown to tan, white rays align on shell margin under primary / secondary ribs, siphonal groove distinct, same colour as shell edge, points to right anterior; spatula white, some specimens may be dark chocolate brown; ADM scar distinct, CMS straight, paler than shell lip; thickening of shell lip translucent, infills and reduces lip scalloping, spatula becomes whitened. Reproductive system (Fig. 44 D; n = 1). Positioned within coelom under the respiratory cavity, hermaphroditic glands positioned to posterior against right foot wall and over foot sole, epiphallic parts positioned over back of BM and to side of RAM; AO prominent, broad bluntly pointed, joins to top of GA; ED relatively short, strongly twisted, very broad with MA on bend; EG small; single short broad looped flagellum F 1 appears as an extension of ED at join with EG; AO, GA and ED all muscular white tissue; BD and CD with opposing connections (bulbous at CD) join into GA between ED, AO and GP; BD longer and narrower than CD with a prominent distal loop without an MA, and looped immediately before BC, both ducts smooth and pass together through RAM connecting into folds of MG (BD above CD); BC embedded in MG, translucent test, large and bulbous; HD brownish long coiled links AG to a small elongated narrow brownish coarsely granulated HG; MG and AG small folded soft white tissue; SV embedded within AG, AG larger than HG, sides match curvature of inner foot wall at right posterior quarter of coelom. Spermatophore (Fig. 44 E). Body cylindrical, thread-like (length = 15.21 mm, n = 1), test thin, smooth, featureless, translucent; head tip tapered bluntly rounded, containing a white gelatinous core, tapers to a thin flagellum and tip; head shorter thicker than flagellum (head length = 7.17 mm; 47 % of SPM length; flagellum length = 8.41 mm; head width = 142 μm; flagellum width = 31 μm). 4 SPM tightly coiled in brown gelatinous mass in BC of one specimen. Radula. Dentition formula 40: 1: 40 (Hubendick 1946: 51).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFEF8269FCCAFD02FD5FFBD6.taxon	diagnosis	Comparative remarks. In our molecular tree (Figs 1, 2), S. sirius (atra group, unit 31) is the sister species of an unidentified species from Tonga (Dayrat et al. 2014, unit 30). Both differ from each other by COI distances of ≥ 12.5 %. Siphonaria sirius differs from other species by COI distances of ≥ 22 % (Table S 9). Throughout its range, S. sirius has been found in sympatry with nine congeners. Two congeners are sympatric in Honshu: For comparison with S. japonica refer to comparative remarks under that species. Siphonaria acmaeoides has a paler, narrower ribbed shell with dual ribs forming the siphonal ridge, a weaker scalloped shell edge, a reduced AO, shorter wider ED and F 1, smaller BC, longer unlooped BD, and a smaller, drop-shaped SPM. Three congeners are sympatric in Cebu, Philippines: For comparisons with S. bifurcata and S. sipho refer to comparative remarks under these species. Siphonaria caubianensis sp. nov. has a dual-ribbed siphonal ridge, a more posterior and left offset apex, stronger scalloped shell edge, a larger AO, a smaller BC, and no distal loop. Two congeners are sympatric in Singapore: For comparison with S. viridis refer to comparative remarks under that species. Siphonaria alba has a dual-ribbed siphonal ridge with a less flared end, a weaker scalloped shell edge, whiter interior colouration, a larger AO and a shorter F 1. Siphonaria radians, sympatric on Riau Islands, S China Sea, has a weaker scalloped shell edge, a dual-ribbed siphonal ridge, a larger wider and more pointed AO, and a smaller BC. Siphonaria camura sp. nov., sympatric in Hong Kong, differs by having a smaller, taller shell with a dual-ribbed siphonal ridge, a shorter, wider ED, a larger, bulbous BC, and barbed SPM. Specimens from Japan figured in Hubendick (1946: pl. 3, figs 24 – 27), Kira (1962: 201, pl. 69, fig. 12 a, b), Habe (1971: pl. 4, fig. 12), Kuroda et al. (1971: pl. 64, fig. 9), Fukuda et al. (1992: pl. 23, fig. 361 a, b), and Dayrat et al. (2014: fig. 5 H) are morphologically consistent with S. sirius as delineated herein. By contrast, a specimen from Korea figured as ‘ S. sirius ’ in Yoo (1976: 89, pl. 19, fig. 5) is a misidentification. A figured specimen from Palawan identified as ‘ S. laciniosa ’ in Springsteen & Leobrera (1986: pl. 81, fig. 19) is S. sirius (single rib forming siphonal ridge); the associated ventral figure appears to be a different specimen (i. e., not matching the scalloped edge; with a multi-rib siphonal ridge; possibly a specimen of S. alba); the stated synonymy of S. atrata (sic atra) with S. laciniosa is incorrect. The figured specimen of S. sirius in Dharma (1992: pl. 17, fig. 2) from Indonesia is a misidentification and likely a specimen of S. alba based on shell features and distribution.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFEF8269FCCAFD02FD5FFBD6.taxon	distribution	Distribution and habitat. Known from Japan, China, Philippines, Vietnam, Singapore and Sumatra (Fig. 45). In this study found in sheltered positions on moderately exposed and exposed rocky shores, mid to upper littoral level, often associated with Lithothamium (Fig. 43 P).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFE8826AFF68FBE2FCF3FDD6.taxon	description	(Figs 43 J – L, S – T, 44 H – I)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFE8826AFF68FBE2FCF3FDD6.taxon	materials_examined	Material examined. Type material. Lectotype of Siphonaria subatra Pilsbry, 1904 from Chichijima, Ogasawara [Japan] (ANSP 86132 a, Fig. 43 J). Two paralectotypes same data as lectotype (ANSP 86132). Other, non-type material. Japan, Okinawa: Tancha Bay, 26 ° 27.897 ’ N, 127 ° 49.131 ’ E, JP 01 - 5 (AM C. 584932 1 p, C. 584932 p [SK 332]), Tancha Bay, rocky point 26 ° 27.941 ’ N, 127 ° 49.194 ’ E, JP 01 - 6 (AM C. 585663 6 p, C. 584930 p [SK 349], C. 584931 p [M 499, SK 318], C. 584933 p [M 498, SK 317]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFE8826AFF68FBE2FCF3FDD6.taxon	discussion	Taxonomic remarks. The original description is evidently based on a series of specimens (Pilsbry, 1904: 36, pl. 6, figs 61, 61 a – b). Baker (1964: 159) subsequently designated the lectotype. Dimensions of the lectotype (Fig. 43 J) match the original figure in Pilsbry (1904: 36) reasonably well. The lectotype has also been figured by Higo et al. (2001: 142, fig. G 4978 a). Our delineation of this species is based on comparative analyses of the morpho-anatomy and mitochondrial genetics of freshly collected topotypes (Fig. 43 K, L, Table S 1). Hubendick (1946: 50) listed a ‘ transitional form’of ‘ S. subatra <> S. atra ’ without mentioning the locality. However, the figured specimen (Hubendick 1946: 50, pl. 5, figs 5 – 6) is here identified as a specimen of S. subatra. Morrison (1972: 56 – 58) treated S. subatra as a synonym of S. laciniosa based on similarity in shell form and ‘ common reproductive development’. Christiaens (1980 b: 79) treated S. subatra as a variety of S. laciniosa based on specimens from Ping Chau, Hong Kong. These samples indeed correspond to shell characteristics considered herein as typical of S. subatra (i. e., siphon ridge formed by 3 – 4 ‘ coalescing’ small ribs). Je (1989: 29) incorrectly listed S. subatra as a synonym of Anthosiphonaria sirius (= S. sirius).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFE8826AFF68FBE2FCF3FDD6.taxon	description	External morphology (Fig. 43 S). Foot wall, cephalic folds and pneumostomal lobe all evenly pale grey in colour with white subepithelial pustules; paler grey to foot edge, foot sole darker; mantle thin, translucent, wider than foot wall, weakly lobed without a thickened edge, pale band on mantle edge align with underside of ribs; no black pigmentation; two black ‘ Eye’ spots prominent centrally on thickened cephalic lobes; pneumostome fold prominent. Shell (Figs 43 J – L; Table S 9). Medium sized (max sl mean = 19.8 mm, SD = 3.3 mm, n = 7), ovate; height low; apex offset sightly posterior and central, apical sides convex, shell thickness thick; protoconch direction undetermined, shell whorl dextral; growth striae indistinct in bands; exterior uneven, dark brown to pale tan, weak radial colour banding, protoconch area pale, central band darker and shell edge dark brown; rib count (mean = 51.4, SD = 8.0, n = 7), primary rib ridges pale, narrow, ribs slightly bent, increasingly raised and strongly protrude beyond shell lip (often> 1 mm) to unevenly scallop and corrugate the edge; 3 – 5 finer secondary ribs between primary ribs, rib interstices darker; siphonal ridge formed by closely paired primary ribs protrudes greatest. Interior shell margin dark chocolate brown to blue / grey, uneven blue / grey rays on shell margin aligned under primary ribs, siphonal groove prominent, often same colour as shell margin; spatula mottled dark chocolate to tan brown; ADM scar distinct, CMS convex, darker than shell lip; thickening of shell lip common, overcoats brown markings on shell margin with blue / grey, infills and reduces lip scalloping. Reproductive system (Fig. 44 H; n = 3). Positioned within coelom under the respiratory cavity, hermaphroditic glands positioned to posterior against right foot wall and over foot sole, epiphallic parts positioned between BM and RAM; AO prominent, large, reasonably long, broad bluntly pointed, joins at top of smallish GA; ED elongated, narrow, twisted; EG medium with folds, single bent thickish flagellum F 1 appears as an extension of ED at join with EG; AO, GA and ED all muscular white tissue; BD and CD elongated, with opposing connections join into GA close to ED and AO; BD longer and narrower than CD with a prominent distal coiling with an MA on bend attaching to inner foot wall, CD broadens to MG end, BD smooth, both ducts pass together through outer side of RAM (BD above CD) and bent before connecting into folds of MG; BC small, prominent with internal purple gel, embedded in MG, translucent test, medium and bulbous; coiled HD with brown markings links AG at lower end of a block-like elongated yellowish finely granulated HG; MG and AG soft white folded tissue; SV embedded within AG, AG larger than HG, sides match curvature of inner foot wall at right posterior quarter of coelom. Spermatophore (Fig. 44 I). Body cylindrical, thread-like, test thin, translucent, smooth, featureless (length = 9.17 mm, head length = 7.85 mm, n = 1, 86 % of SMP length, AL = 16 mm; head tip tapered bluntly rounded, section containing a white core, tapers to a thin short flagellum; two SPM tightly folded in brown gelatinous mass in BC of one specimen. Radula. Dentition formula 34: 1: 34 (Hubendick 1946: 51).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFE8826AFF68FBE2FCF3FDD6.taxon	diagnosis	Comparative remarks. In our phylogeny (Figs 1, 2), S. subatra (atra group, unit 38) is the sister species of S. tenebrae sp. nov. (unit 92) from CI. Both species combined from the sister lineage of S. vudaensis sp. nov. (unit 37) from Fiji. Siphonaria subatra differs from S. tenebrae by COI distances of ≥ 15.1 % and from S. vudaensis by distances of ≥ 13.3 % (Table S 3). Siphonaria subatra has been found in sympatry with three congeners in Okinawa. Siphonaria rucuana has a smaller, taller, shell with stronger ribbing and weaker edge scalloping, a larger, bulbous, blunt AO, and a longer BC. Siphonaria camura sp. nov. has a smaller, taller, more fragile shell with a less scalloped edge, larger, bulbous BC, BD without a distal loop, and a barbed SPM. Siphonaria tanchaensis sp. nov. has a larger, taller, paler shell with greater edge scalloping, patterned exterior, and a smaller AO and BC. For comparison with S. sipho refer to comparative remarks under that species. Siphonaria subatra exhibits a similar shell morphology to other species in the atra group. However, the extended projection of the multi rib formed siphonal ridge beyond shell edge is a major difference. A shell figured as ‘ S. atra ’ from Japan in Hirase (1941: pl. 121, fig. 17) corresponds well with characteristics typical of S. subatra. It is well outside the known distribution of this species. Specimens depicted as ‘ S. subatra ’ in Hubendick (1946: pl. 3, figs 32 – 35) are of two different species; figs 32, 35 from ‘ Japan’ are specimens of S. subatra and figs 33, 34 from Mindanao and Java Sea are likely specimens of S. alba. Figured specimens in Kira (1962: pl. 69, figs 10 a, b) from Amami Islands as well as from Okinawa (atra group, unit 38) in Dayrat et al. (2014: fig. 5 R, UF 351784) exhibit features typical of S. subatra.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFE8826AFF68FBE2FCF3FDD6.taxon	distribution	Distribution and habitat. Recorded from Okinawa, Japan (Fig. 45). In this study, found on exposed rocky shores at sheltered positions across the mid-littoral level (43 T).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFEB8264FCCAFDE2FC39FDB6.taxon	description	(Figs 43 G – I, Q – R, 44 F – G)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFEB8264FCCAFDE2FC39FDB6.taxon	materials_examined	Material examined. Type material. Lectotype of Siphonaria rucuana Pilsbry, 1904 from ‘ Riukiu Island’ [Japan] (ANSP 86131 a, Fig. 43 G). Three paralectotypes, same data as lectotype (ANSP 86131). Other, non-type material. Japan, Okinawa: Tancha Bay, 26 ° 27.897 ’ N, 127 ° 49.131 ’ E, JP 01 - 5 (AM C. 585662 4 p, C. 584912 p [SK 409, protoconch H 9], C. 584915 p [M 493, SK 312]; C. 584916 p [SK 355]; C. 584917 p [SK 377], C. 584919 p [SK 345], C. 585082 p [SK 406], C. 585914 p [SK 354], C. 585915 p [SK 383], rocky point, 26 ° 27.941 ’ N, 127 ° 49.194 ’ E, JP 01 - 6 (AM C. 585627 6 p, C. 585917 p [M 397], C. 584918 p [M 492, SK 311]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFEB8264FCCAFDE2FC39FDB6.taxon	discussion	Taxonomic remarks. The original description of Pilsbry (1904: 36, pl. 6, figs 60 a – b) is based on a series of specimens. Baker (1964: 159) subsequently designated the lectotype. The dimensions of the lectotype (Fig. 43 G) match the original dimensions in Pilsbry (1904: 36) reasonably well. The lectotype has also been figured by Higo et al. (2001: 142, fig. G 4973). Our delineation of this species is based on comparative analyses of the morpho-anatomy and mitochondrial genetics of freshly collected topotypes (Fig. 43 H – I, Table S 1). ‘ Siphonaria zebra (?) ’ figured in Kuroda (1941: 137, pl. 3, figs 40, 50) is a specimen of S. rucuana. The ‘ uncertain’ records of ‘ S. rucuana’ in Hubendick (1955: 7) are likely misidentified specimens of S. denticulata (from Etty Bay, Qld, MV F 15040) and S. viridis (from Cape Edgecumbe [sic], Bowen, Qld, MV F 15041). Habe (1962: 96, pl. 44, fig. 16) treated S. rucuana as an accepted species; however, Habe (1964: 144, pl. 44, fig. 16) treated it as a subspecies of S. laciniosa. This later treatment is not accepted.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFEB8264FCCAFDE2FC39FDB6.taxon	description	External morphology. Foot sole pale brown, paler to foot edge; foot wall, mantle, cephalic folds and pneumostomal lobe all evenly dark yellowish; mantle thin, translucent, weakly lobed with a thickened dark yellowish banded edge; faint irregular black blotches of pigmentation on foot wall and cephalic lobes; pneumostome fold long between right ADMs and within mantle; genital pore inconspicuous, located on foot wall to right anterior of right cephalic fold; two small black epithelial eye spots centralised on two centrally touching unpigmented cephalic folds. Shell (Figs 43 G – I, R; Table S 9). Small sized (max sl me = 9.2 mm SD = 0.7 mm, n = 10); ovate to elongate, height medium; apex offset central to weakly posterior; protoconch direction weakly heterostrophic (n = 2; Fig. 43 R), shell whorl dextral; apical sides convex, posterior weakly concave to straight; radial colour banding, protoconch area dark brown, centre pale grey, darker to uneven shell edge; growth lines distinct; rib count (mean = 28, SD = 3.1, n = 10) primary ribs whitish grey, straight, rib interstices dark brown / black; rib ridges rounded, broader to and weakly extend beyond shell lip, paired primary ribs form siphonal ridge, 0 – 1 finer grey secondary ribs between primary ribs, number greater either side of siphonal ridge. Interior shell lip weakly corrugated, with white rays aligning under primary ribs, and narrower dark brown / black markings under rib interstices; ADM scar prominent, CMS straight to weakly convex; shell margin and siphonal groove evenly dark chocolate brown, spatula white / grey; thickening or whitening of inner shell lip and spatula not observed. Reproductive system (Fig. 44 F; n = 2). Positioned within coelom under the respiratory cavity, occupies the entire right side of coelom, hermaphroditic glands positioned to posterior against right foot wall and over foot sole, epiphallic parts positioned over back of BM; GA prominent with singular GP through foot wall; AO medium, broad, blunt, joined to upper GA alongside ED; ED elongated, broad, slightly bent (no MA), joins to side of GA; GA, AO, ED all white muscular fibrous tissue; EG soft whitish tissue, slightly folded, joins ED; single broad flagellum (F 1) with possible 2 nd shorter flagellum; appears as a continuous extension of ED to EG, laid over BM; BD and CD connect together in opposing directions into GA between ED / AO joint and GP, both ducts short, straight, smooth, thickened, whitish, featureless, pass closely together through outer side of RAM (BD over CD) into soft white folded tissues of MG; MG / AG complex medium; BD narrower than CD, straight looped, end of loop attached to inner foot wall; BC medium, spherical, thin translucent test, embedded in outer folds of AG / MG; HD short, thickened, coiled, links ducts in soft white folded tissues of AG to yellowish granulated HG; outer edge of MG lobbed; AG larger than HG, outer sides of both matches curvature of inner foot wall. Spermatophore (Fig. 44 G). Thread-like (length = 6.88 mm, n = 1), translucent, test thin; head section bluntly rounded, cylindrical, containing a core white gelatinous mass, tapers along the transparent flagellum to a thin tip; both sections smooth, featureless. Head section shorter wider than flagellum (head length = 2.74 mm, ~ 40 % of SPM length; head width = 90 μm, flagellum width = 17 μm); 1 SPM in one BC (AM C. 584917).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFEB8264FCCAFDE2FC39FDB6.taxon	diagnosis	Comparative remarks. Siphonaria rucuana (normalis group, unit 83) differs from other species by COI distances of ≥ 24 % (Table S 6, Figs 1, 4). The species has been found in sympatry with four congeners on Okinawa. Siphonaria camura sp. nov. has a smaller, taller, paler, fragile shell with a more offset apex and prominent ribbing, a smaller BC and AO, and a barbed SPM. Siphonaria tanchaensis sp. nov. has a larger, paler shell, with a patterned exterior and greater edge scalloping, and a smaller BC and AO. For comparison with S. sipho and S. subatra refer to comparative remarks under these species. Specimens figured as ‘ Siphonaria radians’ in Habe & Kosuge (1966: pl. 42, figs 24, 25) are attributed herein to S. rucuana. Siphonaria radians has less raised, finer ribbing, and a less prominent siphonal ridge.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFEB8264FCCAFDE2FC39FDB6.taxon	distribution	Distribution and habitat. Known only from Okinawa, Japan (Fig. 45). In this study, found on exposed rocky shores in sheltered positions (i. e., rock crevices and hollows; Fig. 43 Q), mid to upper littoral level (amongst green algae).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFE58266FCCAFB42FBEBF7D6.taxon	description	(Figs 46 A – C, M – O, 47 A – B)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFE58266FCCAFB42FBEBF7D6.taxon	materials_examined	Material examined. Type material. Lectotype of Siphonaria stowae (SAM D. 33484; figured in Verco 1906: figs 3 – 5; Jenkins, 2018: fig. 2 A – C). Eight paralectotypes of Siphonaria stowae from Pandolowie [sic Pondalowie] Bay, Spencers Gulf], SA (SAM D. 13590, lot labelled as ‘ holotype’, largest figured in Jenkins 2018: fig. 2 D – F). Three syntypes of Pugillaria stowae comita from Twofold Bay, NSW, [Australia] coll. R. Bell [undated], T. Iredale coll. (AM C. 265927, two figured in Jenkins 2018: fig. 2 J – O). Other, non-type material. Australia, NSW: Woody Head, near Iluka, 29 ° 22.0 ′ S, 153 ° 22.50 ′ E (AM C. 116737 6 d); Clarence River, 29 ° 25.50 ′ S, 153 ° 21.00 ′ E (AM C. 398326 2 d), 29 ° 25 ′ S, 153 ° 21 ′ E (AM C. 265949 2 d); Woolgoolga, 30 ° 06.70 ′ S, 153 ° 12.300 ′ E (AM C. 398328 d); Port Stephens, between beacons, 32 ° 42.397 ′ S, 152 ° 11.502 ′ E (AM C. 265967 d); North Fingal Bay, 32 ° 44.750 ′ S, 152 ° 10.500 ′ E (AM C. 398325 6 d); Fingal Bay, 32 ° 45.0 ′ S, 152 ° 10.500 ′ E (AM C. 398324 d); Patonga, BrokenBay, 33 ° 33.111 ′ S, 151 ° 16.570 ′ E (AMC. 2659614 d); Collaroy Beach, N of Sydney, 33 ° 44.0 ′ S, 151 ° 18.0 ′ E (AM C. 398336 d), 33 ° 43.700 ′ S, 151 ° 18.0 ′ E (AM C. 398335 d); Manly Beach, 33 ° 47.817 ′ S, 151 ° 17.368 ′ E (AM C. 265964 d); Ocean Beach, Manly, 33 ° 47.853 ′ S, 151 ° 17.398 ′ E (AM C. 265970 3 d); Middle Harbour, between Grotto and Dobroyd Points, 33 ° 48.897 ′ S, 151 ° 16.085 ′ E (AM C. 265966 d); Balmoral Beach, 33 ° 49.700 ′ S, 151 ° 15.030 ′ E (AM C. 030179 2 d); Off Chinamans Beach, 33 ° 48.870 ′ S, 151 ° 14.965 ′ E (AM C. 265976 2 d); Sydney Harbour, Quarantine Bay, 33 ° 50.863 ′ S, 151 ° 14.438 ′ E (AM C. 265973 d); Bradleys Head, 33 ° 51.300 ′ S, 151 ° 14.700 ′ E (AM C. 398327 2 d); Sydney, Little Coogee Bay, 33 ° 55.300 ′ S, 151 ° 15.600 ′ E (AM C. 398322 d); Botany Bay, Kurnell, 34 ° 0.580 ′ S, 151 ° 12.380 ′ E (AM C. 398332 2 d); Port Hacking, S of Sydney, Cronulla, Gunnamatta Bay, 34 ° 3.950 ′ S, 151 ° 8.550 ′ E (AM C. 398330 d); SW end Gunnamatta Bay, 34 ° 4.300 ′ S, 151 ° 8.700 ′ E (AM C. 398334 d); Sussex Haven and Wreck Bay, 35 ° 10.203 ′ S, 150 ° 41.293 ′ E (AM C. 22532 d; AM C. 265932 d); Off Montague Island, Narooma, 36 ° 14.347 ′ S, 150 ° 13.015 ′ E (AM C. 265974 4 d); Twofold Bay, Murrumbulga Point, 37 ° 04.702 ′ S, 149 ° 53.103 ′ E (AM C. 150582 p). Vic: 1.6 km N of Gabo Island, 37 ° 34.0 ′ S, 149 ° 56.0 ′ E (AM C. 398323, 1 d); Mallacoota, 37 ° 34.0 ′ S, 149 ° 46.600 ′ E (AM C. 50394 2 d); Bear Gully, Waratah Bay, 38 ° 20 ′ S, 146 ° 00 ′ E (MV F 169206 1 p); Inverloch, 38 ° 38 ′ S, 145 ° 43 ′ E (MV F 161257 2 p); San Remo, Western Port, 38 ° 32.0 ′ S, 145 ° 23.0 ′ E, V 07 - 1 (AM C. 030683 2 d, C. 585616 p [SK 393]), 38 ° 31.489 ’ S, 145 ° 21.858 ’ E (AM C. 585583 3 p); Flinders, Western Port Bay, 38 ° 29.0 ′ S, 145 ° 1.0 ′ E (AM C. 398366 d, AM C. 265936 d); ‘ Clondrisse’ E of Cape Schanck, 38 ° 29.583 ′ S, 144 ° 53.654 ′ E (MV F 193073 p); Port Phillip Bay, 38 ° 09 ′ S, 144 ° 46 ′ E (MV F 185072 2 p); Cheviot Beach, Point Nepean, 38 ° 18 ′ S, 144 ° 40 ′ E (F. 87801 p); Andersons Point, Portland, 38 ° 19.967 ′ S, 141 ° 36.604 ′ E (MV F 126944 p). Tas: King Island: Gulchway, S of Surprise Bay, 40 ° 08.023 ′ S, 143 ° 54.205 ′ E (MV F 193075 p); Fraser Beach, Sea Elephant Bay, 39 ° 54.410 ′ S, 144 ° 06.584 ′ E (AM C. 265930 d); West Head, Greens Beach, Tamar River mouth, 41 ° 5.0 ′ S, 146 ° 45.0 ′ E (AM C. 398367, 1 d); Park Beach Dodges Ferry 42 ° 51.716 ’ S, 147 ° 36.665 ’ E (AM C. 585469 11 p, C. 584835 p [M 109, SK 018], C. 584914 p [SK 216], C. 585264 p [SK 007], C. 585265 p [M 110]). SA: Guichen Bay, near Cape Dombey, 37 ° 7.233 ′ S, 139 ° 45.967 ′ E (SAM D. 33486 3 d); Stokes Bay, N coast of Kangaroo Island, 35 ° 37 ′ S, 137 ° 12 ′ E (AM C. 265968 2 d); Normanville, S of Adelaide, 35 ° 26.800 ′ S, 138 ° 18.500 ′ E (AM C. 398369 d); Adelaide, Glenelg Beach, S of Adelaide, 34 ° 58.0 ′ S, 138 ° 32.0 ′ E (AM C. 265963 d, AM C. 398370 d); Henley, 34 ° 55.430 ′ S, 138 ° 29.595 ′ E (AM C. 265962 2 d); Grange, 34 ° 58.215 ′ S, 138 ° 30.471 ′ E (SAM D. 33490 4 d); Spencer Gulf, Eyre Peninsula, Tumby Bay, 34 ° 22.0 ′ S, 136 ° 8.0 ′ E (AM C. 398372,1 d); Arno Bay, 33 ° 56.0 ′ S, 136 ° 35.0 ′ E (AM C. 398368 10 + d); Pondalowie Bay, 35 ° 13.989 ’ S, 136 ° 49.892 ’ E (AM C. 585492 p); Near Salmon Point, Elliston Bay, 33 ° 39.0 ′ S, 134 ° 53.0 ′ E (AM C. 265905 1 d); Sceale (sic Sceales) Bay, Cape Blanche, 33 ° 00.237 ′ S, 134 ° 11.502 ′ E (SAM D. 33485 11 d); Franklin Islands, Investigator Group, 32 ° 26.450 ′ S, 133 ° 39.735 ′ E (SAM D. 33489 3 d); Point Sinclair, 32 ° 6.0 ′ S, 132 ° 59.0 ′ E (AM C. 398371 d). WA: Gnarabup Beach, S of Margaret River, 34 ° 1.0 ′ S, 114 ° 59.0 ′ E (AM C. 398365 d); Kilcarnup, N side Margaret River, 33 ° 57.0 ′ S, 114 ° 59.0 ′ E (AM C. 398360 3 d); Ellensbrook (S Cowaramup), near Margaret River mouth, 33 ° 53.0 ′ S, 114 ° 59.0 ′ E (AM C. 398361 d, AM C. 265965 9 d, AM C. 398363 d, AM C. 398364 9 d); N side Cape Naturaliste Lighthouse, 33 ° 32.752 ′ S, 115 ° 0.418 ′ E (AM C. 265975 d); Bunbury, 33 ° 18.750 ′ S, 115 ° 39.061 ′ E (SAM D. 33488 2 d); Point Peron, 48 km S of Perth, 32 ° 16.0 ′ S, 115 ° 41.0 ′ E (AM C. 398358 d, AM C. 398357 3 d); Garden Island, S of Perth, 32 ° 14.0 ′ S, 115 ° 41.0 ′ E (AM C. 398359 11 d); Cockburn Sound, Jervoise Groyne, 1.6 km S of Woodmans Point, 32 ° 9.0 ′ S, 115 ° 46.0 ′ E (AM C. 398362 d); Rottnest Island, 32 ° 00.683 ′ S, 115 ° 30.993 ′ E (SAM D. 33492 7 d); Geraldton, 28 ° 46.143 ′ S, 114 ° 36.283 ′ E (SAM D. 33491 d).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFE58266FCCAFB42FBEBF7D6.taxon	discussion	Taxonomic remarks. The lectotype has been designated by Jenkins (2018: 3). Examinations of freshy collected topotypic specimens herein (Fig 46 A – C) validate the identity of S. stowae and confirm that S. stowae comita is its junior synonym. Hedley (1916 a: 220) transferred S. stowae to the genus Kerguelenia. Refer to Jenkins (2018: 276) for comments on type specimens, including labels. The present description is based on the re-description of this species in Jenkins (2018: 276) and is expanded upon for completeness and taxonomic consistency.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFE58266FCCAFB42FBEBF7D6.taxon	description	External morphology (Fig. 46 O). Foot wall, mantle, pneumostomal lobe and cephalic folds pale grey to cream, foot sole darker grey paling to foot edge; mantle narrow, thin, translucent with thickened unlobed fringe, even pigmented shading only at foot wall mantle join; pneumostomal lobe on right side within mantle, covering inconspicuous anus in foot wall; two black ‘ Eye’ spots prominent centrally on touching cephalic folds; genital pore inconspicuous, positioned in foot wall posterior to right cephalic fold. Shell (Figs 46 A – C, M; Table S 9). Small sized (max sl <10 mm), ovate, cap-like, arched dorsally, height medium to tall; apex lower than shell height, dorsally aligned close to or over posterior edge (Jenkins 2018: fig. 2), laterally offset 10 – 15 ° to left of transverse centre line (cl); exterior white to cream with irregular red-brown apical striations and blotches, coloration and patterning variable; Shell whorl dextral, protoconch direction homostrophic (n = 1; Jenkins 2018: fig. 2 V); posterior margin weakly concave, other margins convex (Jenkins 2018: fig. 2); in larger specimens (sl> 8 mm), lip thickened, opaque, ribs prominent, irregularly spaced, flattened, weakly corrugated, often distorted by prominent growth striae (Jenkins 2018: fig. 2 F, M); rib interstices are irregular and red-brownish denoted by coloured apically aligned bands or streaks (Jenkins 2018: fig. 2 A – C); growth striae prominent and uneven; in small specimens, shell translucent, exterior smooth and polished, ribs indistinct, lip thin, unscalloped and translucent to edge, red-brown exterior of irregular rib interstices shows through to interior between ADM scar and lip (Jenkins 2018: fig. 2 H, Q); interior smooth, cream coloured, spatula smooth and white; ADM scar indistinct, weakly indented, posterior of scar shaded red-brown (Jenkins 2018: fig. 2), cephalic ADM scar (cam) indistinct, straight to slightly convex; siphonal groove weakly indented. Reproductivesystem (Fig. 47 A; n = 3). Predominantly located in posterior of coelom; HG granulated, HD and folds of AG situated to posterior under digestive gland and pallial cavity; a small rounded brownish BC and BD located behind pneumostomal opening and beside posterior RAM; BD narrow elongate; HG connected by several short thin translucent ducts; MG and AG with translucent folds; SV positioned to side of these folds; EG, ED and small bulbous GA are situated to right side of BM just behind RAM; two short thick flagellum present at join of ED and EG; outer layer of GA translucent whitish with white opaque central layer; BD passes through the RAM; a thickened coiled CD decreasing in diameter opens into GA; a single (monoaulic) small GP opens from GA through foot wall posterior to right cephalic fold and in front of right anterior RAM; BD and CD open separately into GA close to GP. Spermatophore (Fig. 47 B). Thread-like (length = 3.63 ± 1.06 mm, n = 2), translucent, test thin; head section, tip bluntly rounded, evenly cylindrical, elongate, containing a white gelatinous mass; taper region into the filamentous transparent flagellum is short; both sections smooth, featureless. Head shorter and thicker than flagellum (head length = 1.46 ± 0.11 mm, ~ 41 % of SMP length, head width = 120 ± 20 μm; flagellum width = 13 ± 15 μm). We found 7 SPM tightly coiled each in two bursas. Radula (Jenkins 2018: fig. 4 A – D). each transverse row has a single narrow central rachidian tooth flanked squarely by mirrored half rows of block-like lateral teeth (fig. 4 A); tooth size in a single row decreases rapidly from the more solid (around 4 × larger) innermost laterals to smaller outermost mid laterals, the gap between rows notably increases (fig. 4 B, D); each half row has on average eight mid and eight outer lateral teeth; inner teeth interlock via forks and notches, centrals anterior dualpronged fork, blunt posterior notch; laterals possess a single outward pointing basal prong and notch (fig. 4 B); central tooth significantly lower and smaller than adjacent lateral teeth with single weakly-pointed mesocone, basal plate of similar length, narrower than adjacent laterals (fig. 4 A); inner laterals (i. e., without ecto / endocones) uncommon, irregular (fig. 4 A); mid lateral mesocones around half length of base, edge shallow ‘ U’ shaped, bluntly bi-cuspidate (some weakly tricuspidate), tips of inner lateral mesocones often erode back of tooth in front; single ectocone increasing from almost no ectocone to very prominent ectocone almost width and half height of mesocone (figure 4 B); outer lateral (fig. 4 B) mesocone single cusped, ecto / endocones variably-shaped bluntlypointed tooth bases progressively widen to outer ribbon edge; outside outer laterals (from tooth 13) possess additional endocone, ecto / endocones appear as separate cones, some almost as large as mesocones size, without an angle of separation from mesocone. Radula dentition agrees with original description (Verco 1906: 224) apart from having four fewer laterals per half row than Verco’s formulae (‘ around 22 ’); and for the lack of evidence that the central tooth ‘ tends to be bilobed’ (i. e., bi-cuspidate). Dentition formula 16: 1: 16 + / − 2 (n = 3; row count not assessed), 22: 1: 22 (Hubendick 1946: 29).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFE58266FCCAFB42FBEBF7D6.taxon	diagnosis	Comparative remarks. Siphonaria stowae (atra group, unit 52) is member of a subclade of Clade H containing the species S. zelandica (unit 26), S. acmaeoides (unit 91), and S. restis (unit 54) (Figs 1, 2). It differs from these species by COI distances of ≥ 17.5 % (Table S 4). Throughout the range of S. stowae we found ten congeners with partly sympatric distributions. Four species are sympatric in south-eastern Australia. Siphonaria emergens has a smaller, mottled orange / brown shell with less edge scalloping, S. pravitas sp. nov. has a larger, lower shell with stronger ribbing and edge scalloping, a more prominent siphonal ridge, a larger AO and BC, and a longer ED. Siphonaria scabra has a larger, taller shell with greater edge scalloping, a larger AO, longer ED, and a longer SPM. For comparison with S. denticulata refer to comparative remarks under that species. Five species have been found in sympatry in southern Australia: For comparisons with S. diemenensis, S. funiculata, S. jeanae, S. tasmanica, and S. zelandica refer to comparative remarks under these species. One species occurs in sympatry in south-western Australia: Siphonaria restis sp. nov. has a larger, lower shell with stronger raised ribbing and greater edge scalloping, a larger, bulbous, blunt AO, and a longer BC. The combined shell geometry, size and colouration of S. stowae is very distinctive and unlike any other siphonariid.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFE58266FCCAFB42FBEBF7D6.taxon	distribution	Distribution and habitat. Southern coasts of Australia, from Iluka, northern NSW, through Vic, Tas, SA to Geraldton, WA (Fig. 48). Found in sheltered rocky intertidal platforms and rocky areas, often on rocks in tidal pools and associated with white Lithothamnion algae, at mid littoral level (Fig. 46 N).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFE18260FF68FF02FCEAF7F6.taxon	description	(Figs 46 G – I, 47 H – I)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFE18260FF68FF02FCEAF7F6.taxon	materials_examined	Material examined. Type material. Lectotype of Siphonaria madagascariensis Odhner, 1919, present designation, from Majunga [Mahajunga, Madagascar]; coll. M. W. Kaudern, 1912 (UUZM UUMS 2000). Two paralectotypes, same data as lectotype (UUMS; not seen). Other, non-type material. Madagascar: Ambatobe, Bavarama, 25 ° 27.9 ’ S, 44 ° 57.6 ’ E, BM 06, (MNHN IM- 2009 - 14095 p [M 582]); Plage de Lavanono, 25 ° 25.2 ’ S, 44 ° 56.3 ’ E, BM 01 (MNHNIM- 2009 - 13779 p [M 577]); SW coast, Itampolo, Fringing reef, 24 ° 50.885 ’ S, 43 ° 59.693 ’ E MA 09 - 1 a (AM C. 585971 10 p, AM C. 584818 d [M 267, SK 372]; C. 584819 p [M 268, SK 370], C. 584820 p [M 269, SK 371]); Inner lagoon shore, 24 ° 50.885 ’ S, 43 ° 59.693 ’ E MA 09 - 1 b (AM C. 608184 7 p, C. 584957 p [M 270], C. 584958 [M 271], C. 584959 p [M 272]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFE18260FF68FF02FCEAF7F6.taxon	discussion	Taxonomic remarks. The largest syntype is herein designated as the lectotype of S. madagascariensis for the stabilisation of the name (Art. 74.1 of the Code). Our delineation of this species is based on comparative analyses of the morpho-anatomy and mitochondrial genetics of freshly collected topotypes (Fig. 46 G – I) and geographic series of additional specimens (Table S 1). Morrison (1972: 56 – 58) treated S. madagascariensis as a junior synonym of S. laciniosa based on similarity in shell form and ‘ common reproductive development’. This synonymy is not supported by examination of type specimens and comparative morpho-anatomy.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFE18260FF68FF02FCEAF7F6.taxon	description	External morphology. Foot wall, mantle, cephalic folds and pneumostomal lobe all evenly pale cream in colour paler to foot edge, darker to foot sole; faint irregular dark brown pigmentation markings over foot wall and centre of cephalic folds; mantle narrower than foot wall, lobed with a thickened cream edge band; mantle lobes align with undulations of primary shell ribs, foot wall and pneumostome pustulose, pneumostome wide between right ADMs and within mantle. Shell (Figs 46 G – I; Table S 9). medium sized (max sl mean = 19.3 mm, SD = 2.1 mm, n = 6), ovate, flattened, shell height low, apex offset strongly to left and posterior of centre, apical sides convex, growth lines prominent, surface uneven; protoconch direction heterostrophic (n = 1), shell whorl dextral; growth striae distinct; ribs raised, whitish, wavy rib count (mean = 42.7, SD = 1.6, n = 6), ridges rounded, width increases strongly to shell lip, rib interstices narrow, dark brown; 11 – 14 prominent primary ribs, extend up to 1 mm beyond shell edge; siphonal ridge prominent, formed by dual primary ribs, 1 – 2 interspersed secondary ribs; shell lip uneven, weakly scalloped and unevenly corrugated aligning with protruding ribs. Interior colouration matches white primary ribs and dark brown rib interstices, from shell lip, over shell margin to the dark chocolate brown coloured spatula; ADM scar impression distinct, same as shell margin colouring, siphonal groove prominent, indented; CMS straight to convex; thickening / faint whitening of shell interior occurs; white layering thickens and covers shell margin, spatula coated white (e. g., Fig. 46 G). Reproductive system (Fig. 47 H; n = 2). Positioned within coelom under the respiratory cavity, hermaphroditic glands positioned to posterior against right foot wall and over foot sole, epiphallic parts positioned to anterior over back of BM. Join of AO and ED to top of GA distinct, AO larger than GA, elongated, broad, bluntly pointed, slightly bent, much thicker than ED; ED longer and narrower than AO; AO, GA and ED all muscular white tissue; EG large, bluntly pointed with a single long broad bent flagellum F 1; BD and CD connect in opposing directions into GA, CD at inner side and BD in front of ED entry; BD with bulbous entry and weak distal loop, in front of GA / ED join, BD and CD similar in thickness, BD longer, both ducts smooth and pass together through RAM connecting into MG (BD over CD), BC large, bulbous, embedded in MG / AG and against digestive tract; HD long broad coiled, links under digestive tract against foot sole, a small broad AG to a separated broad yellowish granulated HG, MG small, AG and MG folded, soft white tissue, AG slightly smaller than curved HG reflecting the close positioning to curvature of inner foot wall at right posterior quarter of coelom; SV embedded on left side of AG close to BC. Spermatophore (Fig. 47 I). Thread-like, test thin, translucent (length = 10.96 ± 4.5 mm, n = 4, mean AL = 9.4 mm); head section cylindrical, bulbous, centrally bent, rounded tip; test thin, smooth, featureless, translucent, contains a white core, tapers into short flagellum; head slightly shorter, wider than translucent flagellum (head length = 8.5 ± 1.7 mm, ~ 82 % of SPM length, head width = 155 ± 20 μm, flagellum width = 47 ± 10 μm, n = 4); 8 SPMs tightly coiled in BC of one specimen. Radula. Dentition formula 30: 1: 30 (Hubendick 1946: 56).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFE18260FF68FF02FCEAF7F6.taxon	diagnosis	Comparativeremarks. Siphonariamadagascariensis (atra group, unit 44) differs from its sister species S. belcheri (Figs 1, 2) by COI distances of ≥ 21.4 % (Table S 5). Siphonaria madagascariensis occurs in sympatry with two other congeners in Madagascar: Siphonaria striata sp. nov. has a smaller, taller, paler shell with a more posteriorly offset apex, less raised ribbing and darker interior, distal and bursal BD loops, a smaller BC and a wider ED. Siphonaria itampoloensis sp. nov. has a smaller, taller, paler shell with less raised ribbing, a smaller AO and BC, and a shorter ED. Based on comparatively ‘ greater number of [shell] ribs’ and ‘ stouter genital retractor’, Hubendick (1946: 56) doubted that S. madagascariensis should be ‘ specifically distinguished’ from S. kurracheensis. However, apart from being genetically clearly different, S. madagascariensis has consistently broader and larger RS epiphallic parts (including genital retractor) than S. kurracheensis.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFE18260FF68FF02FCEAF7F6.taxon	distribution	Distribution and habitat. Endemic to Madagascar (Fig. 48). Found in sheltered positions on moderately exposed fringing reef and inner lagoon shores, mid littoral level.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFE08262FF68FF02FA8BF7F6.taxon	description	(Figs 46 J – L, Q, 47 J – K)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFE08262FF68FF02FA8BF7F6.taxon	materials_examined	Material examined Type material. Lectotype of Mestosiphon lentulus Iredale, 1940, present designation, from LHI; coll. R. Bell, 1912 – 14 (AM C. 103712, Fig. 46 J). Forty-one paralectotypes, same data as lectotype (AM C. 103713). Other, non-type material. Australia, LHI: Off NE side; 31 ° 30.43 ’ S, 159 ° 04.22 ’ E (AM C. 356975 d); 31 ° 32.5 ’ S, 159 ° 03.75 ’ E (AM C. 103712 d; AM C. 103713 41 d); near runway 31 ° 32.176 ’ S, 159 ° 04.258 ’ E, LHI 2017 Apr 04 - 102 (AM C. 482039 p); lagoon near wharf 31 ° 31.5 ’ S, 159 ° 03.45 ’ E (AM C. 356974 d); Signal Point, 31 ° 31.5 ’ S, 159 ° 03.88 ’ E (AM C. 356977 d); 31 ° 31.501 ’ S, 159 ° 03.580 ’ E, LHI 2017 Apr 04 - 104 (AM C. 585957 3 p); 31 ° 31.501 ’ S, 159 ° 03.578 ’ E, LHI 2017 Apr 04 - 099 (AM C. 546717 5 p; C. 546718 14 p, C. 585955 p [SK 053], C. 585956 p [SK 054], C. 585958 p [SK 234], C. 595975 p [M 040]); between Old Settlement Beach and Dawson Point, 31 ° 31.18 ’ S, 159 ° 03.45 ’ E (AM C. 356978 6 p, C. 608189 p [SK 051]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFE08262FF68FF02FA8BF7F6.taxon	discussion	Taxonomic remarks. The description contains no original type designation (Iredale, 1940: 439). There are two registered type lots in AM reference collection, one labelled ‘ holotype’ (AM C. 103712) and another with same data as the ‘ holotype’ (AM C. 103713, 41 d, paralectotypes). We consider all these specimens as syntypes. The specimen AM C. 103712 is herein designated as the lectotype of M. lentulus for the stabilisation of the name (Art. 74.1 of the Code). Our delineation of this species is based on comparative analyses of the morpho-anatomy and mitochondrial genetics of freshly collected topotypes (Figs 46 K – L; Table S 1). The lectotype is the specimen figured in Iredale (1940: 441, pl. 34, fig. 14 – 15) matching the shell dimensions given in the original description. Hubendick (1946: 49) incorrectly treated S. lentula as a synonym of ‘ S. australis’ without examining types or topotypes. However, S. australis is a distinct species (Jenkins 1983: 1, fig. 3 a). Morrison (1972: 56 – 58) treated ‘ M. lentulus’ as a synonym of S. laciniosa based on similarity in shell form and ‘ common reproductive development’. This synonymy is not supported by examination of type specimens and comparative morpho-anatomy.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFE08262FF68FF02FA8BF7F6.taxon	description	External morphology. Foot sole, foot wall, cephalic folds, mantle evenly dark grey / brown; foot edge; mantle narrow, mantle edge unlobed thickened paler band, bands dark pigmentation aligning with rib interstice furrows; genital pore inconspicuous, located on foot wall to right anterior of right cephalic fold; two small black epithelial eye spots centralised on two thick centrally touching dark grey cephalic folds; thin, pale grey, large pneumostomal lobe part of the mantle, between the right ADMs, closes the pneumostome and anus at the mantle edge. Shell (Figs 46 J – L, Q; Table S 9). Small sized (max sl mean = 9.9 mm, SD = 1.0 mm, n = 13); ovate often elongate; height medium; flattened, apex offset central slightly to posterior; apical sides anteriorly concave posteriorly convex; protoconch direction homostrophic (n = 3; Fig. 46 Q), shell whorl dextral; growth striae prominent growth lines often distorted, undulating; rib count (mean = 22, SD = 2.5, n = 13), whitish primary ribs raised, bent and crooked, broaden to, extend beyond and scallop the often growth deformed shell lip; 2 – 3 interspersed darker secondary ribs, black to brown rib interstices; fairly distinctive primary rib pattern, abutting paired raised primary ribs on siphonal ridge, flare out shell lip; segment area immediately behind siphonal ribs clear of any ribbing, 4 – 5 primary ribs span posterior end, 4 primary ribs span anterior end. Interior shell margin brown, shell lip coloured cream / white rays and black / brown infilling under primary / secondary ribs and rib interstices respectively; spatula dark brown, siphonal groove dark brown or white, ADM scar indistinct, coloured same as shell margin, cephalic muscle car concave; Thickening of shell lip and spatula occurs across larger shell sizes, translucent, infills and reduces lip scalloping, spatula becomes whitened; Reproductive system (Fig. 47 J; n = 2). Positioned within coelom under the respiratory cavity, hermaphroditic glands positioned to posterior against right foot wall and over foot sole, epiphallic parts positioned between BM and to side of RAM; AO large, elongated bulbous, bluntly pointed (embeds into MG), centrally bent, merges with top of indistinct GA; ED long narrow, slightly twisted; EG relatively large, folded, elongated and pointed; flagellum F 1 absent; AO, GA and ED all muscular white tissue; BD and CD jointly connect into GA between ED, AO and GP; BD longer and much narrower than CD with a prominent loop close to BC, both ducts smooth curved and pass together (BD above CD) through RAM connecting into folds of MG; BC small translucent test bulbous, embedded in soft white folds of AG / MG complex; HD short coiled, links AG to a small brownish finely granulated HG; dark SV embedded within AG, AG larger than HG. Spermatophore (Fig. 47 K). Test thin, translucent (length = 1.47 mm, n = 1, possibly longer as flagellum appears incomplete); head bulbous, tip bluntly rounded, containing a white gelatinous mass; very short taper region into the filamentous transparent flagellum; both sections smooth, featureless; head shorter and much thicker than flagellum (head length = 0.69 mm, ~ 46 % of SPM length; flagellum length = 0.78 mm; head width = 120 μm; flagellum width = 24 μm, n = 1); single SPM coiled in one bursa (AM C. 356978).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFE08262FF68FF02FA8BF7F6.taxon	diagnosis	Comparative remarks. Siphonaria lentula (atra group, unit 73) is the sister group of an unidentified species from Molokai (‘ S. atra group, unit 36 ’ by Dayrat et al. 2014). Both sequences together form a well-differentiated subclade (Figs 1, 2). Siphonaria lentula differs from its sister lineage by COI distances of ≥ 16.6 % (Table S 3). Siphonaria lentula occurs in sympatry with two congeners on LHI: For comparison with S. exulum refer to comparative remarks under that species. Siphonaria pravitas sp. nov. has more raised ribbing and edge scalloping, a shorter ED and BD, with no distal loop, a larger BC, and a more threadlike SPM. The SPM of S. lentula is not typical of the atra group. It is very short, bulbous (Fig. 47 K).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFE08262FF68FF02FA8BF7F6.taxon	distribution	Distribution and habitat. Endemic to LHI (Fig. 48). In this study, found in sheltered to exposed places on rocky shores, at upper to mid-littoral level.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFE2829CFF68FF02FD53FA16.taxon	description	(Figs 46 D – F, R – T, 47 C – G)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFE2829CFF68FF02FD53FA16.taxon	materials_examined	Material examined. Type material. Lectotype of Mallorisiphon oppositus Iredale, 1940, present designation, from Keppel Bay, [Qld, Australia]; coll. H. Bernhard [1940] (AM C. 103714, Fig. 46 D). Four probable paralectoypes from North Keppel Is, [Qld]; coll. H. Bernhard, [1 Jan 1934] (AM C. 108527). Probable paralectotype of Mallorisiphon oppositus Iredale, 1940 from Emu Park, Keppel Bay, Qld. Pres. T. Iredale, 22 May 1953 ‘ Paratype’ (MV F 13839) Other, non-type material. PNG: Kranket Is, N end, Madang, 5 ° 11 ’ S, 145 ° 51 ’ E (AM C. 595915 p [SK 545], C. 595916 p [SK 544]); Biliau I., 05 ° 11.8 ’ S, 145 ° 48.2 ’ E, PM 38 (MNHN IM- 2013 - 15195 p [M 560]). Australia, Qld: W side Kissing Pt, Townsville, 19 ° 14.332 ’ S, 146 ° 48.040 ’ E, Q 26 - 2 (AM C. 585150 p [M 189]); Mackay breakwater wall, 21 ° 06.415 ’ S, 149 ° 14.033 ’ E, Q 14 - 2 (AM C. 584998 d [R 21282]); Yeppoon, Wreck Pt, 23 ° 08.736 ’ S, 150 ° 45.865 ’ E, Q 08 - 4 (AM C. 585598 4 p, C. 585135 p [M 424, SK 237], C. 585136 p [M 425], C. 585137 p [SK 237]), Double Head, 23 ° 09.908 ’ S, 150 ° 47.638 ’ E, Q 08 - 3 (AM C. 585701 8 p); Buff Pt, 23 ° 11.123 ’ S, 150 ° 47.830 ’ E, Q 08 - 2 (AM C. 585597 4 p, C. 585860 p [SK 113], C. 585867 p [M 210], C. 585868 p [M 211]); Zilzie, 23 ° 16.778 ’ S, 150 ° 49.553 ’ E, Q 08 - 1 (AM C. 585343 p). Canoe Pt Gladstone 23 ° 56.155 ’ S, 151 ° 21.964 ’ E, Q 07 - 2 (AM C. 585671 6 p, C. 585865 p [M 212], C. 585866 p [M 213]); Bagara, Hervey Bay, 24 ° 49.180 ’ S, 152 ° 28.011 ’ E, Q 06 - 1 (AM C. 585700 8 p, C. 585129 p [SK 180]); Urangan, Hervey Bay 25 ° 17.504 ’ S, 152 ° 54.664 ’ E, Q 05 - 1 (AM C. 585341 1 p, C. 585864 p [M 209]); Scarborough, Nth Reef, 27 ° 11.432 ’ S, 153 ° 06.755 ’ E, Q 03 - 5 (AM C. 585126 p [SK 136]); 27 ° 11.451 ’ S, 153 ° 06.722 ’ E, Q 03 - 4 (AM C. 585596 4 p); 27 ° 11.589 ’ S, 153 ° 06.892 ’ E, Q 03 - 6 (AM C. 585637 5 p, C. 585819 p [M 458, SK 181], C. 585863 p [M 431, SK 135]), Drury Pt, Q 03 - 7 (AM C. 585409 10 + p), Scarborough Pt, 27 ° 12.168 ’ S, 153 ° 06.980 ’ E, Q 03 - 8 (AM C. 585670 6 p).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFE2829CFF68FF02FD53FA16.taxon	discussion	Taxonomic remarks. Iredale (1940: 440) stated in the original description that M. oppositus was collected ‘ from many places in Qld’. The description was evidently based on more than one specimen and did not contain a type designation. Therefore, all types are syntypes. The largest syntype labelled ‘ holotype’ (Fig. 46 L) is herein designated as the lectotype of M. oppositus for the stabilisation of the name (Art. 74.1 of the Code). Through this designation, the type locality is herein restricted to Keppel Bay, Qld. Our delineation of this species is based on comparative analyses of the morpho-anatomy and mitochondrial genetics of freshly collected topotypes (Fig. 46 E – F) and geographic series of additional specimens (Table S 1). Hubendick (1946: 52) and Cernohorsky (1972: 210) incorrectly treated S. opposita as a synonym of S. atra. However, the figured specimen of ‘ S. atra ’ in Cernohorsky (1972: 210, pl. 60, fig. 1) is a specimen of S. vudaensis sp. nov. Morrison (1972: 56 – 58) treated ‘ Mallorisiphon oppositus ’, as a synonym of S. laciniosa based on similarity in shell form and ‘ common reproductive development’. This synonomy is not supported by examination of type specimens and comparative morpho-anatomy.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFE2829CFF68FF02FD53FA16.taxon	description	External morphology (Fig. 46 R). Foot sole, foot wall, foot edge, cephalic folds and pneumostomal lobe evenly cream; mantle thick translucent in larger specimens, thin in smaller specimens, edge thickened, whitish, lobed, with very light black pigmentation at mantle edge aligning with large primary rib interstices; genital pore inconspicuous, located on foot wall to right anterior of right cephalic fold; two small black epithelial eye spots centralised on two centrally touching, centrally black pigmented cephalic folds; pneumostomal lobe long, under the mantle, unpigmented, behind right cephalic fold. Shell (Figs 46 D – F, R, T; Table S 9). Medium sized (max sl mean = 20.8 mm, SD = 1.9 mm, n = 11), ovate; height very low; apex offset central and strongly to left, apical sides convex to straight, protoconch direction homostrophic (n = 1; Fig. 46 T), shell whorl dextral; growth striae prominent, without radial banding, exterior often evenly light brown, whitened if eroded; shell thickness thin to thickened; rib count (mean = 50, SD = 3.7, n = 11), 17 – 21 primary ribs pale, fairly evenly spread radially, narrow, raised and protrude strongly (often> 1 mm) beyond shell lip to unevenly scallop and corrugate the edge; 1 – 2 distinctly smaller, finer secondary ribs between primary ribs; rib interstices slightly darker; dual slightly spaced primary ribs over siphonal ridge, slightly more prominent, protruding more than other primary ribs, interstice gap wider either side of siphonal ridge. Interior; shell margin dark brown to white, narrow white rays align on grooves over shell margin under primary ribs, weakly furrowed, extend from lip to spatula; siphonal groove distinct, bent, same colour as shell edge; spatula evenly dark chocolate brown to mottled tan, ADM scar distinct, CMS convex, similar colour shell margin; thickening of shell lip commonly occurs, infills interior rib furrows and margin reducing lip scalloping, spatula not whitened. Reproductive system (Fig. 47 C, F; n = 4). HG / AG / MG complex positioned within right side of coelom, against foot wall over foot muscle, under the respiratory cavity; epiphallic parts relatively large, positioned between BM and RAM; GA small indistinct, with singular GP through foot wall; AO large broad elongated pointed, joined to upper GA; ED short, broad, coiled, twisted, joins to side of GA; GA, AO, ED all white muscular fibrous tissue; EG large, longer than ED, soft whitish tissue, folded, joins ED; single short broad flagellum (F 1), shorter and narrower than ED, appears as an extension of ED; BD and CD connect from opposite directions into GA between ED join and GP, distal loop in BD, both ducts long straight smooth thickened whitish featureless, pass closely together through RAM (BD over slightly wider CD) into soft white folded tissues of MG; MG / AG complex relatively large; BC embedded in folds of AG / MG complex close to embedded blackish SV; BD with short distal twisted loop, twisted coiling immediately prior to BC, slightly thinner but much longer than CD; BC relatively small, bulbous, thin whitish translucent test; HD short wide coiled, links ducts in soft white folded tissues of AG to yellowish granulated HG; AG usually larger than HG. Spermatophore (Figs 47 D, E, G). Relatively short and wide (length = 6.37 ± 0.93 mm, n = 6), head section cylindrical, tip bluntly rounded, central white core; flagellum very thin, transparent, tapering to a thread-like end; both sections smooth, featureless; test thin, translucent (head length = 4.9 ± 0.76 mm, ~ 80 % of SPM length, head width = 114 ± 20 μm; flagellum width = 14 ± 2 μm, n = 6); prominent twist before short taper region into filamentous transparent flagellum; three SPMs coiled, embedded in brown gelatinous mass in BC of two specimens.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFE2829CFF68FF02FD53FA16.taxon	diagnosis	Comparative remarks. Siphonaria opposita (atra group, unit 34) is the sister species of S. plana (unit 35); both species together are the sister group of S. denticulata (unit 33) (Figs 1, 2). Siphonaria opposita differs from S. plana by COI distances of ≥ 8.1 % and from S. denticulata by distances of ≥ 15.8 % (Table S 3). Throughout its range, S. opposita has been found in sympatry with eight congeners: For comparisons with S. zelandica, S. denticulata, S. normalis, S. scabra, S. atra, S. viridis, and S. javanica refer to comparative remarks under these species. The specimen figured as ‘ atra group, unit 34 ’ in Dayrat et al. (2014: fig. 5 L) exhibits features consistent with S. opposita. The shell of. S. opposita has frequently been mistaken for S. eumelas (= S. atra).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFFE2829CFF68FF02FD53FA16.taxon	distribution	Distribution and habitat. Endemic to Qld, between Townsville and MacKay (Fig. 48). In this study, found on sheltered rocky shores, at upper to mid littoral level.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF1D829DFF68F802FCE4FDD6.taxon	description	(Figs 49 A – C, O – P, T, 50 A – D)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF1D829DFF68F802FCE4FDD6.taxon	materials_examined	Material examined. Type material. Lectotype of Hebesiphon monticulus Iredale, 1940, present designation, from ‘ Lifu’ [Lifou], Loyalty Islands, NC; coll. 1905 (AM C. 103720 ‘ holotype’, Fig. 49 A). Paralectotype, same data as lectotype (AM C. 410720). Other, non-type material. NC, Lifou: Drueulu, 20 ° 55.570 ’ S, 167 ° 05.067 ’ E LFU 02 - 1 (AM C. 585397 10 + p, C. 585875 p [SK 058], C. 584948 p [M 384], C. 584949 p [M 385], C. 584950 p [M 386], C. 584951 p [M 387]); We Baie de Chateaubriand East coast, 20 ° 54.779 ’ S, 167 ° 15.636 ’ E LFU 01 - 1 (AM C. 584944 p [SK 057], C. 584945 p [SK 056]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF1D829DFF68F802FCE4FDD6.taxon	discussion	Taxonomic remarks. The original description does not contain a type designation. Hence all types are considered as syntypes. The specimen labelled as ‘ holotype’ (Fig. 49 A) is herein designated as lectotype of H. monticulus for the stabilisation of the name (Art. 74.1 of the Code). This specimen is probably the shell figured in Iredale (1940: 441, pl. 34, fig. 11 – 13) and matches the shell dimensions given in the original description. The lectotype is a small specimen displaying a particularly tall shell profile. Our delineation of this species is based on comparative analyses of the morpho-anatomy and mitochondrial genetics of freshly collected topotypes (Fig. 49 C) and geographic series of additional specimens (Table S 1). While the description of S. monticulus in Hubendick (1946: 45) agrees with our concept of this species, the specimen figured from ‘ Wijnkoopsbai, south coast of Java’ (Hubendick 1946: pl. 3, figs 7 – 9) is not of H. monticulus. Moreover, Hubendick (1946) did not examine any specimens from Lifou, Fiji or Tonga. Hence, he has probably misidentified specimens of S. javanica.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF1D829DFF68F802FCE4FDD6.taxon	description	External morphology (Fig. 49 T). Foot sole grey; foot wall, pneumostome, cephalic folds and mantle evenly cream; two small black epithelial eye spots centralised on two thick centrally touching cephalic folds; genital pore inconspicuous, located on foot wall posterior to right cephalic fold; mantle thin translucent extends to shell edge, edge weakly lobed with white band and light black bands aligning to rib interstices; pneumostomal lobe thin and within mantle between the right ADMs, closes the pneumostomal and anal openings at the mantle edge; light black pigmentation over centre of cephalic folds. Shell (Figs 49 A – C, O; Table S 9). Small sized (max sl mean = 12.56 mm, SD = 1.2 mm, n = 5), ovate; height medium to tall; apex offset central to sightly posterior, apical sides convex, protoconch direction central to weakly heterostrophic (n = 1; Fig. 49 O), shell whorl dextral; growth striae distinct, exterior uneven, shell thickness thick; rib count (mean = 42.8, SD = 4.6, n = 5), no clear distinction between primary and secondary ribs, ribs pale white, fairly straight, rib interstices paler, mottled radial colouration band appears to be algal growth in hollows; increasingly raised and protrude slightly beyond shell lip, edge uneven weakly scalloped, strongly corrugated; 3 – 4 fused ribs form an indistinct siphonal ridge. Interior shell lip and margin white, dark to pale brown rays aligning under primary / secondary ribs, span shell margin to golden / whitish spatula, siphonal groove distinct raised above shell lip, same colour as shell edge / margin, ADM scar distinct, CMS straight, paler than shell lip; thickening of shell lip translucent, infills and reduces lip scalloping, spatula becomes whitened. Reproductive system (Figs 50 A, C; n = 3). Positioned within and right side of coelom under the respiratory cavity, hermaphroditic glands positioned to posterior against right foot wall and over foot sole, epiphallic parts positioned to anterior over BM against digestive organs; GA prominent with singular GP through foot wall; AO large, bluntly pointed, joined to upper GA alongside ED; ED elongated, broad, twisted and coiled (without prominent MA), joins to side of GA above BD; GA, AO, ED all white muscular fibrous tissue; EG large, soft whitish tissue, slightly folded, joins ED; single long thick flagellum (F 1), similar length to ED, appears as a continuous extension of ED to EG, laid over BM; BD and CD connect close in opposing directions into GA between AO join and GP, BD with distal loop and prominent MA; both ducts long, straight, smooth, similar thickness of narrow, whitish, featureless, pass closely together through centre of RAM (BD over CD) into soft white folded tissues of MG; MG / AG complex relatively large; BC embedded in folds of AG / MG close to embedded SV; BC large, flat rounded, thin translucent test; HD long, thickened, coiled, brown markings, links ducts in soft white folded tissues of AG to yellowish granulated HG; outer edge of MG lobbed; AG / MG much larger than HG. Spermatophore (Figs 50 B, D). Thread-like, test thin, translucent (length = 13.09 mm, n = 1); head section cylindrical, tip bulbous bluntly rounded, containing a white gelatinous core, tapers into the filamentous transparent flagellum; both sections smooth, featureless; head longer and much thicker than flagellum (head length = 7.71 mm, n = 1; head ~ 58.8 % of SPM length; head width = 80 μm; flagellum width = 13 μm). Single SPM embedded in red-brown gelatinous mass in one BC (AM C. 584951). Radula. Dentition formula 39: 1: 39 (Hubendick 1946: 46).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF1D829DFF68F802FCE4FDD6.taxon	diagnosis	Comparative remarks. Siphonaria monticulus (plicata group, unit 57) represents a well-differentiated lineage. It is the sister species of a subclade containing S. lirata, S. tanguissonensis sp. nov., S. mauiensis sp. nov. and S. undans sp. nov. (Figs 1, 3). S. tanguissonensis sp. nov. and S. lirata differ by COI distances of ≥ 15.4 % and ≥ 17 %, respectively (Table S 7). Siphonaria monticulus has been found in sympatry with three congeners on Lifou, Loyalty Islands: For comparison with S. normalis refer to comparative remarks under that species. Siphonaria hienghenensis sp. nov. has a larger, darker shell with a more prominent and flared siphonal ridge, stronger edge scalloping, a shorter ED, narrower BD, smaller BC, and a less thread-like SPM. Siphonaria bourailensis sp. nov. has a lower, darker shell with more prominent and fewer primary ribs, and a shorter ED and F 1. Siphonaria monticulus exhibits a shell morphology similar to that other species of the plicata group, such as S. nuttallii (Hawaii), S. tongatapuensis sp. nov. (Tonga), S. lirata (Guam), S. plicata (Lifou), S. namukaensis sp. nov. (Fiji), and S. yagasaensis sp. nov. (Fiji). A specimen from Java figured as ‘ S. monticulus’ in Hubendick (1946: 91, pl. 3, fig. 7 – 9) is a misidentification of S. javanica.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF1D829DFF68F802FCE4FDD6.taxon	distribution	Distribution and habitat. Endemic to Lifou, Loyalty Islands, NC (Fig. 48). In this study found in sheltered positions (e. g., hollows of rocky platforms, cliff bases) on exposed rocky shores, mid and lower littoral levels (Fig. 49 P).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF1C8298FCCAFDE2FE84F8B6.taxon	description	(Figs 49 D – G, U – V, 50 E – F)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF1C8298FCCAFDE2FE84F8B6.taxon	materials_examined	Material examined. Type material. Lectotype of Siphonaria alba Hubendick, 1943; 2. fig. 3, 7, present designation, from Nordwatcher, Javasee [near Singapore, Java Sea]; coll. C. Aurivillius, 1891. (UUZM 1574 - 3957, colour image unavailable). Paralectotype, same data as lectotype (UUZM 1574 - 3957). Other, non-type material. Singapore: Lazarus Island causeway, St Johns Island, 01 ° 13.288 ’ N, 103 ° 51.195 ’ E SI 04 - 3 (AM C. 585229 p [M 332], C. 585230 p [M 334], C. 585231 p [M 335], C. 585232 p [M 337], C. 585237 p [SK 175]); Lazarus Island, 01 ° 13.355 ’ N, 103 ° 51.148 ’ E SI 04 - 2 (AM C. 585352 p); Fort Road, drain seawall 01 ° 17.605 ’ N, 103 ° 53.809 ’ E SI 01 - 3 (AM C. 585602 5 p); East Coast Park, seawall, 01 ° 18.643 ’ N, 103 ° 57.077 ’ E SI 01 - 2 (ZRC Moll. 9121 7 p, ZRC. MOL. 24914 p [M 474, SK 281], ZRC. MOL. 24915 p [SK 294]). PNG: Rempi Area, S Dumduman Is., 05 ° 00,2 ’ S, 145 ° 47,6 ’ E PM 12 (MNHN IM- 2013 - 12000 p [M 557]). Timor-Leste: N of Dili, Christi Rea Beach, 8 ° 32.072 ’ S, 125 ° 36.868 ’ E TL 01 - 2 (AM C. 585905 4 p, C. 585274 p [M 440]. Australia, WA: Kimberley, between the Maret Islands, 14 ° 24 ’ S, 124 ° 58 ’ E (WAM S 72337 p); Caffarelli Is., 16 ° 01.991 ’ S, 123 ° 18.625 ’ E, WA 19 - 1 (AM C. 584678 p [M 323]); Conilurus Is, Kimberley, 16 ° 08.875 ’ S, 123 ° 35.234 ’ E, WA 18 - 1 (AM C. 585653 3 p, WAM S 74082 2 p); Catamaran Bay, Cape Levique, Kimberley, 16 ° 27.622 ’ S, 123 ° 00.242 ’ E, WA 22 - 1 (AM C. 585655 4 p, C. 585296 p [SK 424], C. 585299 p [M 070]); Gantheaume Point, Broome, 17 ° 58.384 ’ S, 122 ° 10.677 ’ E, WA 26 - 2 (AM C. 584735 15 p, WAM S 74083 5 p), CI: Ethel Beach, 10 ° 27.827 ’ S, 105 ° 42.497 ’ E CI 02 - 1 (AM C. 584845 p [M 304]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF1C8298FCCAFDE2FE84F8B6.taxon	discussion	Taxonomic remarks. The largest syntype (UUZM UUMS 3957 / 1574) is herein designated as the lectotype of S. alba for the stabilisation of the name (Art. 74.1 of the Code). Our delineation of this species is based on comparative analyses of the morpho-anatomy and mitochondrial genetics of freshly collected topotypes (Fig. 49 D, E – F) and geographic series of additional specimens (Table S 1). We attribute the record of ‘ S. ferruginea ’ from Christmas Is in Smith (1909: 369) to S. alba based on the similar shell and because S. alba occurs on CI while there is no confirmed record of S. ferruginea (= S. plana). Throughout the taxonomic literature, Siphonaria alba has frequently been misidentified as S. atra. Records in Berry (1977), Morton & Morton (1983) from Hong Kong, Tan & Chou (2000) from Singapore, Tan & Kastoro (2004) from South China Sea, Tan & Woo (2010) from Singapore, and Tan & Low (2014) from CKI are attributed here to S. alba because these records are within the range of S. alba but outside of the range of other species in the atra group that exhibit similar shells, such as S. bifurcata (Philippines), S. sirius, S. atra and S. subatra (Japan). Morrison (1972: 56 – 58) treated S. alba as a synonym of S. laciniosa based on similarity in shell form and ‘ common reproductive development’. This synonymy is not supported by examination of type specimens and comparative morpho-anatomy.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF1C8298FCCAFDE2FE84F8B6.taxon	description	External morphology (Fig. 49 V). Foot sole, foot wall, foot edge, mantle, pneumostomal lobe and cephalic folds evenly cream; faint irregular black blotches of pigmentation over centre of cephalic lobes and parts of foot wall; mantle narrow with edge thickened, heavily lobed and white band; genital pore indistinct, located on foot wall to right anterior of right cephalic fold; small black epithelial eye spots centralised on each centrally touching cephalic folds; pneumostomal lobe under the mantle, between the right ADMs. Shell (Figs 49 D – G; Table S 9): medium sized (max sl mean = 21.9 mm, SD = 3.1 mm, n = 9); height low; circular ovate; apex offset central (commonly eroded); apical sides convex, protoconch direction homostrophic (n = 1,), shell whorl dextral; growth striae indistinct, shell margin thick; rib count (mean = 40.8, SD = 4.6, n = 9), primary ribs narrow to broad, solidly raised, pale white protrude beyond shell lip to unevenly scallop and corrugate the edge, some primary ribs protrude 1 – 2 mm beyond shell lip, protrusion at shell lip greater at siphonal ridge and the forming primary ribs larger, interspersed with 1 – 3 pale white finer secondary ribs size; radial colour banding occurs, protoconch area brown, central band paler and shell edge dark brown; rib interstices darker. Interior; spatula, shell margin, ADM scar and siphonal groove evenly white (Fig. 49 D) or dark chocolate brown (Fig. 49 F), white rays from shell lip to margin, align under ribs and match rib width; siphonal groove distinct, same colour as shell edge, slightly curved to right anterior; CMS convex thin; thickening and whitening of shell lip occurs (e. g. Fig. 49 D). Reproductive system (Fig. 50 E; n = 2). Positioned within coelom under the respiratory cavity, hermaphroditic glands positioned to posterior against right foot wall and over foot sole; GP small, singular, positioned through foot wall behind right cephalic fold, GA small, prominent; AO medium sized, elongated, bluntly pointed, slightly bent centrally, joins to top of GA; ED long, narrow, slightly twisted, thickened, joins to back side of GA alongside AO; EG very small, white, folded; single short broad flagellum (F 1), appears as extension of ED; AO, GA and ED all muscular white tissue; BD and CD connect closely but in opposing directions into GA between ED and AO joins, both ducts long narrow smooth featureless, pass together through RAM connecting into MG (BD above CD), BD with distal loop and short MA attached to inner foot wall, additional loop immediately in front of BC; BC large spherical, embedded in folds of MG, test translucent; SV embedded on left side of AG; HD short, thick coils, links small AG to similar sized yellowish granulated HG; MG and AG folded, soft white tissue; sides match curvature of inner foot wall on right posterior of coelom. Spermatophore (Fig. 50 F). Relatively long thread-like (length = 13.87 ± 3.24 mm, n = 2, possibly longer as flagellum on one appears incomplete); test thin, translucent, containing a white gelatinous core mass; over half-length comprises a translucent bulbous cylindrical head section (head length = 8.86 mm, SD = 0.65 mm, n = 2; mean ~ 65 % of SPM length); tip bluntly rounded; head section much thicker than flagellum (head width = 116 ± 16 μm, flagellum width = 12 ± 0 μm, n = 2); both sections smooth, featureless; 7 – 8 SPM tightly coiled in one BC (AM C. 585237). Radula. Dentition formula 44: 1: 44 (Hubendick 1946: 56).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF1C8298FCCAFDE2FE84F8B6.taxon	diagnosis	Comparative remarks. Siphonaria alba (atra group, unit 39) is the sister species of a subclade formed by S. atra and S. hienghenensis sp. nov. (Figs 1, 2). It differs from these two species by COI distances of ≥ 13 % (S. atra) and ≥ 8.3 % (S. hienghenensis sp. nov.) (Table S 3). Throughout its range, S. alba has been found in sympatry with fifteen congeners. Four congeners are sympatric in Singapore: Siphonaria costellata sp. nov. has a taller shell with slightly more raised and even ribbing, a shorter ED and BD, smaller AO and BC, and a shorter SPM. Siphonaria caubianensis sp. nov. has a darker shell exterior and interior, broader and more raised primary ribs, a more posterior and left offset apex, a stronger scalloped shell edge, a smaller AO and BC, no BD distal loop. For comparisons with S. normalis and S. sirius refer to comparative remarks under these species. Two species are sympatric in Cebu, Philippines: For comparisons with S. bifurcata and S. sipho refer to comparative remarks under these species. Siphonaria umbra sp. nov. is sympatric in CI. It has a smaller, darker shell with more even ribbing, smaller AO, shorter, wider ED, and larger BC. Six species are sympatric in Timor-Leste: For comparisons with S. atra, S. viridis, and S. javanica refer to comparative remarks under these species. Siphonaria campestra sp. nov. has a smaller, darker shell with less prominent siphonal ridge and weaker edge scalloping, a smaller AO and BC, shorter ED and BD, and a shorter SPM. Siphonaria forticosta sp. nov. has a larger shell with a slightly more posteriorly offset apex, weaker edge scalloping and darker interior, a smaller AO, shorter ED, and a larger F 1. Siphonaria planucosta sp. nov. has a smaller, darker shell with less raised ribbing and weaker edge scalloping, a smaller AO, shorter ED, and shorter SPM. Three species are sympatric in WA (along with S. atra, and S. viridis): Siphonaria gemina sp. nov. has smaller, taller, paler shell with stronger edge scalloping, a smaller AO, shorter ED and BD, and a less thread-like SPM. Siphonaria restis sp. nov. has a paler shell with more uneven ribbing and stronger edge scalloping, a smaller AO, shorter wider ED, and larger BC. For comparison with S. zelandica refer to comparative remarks under that species. The RS (Fig. 50 E) and SPM (Fig. 50 F) of S. alba shown herein correspond well the RS and SPM figured in Hubendick (1943: 3, fig. 10 – 11; 1945: 3 figs 10 – 11) although no serration on SPM flagellum was observed. A specimen from Mindanao figured as ‘ S. acervus ’ in Hubendick (1946: pl. 3, figs 10 – 12) is probably a specimen of S. alba for exhibiting typical shell characteristics. It is within the known distribution of S. alba. A specimen from Palawan figured as ‘ Siphonaria sirius ’ in Springsteen & Leobrera (1986: 285, pl. 81, fig. 20) is a misidentification and likely a specimen of S. alba for the siphonal ridge exhibiting multiple and not just a single rib. A specimen from Seribu Islands, Java Sea figured as ‘ S. laciniosa’ in Dharma (2005: pl. 79, figs 20 a, b) is a misidentification and likely a specimen of S. alba as it matches the shell morphology of this species (Fig. 49 D – G; paired siphonal ribs) and is from within the known range of this species. A specimen from Long Dong, Taiwan figured as ‘ atra group, unit 39 ’ in Dayrat et al. (2014: 263, fig. 5 S) is morphologically consistent with S. alba.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF1C8298FCCAFDE2FE84F8B6.taxon	distribution	Distribution and habitat. Widely distributed through tropical northwestern Pacific, including Taiwan, Sulawesi, Singapore, Thailand, Philippines, CI, northern Australia (Kimberey, WA) and Timor-Leste, Dili (Fig. 45). In this study found at sheltered positions on moderately exposed rocky shores, at upper and lower littoral levels (Fig. 49 U).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF19829AFF68F802FAB9FE96.taxon	description	(Figs 49 H – N, Q – S, 50 G – H)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF19829AFF68F802FAB9FE96.taxon	materials_examined	Material examined. Type material. Holotype of S. asghar Biggs, 1958 from Hormuz Island, Persian Gulf; coll. 12 May 1933 (NHMUK 1958.6.13.13, Fig. 49 H). Six paratypes of S. asghar Biggs, 1958 same data as holotype (NHMUK 1958.6.13.14 – 19, Figs 49 I – K). Other, non-type material. Pakistan: Karachi, Clifton Beach, 24 ° 45.500 ’ N, 67 ° 05.968 ’ E, PA 02 - 1 (AMC. 585895 p); French Beach, 24 ° 50.345 ’ N, 66 ° 49.244 ’ E, PA 01 - 2 (AM C. 585899 5 p); 24 ° 50.367 ’ N, 66 ° 49.387 ’ E, PA 01 - 1 (AM C. 585818 13 p, C. 585843 p [M 243], C. 585844 p [M 244], C. 585845 p [M 245], C. 585846 p [M 456, SK 190]); Karapir Beach, 24 ° 50.590 ’ N, 66 ° 53.927 ’ E, PA 01 - 3 (AM C. 585901 7 p, C. 585847 p [SK 144], C. 585854 p [M 235], C. 585855 p [M 236], C. 586001 p [SK 532]); Bubiji Beach, 24 ° 53 ’ N, 67 ° 01 ’ E 24 ° 53 ’ N, 67 ° 01 ’ E (WAM S 72336 9 p, S 74084 p [SK 408], S 74085 p [SK 407], S 74086 p [SK 303], S 74087 p [SK 359], S 74088 p [SK 421]), S 113648 [SK 530], S 113649 [SK 531]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF19829AFF68F802FAB9FE96.taxon	discussion	Taxonomic remarks. The holotype of S. asghar (Fig. 49 H) is a worn or malformed specimen, unlike the paratypes (Fig. 49 I – K). Our delineation of this species is based on comparative analyses of the morpho-anatomy and mitochondrial genetics of freshly collected topotypes (Fig. 49 L, M; Table S 1). Smythe (1979: 69) incorrectly considered S. asghar as a synonym of S. rosea (= S. crenata). However, both species differ conchologically and anatomically. The RS structure of S. tenuicostulata (= S. carbo) figured in Hubendick 1947 a: fig. 1) differs from that of S. asghar. A specimen figured as ‘ S. asghar ’ in Dayrat et al. 2014: fig. 3 C) corresponds well with the paratypes; Figs 49 I – K). Compared to freshly preserved specimens from Karachi (Figs 49 L – M), the types (Figs 49 H – K) are rather small, around half the size of the topotypes.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF19829AFF68F802FAB9FE96.taxon	description	External morphology. Foot sole, foot wall, foot edge cephalic folds and pneumostomal lobe evenly cream coloured; black pigmentation absent except for central patch on cephalic folds; mantle cream coloured, narrower than foot wall, edge lobed thickened; pneumostome under mantle between right ADMs. Shell (Figs 49 H – N, Q, R; Table S 9). Small sized (max sl mean = 15.1 mm, SD = 2.2 mm, n = 8), ovate, height medium; apex offset weakly to posterior and left, apex usually eroded (often upper 2 / 3 rds eroded), apical sides strongly convex, protoconch direction undetermined, shell whorl dextral; growth striae inconspicuous, shell thickness medium; rib count (mean = 45, SD = 4.9, n = 8), ribs flat / unraised, narrow, sometimes wavy, very even in width, pale grey / brown; non-protruding at shell lip; shell edge weakly corrugated, not scalloped; few secondary ribs, rib interstices darker; siphonal ridge indistinct. Interior shell margin brown and white; brown rays even in width and depth, widen from lip to margin, align under rib interstices, siphonal groove indistinct; spatula and ADM scar chocolate brown paler in parts; ADM scar distinct, CMS weakly convex. Smaller shells tend to display greater white rays on shell margin, a paler brown / white spatula. Juvenile specimens show paired primary ribs over siphonal ridge. Reproductive system (Fig. 50 G; n = 4). Positioned within coelom under the respiratory cavity, hermaphroditic glands positioned to posterior against right foot wall and on foot muscle, epiphallic parts positioned between RAM and BM, EG in front of BM, F 1 lies on foot muscle to left side / under BM; AO small, broad, rounded, MA on side, bluntly pointed, joins at top of larger GA; ED relatively short (often embedded in soft tissue), narrow, straight; EG medium with folds, broad flagellum F 1 appears as an extension of ED at join with EG; AO, GA and ED all muscular white tissue; BD and CD closely connecting, join into GA close to ED and AO (very distinct); BD longer and narrower than CD without distal loop or MA, CD and BD smooth, narrow, both pass together through RAM (BD above CD) and bent before connecting into folds of MG; BC embedded in MG, size medium, bulbous test soft translucent; HD long, brown flecked, sac-like, edge lobed, links AG to small elongated narrow brownish granulated HG; MG and AG small folded soft white tissue; SV embedded within AG, AG larger than HG, sides match curvature of inner foot wall at right posterior quarter of coelom. Spermatophore (Fig. 50 H). Elongated drop-shaped, test thin, translucent (length = 1.28 ± 0.3 mm, n = 3), head bulbous, broad, tip bluntly rounded, containing a white gelatinous mass; taper region to flagellum reduced; both sections; head much larger than flagellum (head length = 0.85 ± 0.03 mm, n = 3, ~ 66 % of SPM length); tip bluntly rounded; head section much thicker than flagellum (head width = 580 ± 51 μm, flagellum width = 120 ± 35 μm, n = 3), both sections smooth, featureless; 3 SPM tightly coiled in one BC (AM C. 586001).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF19829AFF68F802FAB9FE96.taxon	diagnosis	Comparative remarks. Siphonaria asghar (pectinata group, unit 3) is the sister species of the species pair S. carbo and S. itampoloensis sp. nov. (Figs 1, 4). It differs from these species by COI distances of ≥ 10.9 % (S. itampoloensis) and ≥ 11.1 % (S. carbo) (Table S 8). Siphonaria asghar has been found in sympatry with four congeners at Karachi, Pakistan: For comparisons with S. crenata, S. belcheri, and S. kurracheensis refer to comparative remarks under these species. Siphonaria perexigua sp. nov. has a taller, internally paler shell with less raised and wider ribs, shorter, bulbous AO, longer and narrower BD, wider ED and a longer F 1. Conchologically similar are S. capensis (SE Africa) and S. striata sp. nov. (Madagascar), but S. asghar exhibits finer, more numerous, and flatter ribs.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF19829AFF68F802FAB9FE96.taxon	distribution	Distribution and habitat. Coasts of the Arabian Sea, recorded at Q’rum, Muscat (Oman), Karapir, Karachi (Pakistan) and western coast of India (Fig. 51). In this study found in sheltered positions on exposed rocky shores, upper and mid littoral levels (Fig. 49 S).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF1B829BFCCAFE22FB55FA55.taxon	description	(Figs 50 I – K, 52 A – D, M – N)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF1B829BFCCAFE22FB55FA55.taxon	materials_examined	Material examined. Type material. Holotype of S. propria, from Lower littoral, S side Kaikoura Peninsula, E coast, South Island, NZ (NMNZ M. 77363, Fig. 52 A). Twenty-six paratypes same data as holotype (NMNZ M. 77364, 10 p; AM C. 130361, 5 d, 11 p). Lectotype of Siphonaria cookiana Suter, 1909 b: 258, designation by Boreham (1959: 71), GNS TM 1197 (figured in Jenkins, 1983: 21, pl. 3 e). Six paralectotypes same data as lectotype (GNS TM 1198 - 1202, 5, figured in Jenkins, 1983: 21, pl. 3 f – g; AM C. 29118, 1). Other, non-type material. NZ, North Island: Fiordland, Centre Island, Beetles, Preservation Inlet, 46 ° 8 ’ S, 165 ° 40 ’ E (MA. 100954 4 p); N end Seaview Marina, Lower Hutt, South Island, NZ, 41 ° 14.85 ’ S, 174 ° 54 ’ E, Stn 2011011 (NMNZ M. 331452 6 p, [M 509, SK 429], [M 510, SK 430], [M 511, SK 431], [SK 428 protoconch H 12]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF1B829BFCCAFE22FB55FA55.taxon	discussion	Taxonomic remarks. The original description of S. cookiana (Suter, 1909 b: 258) was based on a series of syntypes that represented two distinct species. Boreham (1959: 71) subsequently designated the lectotype without realising that the syntypes were a mixed series. The lectotype was subsequently identified as a juvenile of S. australis (Jenkins 1983: 29, pl. 3 e) rendering the name S. cookiana a junior synonym of the latter. However, the paralectotypes of S. cookiana represented an unnamed species for which Jenkins (1983) described S. propria. The statement in Hutton (1873: 55) that ‘ S. denticulata is also found in Tasmania and Australia’ is erroneous and probably refers to specimens of S. propria rather than S. australis. Siphonaria denticulata as delineated herein does not occur in New Zealand. Raven & Bracegirdle (2010: 46) erroneously listed S. cookiana as a synonym of S. propria.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF1B829BFCCAFE22FB55FA55.taxon	description	External morphology. Foot sole broad, smooth and cream to pale yellow. Foot wall smooth to weakly pustulose, superficially unevenly mottled with grey to black markings, concentrated over two cephalic folds; mantle wide and thin, greyish with irregular black bands corresponding under rib interstices, mantle edge banded white to cream; small black eye spot centralised on each cephalic fold; pneumostomal lobe long, whitish, weakly shaded with mottled black markings. Shell (Figs 52 A – D, M – N; Table S 9). Ovate, small sized (max sl mean = 12.5 mm, SD = 1.4 mm, n = 5), posteriorly wide, height medium; apex weakly offset to posterior and left (Jenkins 1983: pl. 5, fig. a – f), and to left of shell centre line; protoconch notched, direction homostrophic (n = 2; Fig. 52 M), shell whorl dextral; exterior pale brown, apical sides weakly convex, growth striae irregular prominent; rib count (mean = 34.6, SD = 1.7, n = 5), ribs slightly raised, irregularly spaced; curve, broaden from apex to protrude weakly beyond shell lip; 1 – 2 secondary ribs between primary, dual juxtapose primary ribs form distinct siphonal ridge. Interior dark brown, spatula tan to cream, siphonal groove clear, purple-brown to dark brown, curving from shell edge to spatula; white rays weakly corrugate shell edge, extend to shell margin, aligned under external ribs; CMS narrow, shallow, concave. Shell lip may be thickened, whitish or purple-brown. Reproductive system (Fig. 50 I; n = 2). Positioned within entire right side of coelom, against foot wall on foot muscle; epiphallic parts positioned between BM and RAM. GA small, with singular GP through foot wall behind right cephalic fold; AO indistinct, ED short, broader than BD and CD, unbent, joins to top of GA; GA, AO, ED all white muscular fibrous tissue; EG small, folds of soft whitish tissue, joins ED; single flagellum (F 1), with minor bends, appears as a slightly longer and narrower extension of ED; BD and CD connect side-by-side into GA between ED join and GP, both ducts long, narrow, slightly bent, minor unevenness, whitish, pass closely together just inside outer RAM (BD over CD) into soft white folded tissues of MG; CD connects to ducts in MG / AG complex; BC embedded in MG folds close to large embedded whitish SV; BC spherical to sack-like, thin whitish translucent test; HD short, wide, coiled, links ducts in soft white folded tissues of AG to yellowish granulated HG; outer edge of MG lobbed; AG larger than HG, outer sides match curvature of inner foot wall. Spermatophore (Figs 50 J – K): Thread-like (length = 4.63 ± 0.26 mm, n = 3), translucent, test thin; head section bluntly rounded, cylindrical, containing a core white gelatinous mass, tapers along the transparent flagellum to a thin tip; both sections smooth, featureless. Head section shorter wider than flagellum (head length = 4.01 ± 0.16 mm, ~ 87 % of SPM length, head width = 112 ± 13 μm; flagellum width = 17 ± 0 μm, n = 3), 3 SPM tightly folded in one BC (NZNM M. 331452). Radula. (Jenkins 1983: 27, pl. 6 g – j). Mean dentition formula is 21: 1: 21 (SD = 3.9, n = 6) with 108 weakly curved (anteriorly convex) transverse rows (SD = 10.9, n = 6), single central tooth with pointed mesocone, flanked by 21 half row laterals, 9 (SD = 2.6) mid and 12 (SD = 1.4) outer lateral teeth means (n = 6), inner lateral teeth are absent; mid lateral mesocone bicuspidate separated by a shallow “ U ” to “ V ” shaped cleft, with the inner cusp longer than the outer, with strongly branching pointed ectocone; aberrant laterals are common appearing as extremely broad teeth; outer laterals have a square basal plate supporting a broad, flat ‘ chisel’ like unicuspidate mesocone flanked by pointed single ecto- and endocones, widths and angles of separation of endo and ectocones are variable.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF1B829BFCCAFE22FB55FA55.taxon	diagnosis	Comparative remarks. In our mitochondrial phylogeny, S. propria (lateralis group, unit 90) is the sister species of S. australis (Figs 1, 4). Both species form a well-differentiated subclade and differ from each other by COI distances of ≥ 10.4 % (Table S 8). In NZ this species has been found in sympatry with two congeners: For comparisons with S. australis and S. obliquata refer to comparative remarks under these species. The RS and SPM of ‘ S. cookiana ’ figured in Hubendick (1945: figs 21, 24, 26; reproduced in Berry 1977, fig. 19) are consistent with RS and SPM of S. propria (Figs 50 I – K) shown here except for details of the RS (CD / BD / GP / GA junction, which Hubendick portrayed as wider and with ducts more separated). The radula was briefly described by Hutton (1883: 143) as ‘ S. zelandica ’, which was repeated by Suter (1913: 601). The cusps and type of lateral teeth as seen here agree with Hutton (1883: 143, pl. 17, fig. D). However, the number of teeth and rows recorded by Hutton (33 - 40: 1: 40 - 33 with 130 – 140 rows) are well above those reported here.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF1B829BFCCAFE22FB55FA55.taxon	distribution	Distribution and habitat. Endemic to New Zealand (southern end of North Island, South Island, Stewart and Chatham Islands; Fig. 51). In this study found in sheltered positions on moderately exposed to exposed rocky shores, mid to lower littoral levels (Fig. 52 N).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF1A8297FCCAFA62FD61FF35.taxon	description	(Figs 52 G – I, Q – S, 53 A – B)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF1A8297FCCAFA62FD61FF35.taxon	materials_examined	Material examined. Type material. Holotype of S. jeanae from near boat wharf, Ceduna, SA; coll. B. B. Collette and J. R. Paxton, 4 Feb 1970 (AM C. 123712, Fig. 52 G); 23 paratypes; same data as holotype (AM C. 123713 3 p, 4 d; SAM D. 16383 6 p); Neptune Bay, SA (MV F 15260 5 p); Radar Reef, Rottnest Island, WA coll. R. Black, 4 Aug 1979 (WAM 1798 – 83 5 d). Other, non-type material. Australia, Vic: Bastion Head Mallacoota, 37 ° 34.429 ’ S, 149 ° 45.927 ’ E, V 09 - 1 (AM C. 585360 p); Cape Conran, 37 ° 48.798 ’ S, 148 ° 43.608 ’ E, V 08 - 2 (AM C. 585727 9 p); Cape Schanck, 38 ° 29.951 ’ S, 144 ° 53.369 ’ E, V 06 - 4 (AM C. 585609 4 p); Point Lonsdale (nr Queenscliff), 38 ° 17.276 ’ S, 144 ° 36.977 ’ E, V 05 - 1 (AM C. 585440 10 + p, C. 585288 p [M 102]); Loutit Bay Lorne, 38 ° 31.190 ’ S, 143 ° 59.429 ’ E, V 03 - 2 (AM C. 585434 10 + p, C. 585285 p [M 103]); Crofts Bay, 38 ° 35.363 ’ S, 142 ° 50.633 ’ E, V 01 - 3 (AM C. 585649 5 p); Armstrong Bay, 38 ° 21.012 ’ S, 142 ° 21.633 ’ E, V 01 - 2 (AM C. 585571 3 p). Tas: Park Beach Dodges Ferry, 42 ° 51.716 ’ S, 147 ° 36.665 ’ E, T 03 - 4 (AM C. 585263 p [M 108]); Haleys Beach Gibson Peninsula, 32 ° 45.084 ’ S, 134 ° 05.490 ’ E, SA 03 - 4 (AM C. 585207 p [M 101]). SA: Cape Northumberland, 38 ° 03.503 ’ S, 140 ° 40.378 ’ E, SA 15 - 1 (AM C. 585547 20 + p, C. 585219 p [M 203], C. 585220 p [M 204]), Port MacDonnell, 38 ° 03.308 ’ S, 140 ° 39.398 ’ E, SA 15 - 2 (AM C. 585546 20 + p, C. 585221 p [SK 017], C. 585222 p [M 202]); Cape Thomas, 37 ° 04.461 ’ S, 139 ° 44.659 ’ E, SA 14 - 1 (AM C. 585545 20 + p, C. 585217 p [M 201]); Fisheries Bay, Lands End, 35 ° 37.999 ’ S, 138 ° 06.921 ’ E, SA 13 - 2 (AM C. 585567 3 p); Groper Bay nr West Cape, 35 ° 14.108 ’ S, 136 ° 49.883 ’ E, SA 10 - 1 (AM C. 585484 19 p, C. 585214 p [SK 011]); Pondalowie Bay, 35 ° 13.989 ’ S, 136 ° 49.892 ’ E, SA 10 - 2 (AM C. 585425 10 + p); Fishery Bay, Cape Wiles, 34 ° 55.107 ’ S, 135 ° 41.086 ’ E, SA 05 - 1 (AM C. 585709 8 p); Port Neill, 34 ° 07.102 ’ S, 136 ° 21.271 ’ E, SA 06 - 1 (AM C. 585424 10 + p, C. 585213 p [SK 012]); Port Moonta, 34 ° 03.273 ’ S, 137 ° 33.592 ’ E, SA 09 - 1 (AM C. 585711 8 p); Salmon Point, 33 ° 38.547 ’ S, 134 ° 51.916 ’ E, SA 04 - 2 (AM C. 585537 20 + p); ESE of Pt Lincoln, Dangerous Reef, 32 ° 49.05 ′ S, 136 ° 13.05 ’ E AM C. 595961 4 p) Wellesley Point, 33 ° 38.483 ’ S, 134 ° 51.963 ’ E, SA 04 - 1 (AM C. 585544, 20 + p, C. 585212 p [M 123]); Haleys Beach Gibson Peninsula, 32 ° 45.084 ’ S, 134 ° 05.490 ’ E, SA 03 - 4 (AM C. 585441, 10 + p, C. 585207 d, C. 585211 p [M 122]); Rocky Point, 32 ° 12.250 ’ S, 133 ° 14.861 ’ E, SA 02 - 4 (AM C. 585472,16 p); Cedunanearboatwharf, 32 ° 8 ’ S, 133 ° 41 ’ E (AM C. 123712 d, AM C. 123713 4 d, 13 p); Port Le Hunte Point Sinclair, 32 ° 05.681 ’ S, 132 ° 59.299 ’ E, SA 02 - 1 (AM C. 585419 10 + p), 32 ° 05.554 ’ S, 132 ° 59.476 ’ E, SA 02 - 2 (AM C. 585420 10 + p); Cactus Beach, Point Sinclair, 32 ° 05.135 ’ S, 132 ° 58.943 ’ E, SA 02 - 3 (AM C. 585439 10 + p, C. 585209 p [M 121]); Wandrilla Beach, nr Cape Nuyts, 32 ° 01.894 ’ S, 132 ° 16.052 ’ E, SA 01 - 1 (AM C. 585418 10 + p). WA: Alexander Bay 2 33 ° 53.467 ’ S, 122 ° 44.995 ’ E, WA 64 - 4 (AM C. 584775 5 p, WAM S 74133 5 p); Alexander Bay, 33 ° 53.374 ’ S, 122 ° 44.922 ’ E, WA 64 - 3 (AM C. 584715 5 p, WAM S 74132 5 p); Salmon Beach Esperance, 33 ° 53.254 ’ S, 121 ° 50.381 ’ E, WA 64 - 2 (AM C. 584685, 2 p, WAM S 74131 2 p); Bremer Bay Boat Harbour 34 ° 25.613 ’ S, 119 ° 23.818 ’ E, WA 63 - 2 (AM C. 584772, 5 p, WAM S 74130 5 p); Point Henry 34 ° 28.177 ’ S, 119 ° 21.708 ’ E, WA 63 - 1 (AM C. 584771, 5 p, C. 585313 p [M 027], WAM S 74129 5 p); Cape Riche 34 ° 36.213 ’ S, 118 ° 45.401 ’ E, WA 62 - 5 (AM C. 584713, 5 p, WAM S 74128 5 p); Lookout Point Point Gardner 34 ° 53.449 ’ S, 118 ° 25.397 ’ E, WA 62 - 4 (AM C. 584770, 5 p, WAM S 74127 5 p); Cave Point 35 ° 06.965 ’ S, 117 ° 54.080 ’ E, WA 62 - 1 (AM C. 584767, 5 p, WAM S 74125 5 p); Whaling Cove 35 ° 03.372 ’ S, 117 ° 55.598 ’ E, WA 62 - 3 (AM C. 584769, 5 p, WAM S 74126 5 p); Peaceful Bay; 35 ° 02.989 ’ S, 116 ° 55.769 ’ E, WA 60 - 8 (AM C. 584746, 15 p, WAM S 74122 5 p), 35 ° 02.865 ’ S, 116 ° 55.722 ’ E, WA 60 - 7 (AM C. 584745, 15 p, WAM S 74121 5 p); Wilson Head Ocean Beach 35 ° 02.250 ’ S, 117 ° 19.894 ’ E, WA 61 - 1 (AM C. 584766, 5 p, WAM S 74124 5 p); Augusta 34 ° 20.451 ’ S, 115 ° 10.069 ’ E, WA 60 - 5 (AM C. 584778, 5 p, WAM S 74120 5 p); Sarge Bay Cape Leeuwin 34 ° 22.091 ’ S, 115 ° 08.820 ’ E, WA 60 - 4 (AM C. 584728, 5 p, WAM S 74119 5 p); Point Dalling Dunsborough 33 ° 35.955 ’ S, 115 ° 06.315 ’ E, WA 59 - 4 (AM C. 584777, 5 p, C. 585312 p [M 130], WAM S 74116 3 p); Point Casuarina Bunbury 33 ° 18.544 ’ S, 115 ° 38.201 ’ E, WA 59 - 3 (AM C. 584686, 2 p, WAM S 74115 3 p); groyne nr Robert Point Mandurah 32 ° 31.270 ’ S, 115 ° 42.409 ’ E, WA 59 - 1 (AM C. 585577, 3 p); Fremantle Hbr breakwater 32 ° 03.342 ’ S, 115 ° 43.987 ’ E, WA 58 - 5 (AM C. 584765, 5 p, WAM S 74114 5 p); Quinns Rock 31 ° 39.822 ’ S, 115 ° 41.345 ’ E, WA 58 - 4 (AM C. 584671, 3 p, WAM S 74113 2 p); Rottnest Is: Longreach Bay point, 31 ° 59.333 ’ S, 115 ° 32.063 ’ E RI 01 (AM C. 585757, 3 p, C. 585820 p [SK 215]); Radar Reef, 32 ° S, 115 ° 18.72 ’ E (AM C. 595962, 10 + p); W end Thomson Bay, 32 ° S, 115 ° 32 ’ E (AM C. 320123 8 p); Strickland Bay, 32 ° S, 115 ° 30 ’ E (AM C. 595963 10 + p). Cape Leschenault, 31 ° 17.508 ’ S, 115 ° 27.089 ’ E, WA 58 - 3 (AM C. 584726 6 p, WAM S 74112 6 p); Grey, 30 ° 39.968 ’ S, 115 ° 08.072 ’ E, WA 58 - 2 (AM C. 584670 1 p, WAM S 74111 2 p); Jurien Bay, 30 ° 17.244 ’ S, 115 ° 02.482 ’ E, WA 58 - 1 (AM C. 584695 3 p, WAM S 74110 2 p); Bunker Bay, 33 ° 32.300 ’ S, 115 ° 01.951 ’ E, WA 60 - 2 (AMC. 58474715 p, WAMS 74118 5 p); Yallingup, 33 ° 38.358 ’ S, 115 ° 01.481 ’ E, WA 60 - 9 (AM C. 584779 7 p, WAM S 74123 5 p); Sugarloaf Rock Cape Naturaliste, 33 ° 33.536 ’ S, 115 ° 00.467 ’ E, WA 60 - 1 (AM C. 584700 3 p, WAM S 74117 4 p); Cowaramup Point, 33 ° 51.934 ’ S, 114 ° 58.904 ’ E, WA 60 - 3 (AM C. 585520 2 p); Freshwater Point, 29 ° 36.256 ’ S, 114 ° 58.464 ’ E, WA 57 - 3 (AM C. 584762 5 p, WAM S 74107 5 p); Cambawarra Head Green Head, 30 ° 04.136 ’ S, 114 ° 57.830 ’ E, WA 57 - 6 (AM C. 584764 5 p, WAM S 74109 5 p); Illawong, 29 ° 42.254 ’ S, 114 ° 57.542 ’ E, WA 57 - 5 (AM C. 584763 5 p, WAM S 74108 5 p); S end Leander Point Port Denison, 29 ° 16.725 ’ S, 114 ° 54.918 ’ E, WA 57 - 2 (AM C. 584761 5 p, WAM S 74106 5 p); Leander Point Port Denison, 29 ° 16.568 ’ S, 114 ° 54.858 ’ E, WA 57 - 1 (AM C. 584759 5 p, WAM S 74105 5 p); Cape Burney Geraldton, 28 ° 52.084 ’ S, 114 ° 38.056 ’ E, WA 54 - 1 (AM C. 584757 5 p, WAM S 74104 5 p); Horrocks, 28 ° 21.469 ’ S, 114 ° 24.751 ’ E, WA 53 - 1 (AM C. 584780 6 p, WAM S 74103 6 p); Chinamans Rock Kalbarri, 27 ° 42.776 ’ S, 114 ° 09.361 ’ E, WA 52 - 1 (AM C. 584742 12 p, WAM S 74102 8 p).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF1A8297FCCAFA62FD61FF35.taxon	discussion	Taxonomic remarks. Our delineation of this species is informed by comparative analyses of the morpho-anatomy and mitochondrial genetics of freshly collected topotypes (Fig. 52 H – I; Table S 1).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF1A8297FCCAFA62FD61FF35.taxon	description	External morphology (Fig. 52 S). Foot sole evenly dark grey; foot wall darker than sole with evenly dispersed subepithelial pustules, paler to foot edge; fringing mantle narrow translucent at foot wall gradually becoming opaque with a thickened paler band at the unlobed mantle edge; genital pore inconspicuous, located on foot wall to right anterior of right cephalic fold; two small black epithelial eye spots centralised on two thick centrally touching dark grey cephalic folds; thin pale grey pneumostomal lobe part of the mantle, between the right ADMs, closes the pneumostome and anus at the mantle edge. Shell (Figs 52 G – I, Q – S; Table S 9). Small sized (max sl mean 8.7 mm, SD = 1.7 mm, n = 10), ovate; height medium, slightly broader posteriorly; thickness thin; apex offset posterior and left; apical sides convex; protoconch direction heterostrophic (n = 3; Fig. 52 Q), shell whorl dextral; siphonal ridge is indicated externally by two broad juxtaposed radial ribs on the right side of the shell. These siphonal ribs don’t project beyond the shell lip. Growth striae may be regular and indistinct (Jenkins, 1984: pl. 1 a, d, g) or more commonly irregular and coarse (Jenkins, 1984: pl. 2 a, c, e). This variation in growth striae, only apparent in specimens from Ceduna SA, may be due to differences in habitat exposure, compared to most locations where S. jeanae occurs, Ceduna is sheltered. The radial ribbing is flat and unraised with irregular, pale blue-grey ribs and narrower brown interstices, both of which narrow and curve ad-apically, rib count (mean = 27, SD = 3.8, n = 10), variable as is rib width (Jenkins, 1984: pl. 1 a – i). The shell interior is smooth and glossy with a white to pale blue spatula. Between the thin, purplish grey ADM scar and the spatula, the interior is purplish brown. The siphonal groove is shallow and straight. The brown to grey outer lip is thickened and unsculptured with white radial bands corresponding with exterior radial ribs. Most specimens from south-western Australia have eroded exteriors, irregular growth striae and thickened shell lips (Jenkins, 1984: pl. 2 a, c, e). Reproductive system (Fig. 53 A; n = 2). HG (ovotestis), AG / MG complex located in posterior region of coelom, against inner foot wall and under the respiratory system; HG large, yellow, granular, joined to anterior of the soft white translucent semicircular folds of smaller AG by a short pink lobed HD; HD passes through folds of AG / MG complex. SV small ovate pinkish, embedded in anterior of AG / MG complex, connected via a short thin duct to the junction of the HD and emerging CD. BC small and embedded in folds of MG / AG close to SV. BD and CD long, thin, white, smooth, non-looped, featureless, pass together through the ADM and both enter adjacently into the GA. BC small, brown, appears spherical when holding SPM but deflated when empty. ED long, thick, white and may be centrally looped (Jenkins, 1984: fig 1 b), enters the GA almost opposite the BD and CD juxtapose points of entry. No accessory organ present; EG, cream, soft, lobed, longer than ED, situated to the right posterior of the BM; Single flagellum, F 1 short, white, branching from the junction of the EG and ED. GP small, opens from the GA through the foot wall, under the mantle and posterior to the right cephalic fold. Spermatophore (Fig. 53 B) (original description in Jenkins, 1983: 115). Short, bulbous (length = 5.5 mm ± 0.14 mm, n = 2); body cylindrical, weakly bulbous; test thin, white opaque core, short tapering section merging head with filamentous flagellum; head bluntly rounded, shorter thinner than translucent flagellum (head length = 2.15 ± 0.07 mm; ~ 39 % of SPM length; head width = 200 ± 40 μm, flagellum width = 50 ± 10 μm, n = 2); 4 SPMs tightly coiled in BC of one specimen (AM C. 585820). Radula and jaw (original description in Jenkins, 1984: pl. 2 g, h, i). The radula has a typically siphonariid morphology; a central tooth with an individually variable number of mid and outer lateral teeth arranged in longitudinal rows. The mean dentition formula is 25: 1: 25 (n = 3, SD = 2.6) with about 95 transverse rows (SD = 5.3). These rows are parallel and weakly curved (anteriorly convex). One central tooth is present and of the 25 half row laterals, 10.5 (SD = 1.2) are mid and 14.5 (SD = 2.1) outer lateral teeth means respectively. All teeth are weakly concave posteriorly. The central tooth is narrow with a short, pointed mesocone (Jenkins 1984: pl. 2 g, h) with a lower profile than the flanking laterals. Inner laterals characterised by having no side denticles, are absent. The central tooth’s base is narrow in the mid-section with an anterior cleft and a posterior notched point providing interlocking articulation with adjacent central teeth. Bases of the mid lateral teeth interlock posteriorly and anteriorly. The mesocones of the inner 5 – 7 mid laterals frequently wear a hole in the back of the tooth in front (Jenkins, 1984: pl. 2 i). Outer laterals do not interlock between transverse rows. The spaces between rows increases to the ribbon edges coupled with a gradual decrease in tooth size. The mid lateral teeth are broad based and elongated without flanking ectocones (outer side denticles). The outer lateral teeth have both ecto and endo cones and a squared flat mesocone. The mid lateral’s mesocone can be either pointed or bicuspidate. Of the 3 radulae examined, 2 – 4 of the inner (mean = 2.66, SD = 1.2) and invariably 2 of the outer mid laterals were bicuspidate, while the central 3 – 7 mid laterals (mean = 5.66, SD = 2.3) were pointed (Jenkins, 1984: pl. 2 h, i).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF1A8297FCCAFA62FD61FF35.taxon	diagnosis	Comparative remarks. In our molecular phylogeny, S. jeanae (lateralis group, unit 75) represents a well-differentiated lineage (Figs 1, 4). It is the sister species of a clade formed by S. australis, S. propria and S. diemenensis. Siphonaria jeanae differs from other species by COI distances of ≥ 16.6 % (Table S 8). Throughout its range, S. jeanae has been found in sympatry with seven congeners. Two congeners are sympatric in SE to SW Australia: For comparisons with S. stowae and S. tasmanica refer to comparative remarks under these species. Three species are sympatric in SE Australia: For comparisons with S. funiculata, S. diemenensis, and S. zelandica refer to comparative remarks under these species. One species is sympatric in western WA: Siphonaria restis sp. nov. has a lower, paler off-white coloured shell with greater uneven ribbing and scalloped edge, a paler, golden-brown spatula, a larger AO and BC, and longer ED and F 1. RS (Fig. 53 A) and SPM (Fig. 53 B) depicted here correspond well with the original description of Jenkins (1984).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF1A8297FCCAFA62FD61FF35.taxon	distribution	Distribution and habitat. Endemic to southern coasts of Australia, from Cape Conran Vic, near Hobart, Tas, SA and to Kalbarri, WA (Fig. 51). In this study, commonly found in exposed and sheltered positions (in crevices, holes) on moderately exposed to exposed intertidal marine rocky shores, upper and mid littoral levels (Fig. 52 R).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF168297FF68FE82FBDDFEB5.taxon	description	(Figs 52 E – F, T)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF168297FF68FE82FBDDFEB5.taxon	materials_examined	Material examined. Type material. Holotype of P. emergens from North Fingal Bay, Port Stephens, NSW, 32 ° 44.750 ’ S, 152 ° 10.500 ’ E, on beach in shell sand; coll. J. Voorwinde, 1950 - 60 (AM C. 532859), two paratypes, same data as holotype (AM C. 532860 2 d). Other, non-type material. Australia, NSW: Off Port Stephens, 32 ° 42 ′ S, 152 ° 15 ′ E (AM C. 265971 2 d); North Fingal Bay, Port Stephens, 32 ° 44.750 ′ S, 152 ° 10.500 ′ E (AM C. 265919 20 + d); Shelly Beach, Bateau Bay, S of The Entrance, 33 ° 22 ′ S, 151 ° 29 ′ E (AM C. 265912 d); Blue Lagoon Beach, S of Tuggerah Lakes, 33 ° 21 ′ S, 15,1 ° 30 ′ E (AM C. 265924 d); Long Reef near Collaroy, 33 ° 44.498 ′ S, 151 ° 19.117 ′ E (AM C. 265920 d); Collaroy Beach, 33 ° 43.700 ′ S, 151 ° 18.0 ′ E (AM C. 532861 d); Manly Beach, 33 ° 47.817 ′ S, 151 ° 17.368 ′ E (AM C. 265913 d); Middle Harbour, 33 ° 49.088 ′ S, 151 ° 15.427 ′ E (AM C. 265921 2 d); off Chinamans Beach, 33 ° 48.870 ′ S, 151 ° 14.965 ′ E (AM C. 265928 d); Bronte, 33 ° 54.355 ′ S, 151 ° 16.252 ′ E (AM C. 265925 9 d); Little Coogee Bay, 33 ° 55.300 ′ S, 151 ° 15.600 ′ E (AM C. 265923 d, C. 265914 2 d, C. 265915 d, C. 265922 4 d, C. 265926 d); 2 km E of Mistral Point, 33 ° 56.700 ′ S, 151 ° 16.700 ′ E (AM C. 265916 d); Cronulla, 34 ° 2 ′ S, 151 ° 10 ′ E (AM C. 265931 4 d, C. 265929 d).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF168297FF68FE82FBDDFEB5.taxon	description	Shell (Figs 52 E – F, T; Table S 9). Small sized (max sl <5 mm); height medium; weakly elongate; apex well offset to posterior, aligning with or beyond shell edge (Jenkins 2018: figs 5 A, G, J), to left of shell centre line; protoconch direction homostrophic (n = 1; Fig. 52 T) below apex and shell height (Jenkins 2018: figs 5 C, F, I, L); anterior and lateral apical sides convex, posterior concave almost notch-like; shell thin glassy translucent; dorsally arched with prominent curvature in shell-growth, growth striae irregular prominent; ribs radiate from apex to lip, indistinct, flat unraised; siphonal ribs not prominent on exterior, exterior coloration tan / red-brown with up to ten irregular white apical rib interstice bands extending and widening over lower half of sides to shell edge; lip fragile, undulated; spatula small, glossy, tan / red-brown, interior coloration reflects exterior coloration through translucent shell (Jenkins 2018: figs 5 B, E, K), ADM scar horseshoe-shaped, weakly indented and shallow; cephalic ADM scar straight to weakly convex (Jenkins 2018: fig. 5 E, H) between rounded left and right ADM scars, siphonal groove indistinct, very weakly indented.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF168297FF68FE82FBDDFEB5.taxon	diagnosis	Comparative remarks. Siphonaria emergens (atra group) has been found in sympatry with six congeners in Sydney Harbour, NSW: For comparisons with S. stowae, S. denticulata, S. diemenensis, S. funiculata, S. scabra, and S. zelandica refer to comparative remarks under these species. This species in known only from shells and its anatomy remains undocumented. Consequently, no DNA sequences have been available for genetic study.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF168297FF68FE82FBDDFEB5.taxon	distribution	Distribution and habitat. Incompletely known, yet to be live collected. Endemic to the, warm temperate central coast of NSW between Port Stephens and Cronulla Sydney, Australia (Fig. 51).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF168290FCCAFE02FB1DFA35.taxon	description	(Figs 52 J – L, O – P)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF168290FCCAFE02FB1DFA35.taxon	materials_examined	Material examined. Type material. Holotype of S. oblia from Hospital Point, Thursday Island, Torres Strait, Qld, 10 ° 35.318 ’ S 142 ° 12.573 ’ E; coll. Ponder and Loch, 29 Jun. 1976, on moderately exposed rocky reef (AM C. 125603, figured in Jenkins 2018: fig. 6), 15 paratypes, same data as holotype (AM C. 125604 11 p, 4 d). Other, non-type material. Australia, Qld: Torres Strait: N end Prince of Wales Island, beach opposite Hospital Point, Thursday Island, 10 ° 35.748 ′ S, 142 ° 12.280 ′ E (AM C. 265974 8 d); S side of Thursday Island, 10 ° 35.133 ′ S, 142 ° 13.257 ′ E (AM C. 532862 8 p); Wednesday Island, 10 ° 31.0 ′ S, 142 ° 17.0 ′ E (AM C. 265909 20 + d); Bampfield Point, W side of Prince of Wales Island, 10 ° 43.043 ′ S, 142 ° 6.833 ′ E (AM C. 532863 9 p; AM C. 532864 10 + p); Cape York Peninsula: Albany Passage, 10 ° 44.278 ′ S, 142 ° 35.758 ′ E (AM C. 265907 d); Mutee Head, 10 ° 54.682 ’ S, 142 ° 15.204 ’ E, Q 50 - 2 (AM C. 585755 p [M 105]); Somerset, S side of Fly Point, 10 ° 45.195 ′ S, 142 ° 36.292 ′ E, Q 47 - 1 (AM C. 265910 8 p, 6 d; C. 584902 p [M 042]); Captain Billy Landing, 11 ° 38.019 ′ S, 142 ° 51.472 ′ E, Q 46 - 1 (AM C. 559470 2 p); Portland Road, 12 ° 35.588 ′ S, 143 ° 24.660 ′ E, Q 45 - 1 (AM C. 559471 p); Point S of Bathurst Head, 14 ° 17.583 ′ S, 144 ° 11.845 ′ E, Q 41 - 1 (AM C. 559473 10 p); Lizard Island, Casuarina Beach, 14 ° 40.447 ′ S, 145 ° 26.703 ′ E, Q 40 - 1 (AM C. 266052 d), 14 ° 40.908 ′ S, 145 ° 27.007 ′ E (AM C. 559473 3 p); Point Archer, 15 ° 35.558 ′ S, 145 ° 19.788 ′ E (AM C. 49513 5 d); Cape Kimberley, 16 ° 16.535 ’ S, 145 ° 28.737 ’ E, Q 35 - 1 (AM C. 585478 17 p, C. 585172 p [M 076], C. 585505 p [SK 392]); Port Douglas, 4 Mile Beach, 16 ° 30.470 ′ S, 145 ° 28.168 ′ E (AM C. 265953 d), 16 ° 29.468 ′ S, 145 ° 28.425 ′ E (AM C. 265908 2 d); Pebbly Beach, Yule Reef, Trinity Bay, 16 ° 35.031 ’ S, 145 ° 30.823 ’ E, Q 32 - 2 (AM C. 608188 2 p); Red Cliff Pt N of Cairns 16 ° 41.294 ’ S, 145 ° 35.080 ’ E, Q 33 - 3 N of Cairns, Buchan Point, 16 ° 44.172 ′ S, 145 ° 39.973 ′ E (AM C. 265951 7 d).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF168290FCCAFE02FB1DFA35.taxon	discussion	Taxonomic remarks. Our delineation of this species is based on comparative analyses of the morpho-anatomy and mitochondrial genetics of freshly collected topotypes (Figs 52 J – L) and a geographic series of additional specimens (Table S 1).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF168290FCCAFE02FB1DFA35.taxon	description	External morphology (Fig. 52 O). Animal tissue translucent; foot sole and foot wall evenly pale grey to cream; mantle thin, translucent with large whitish subepithelial pustules in thickened fringe, positioned along top edge of foot wall, forming a broad flat band, radial markings on mantle aligned with shell colour bands; foot wall narrow; faint brown / red splotches present on lower foot wall around body but not extending under mantle; pneumostomal lobe on right side narrow translucent with pointed lobe, covering inconspicuous anus in foot wall; brown markings on posterior lobe of pneumostomal lobe; two black ‘ Eye’ spots prominent centrally on thickened cephalic lobes, centrally touching and marked with brown / reddish splotches; genital pore inconspicuous, positioned in foot wall posterior to right cephalic fold. Shell (Figs 52 J – L; Table S 9). Small sized (max sl <5 mm), ovate, height medium; centrally broad, thickness thin, maybe glassy translucent; apex well offset to posterior edge and left of centre line, apex below shell height, dorsally dark tan / brown in colour; offset growth reflected in apical ridge; protoconch direction weakly heterostrophic (n = 1; Jenkins 2018: 282, fig. 6 J), shell whorl dextral; anterior and lateral apical sides convex, posterior weakly concave; apex ribs radiate from apex to shell lip, flat unraised, appear as bands, tan in colour, rib count (mean = 35, SD = 3.7, n = 32); siphonal ribs not prominent on exterior, apical cream banding of rib interstices prominent, colour bands widen slightly over lower half of sides to lip; number of apical bands varies within individuals, lip fragile, unscalloped, even; growth striae irregular, prominent; light periostracum covers lower parts of exterior sides, often freely extending; spatula glossy, brown, interior colouration reflects exterior colouration through translucent shell; ADM scar horseshoe-shaped, weakly indented shallow; cephalic ADM scar straight to weakly convex, between left and right rounded ADM scars; siphonal groove very weakly indented. Reproductive system (Jenkins 2018: 278, fig. 3 C – D). Located in posterior region of coelom and partly covered by pallial cavity and digestive gland; HG granulated; MG and AG folded; a thickened weakly coiled brown-coated HD emerges from HG; small rounded SV enveloped within AG folds; CD emerges from base of AG folds, progressively decreasing in diameter anteriorly and narrowing to open in rear of GA, AO indistinct; bulbous EG and narrow elongate ED are located between BM and RAM; ED slightly coiled prior to entering GA close to single GP; what appear to be two prominent flagellum both close to join of ED and EG, one broad and blunt, other narrow and elongate (Jenkins 2018: fig. 3 B – C); BC pale brown bulbous with a thin translucent test, located beside posterior RAM; BC may appear deflated when without SPM (Jenkins 2018: fig. 3 B); elongate thickened BD leads from BC and, coupled with anterior section of CD, passes through RAM; both ducts enter separately into GA and opposite to entry of ED. Spermatophore (Jenkins 2018: fig. 4 E). Over half length comprises a translucent cylindrical head section, tip bluntly rounded; flagellum very thin, transparent, tapering to a thread-like end; both sections smooth, featureless (length = 0.49 mm; head length = 0.19 mm, ~ 38 % of SPM length, head width = 17 μm, flagellum width = 4 μm, n = 1). Two SPMs coiled, embedded in a white gelatinous mass in BC of one paratype (AM C. 125604). Radula (Jenkins 2018: 284, fig. 3 E – G, 4 E, F). Each half row has around six mid and nine outer lateral teeth; central tooth with single pointed mesocone, about one-third base length; inner laterals (i. e., without ecto / endocones) absent; first mid lateral has prominent bicuspidate pointed mesocone, other mid laterals possess single cusped elongate mesocone, single cusped ectocones offset from side of mesocone; central tooth basal plate as long as adjacent mid-lateral teeth; mid-lateral teeth possess a single outward pointing forked prong and notch, rows interlock via these basal forks and notches; outer lateral teeth distinctly different in shape to mid laterals, teeth block-like with single cusped chisel-like mesocone, all possess a pointed endocone; starting around laterals 8 – 9 the number of ectocones is doubled on outer teeth, endocones and ectocones appear as separate cones, are variably-shaped, bluntly pointed, some almost as large as the mesocone. Dentition formula 15: 1: 15 + / − 1 (n = 3; row count was not assessed). Jaw (Jenkins 2018: 283, fig. 4 H). Located inside front of buccal cavity, orange-brown, arch shaped with unevenly ‘ shingle’ - arranged cone-like rods, ~ 80 rods wide (~ 0.8 mm) by ~ 9 – 10 rods deep (length = 27 μm, width = 8 μm, n = 12); tip bluntly rounded.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF168290FCCAFE02FB1DFA35.taxon	diagnosis	Comparative remarks. Genetically S. oblia (normalis group, unit 77) is a well-differentiated lineage (Figs 1, 4). It differs from other species by COI distances of ≥ 15.6 % (Table S 8). In N Qld, this species has been found in sympatry with five congeners: For comparisons with S. viridisS. atra, and S. normalis refer to comparative remarks under these species. Siphonaria jiigurruensis sp. nov. has a larger, taller, paler shell with greater edge scalloping, a larger AO, longer ED, and a smaller BC. The combined shell geometry, size, and colouration of S. oblia is highly distinctive.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF168290FCCAFE02FB1DFA35.taxon	distribution	Distribution and habitat. Endemic to tropical NE coast of Australia, Qld, from Torres Strait to just N of Cairns (Fig. 51). In this study, found on moderately exposed rocky intertidal marine shores in sheltered positions (e. g., on under-surface of rocks, amongst oysters, amongst fringes of coralline algae and in crevices), upper and mid littoral levels (Fig. 52 P).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF118291FCCAF982FA08FD56.taxon	description	(Figs 53 C – D, 54 A – C, M – N)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF118291FCCAF982FA08FD56.taxon	materials_examined	Material examined. Type material. Holotype, from Dolokoan Beach 8 ° 31.424 ’ S, 125 ° 37.091 ’ E, N of Dili, Timor-Leste; coll. B. W. Jenkins, TL 01 - 1, 14 July 2019 (AM C. 584823 [M 447, SK 230 (RS)], Fig. 54 A). Three paratypes, same data as holotype (AM C. 585353 p [SK 265], AM C. 585354 p [SK 266], Fig. 54 B; AM C. 585355 p [SK 267], Fig. 54 C).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF118291FCCAF982FA08FD56.taxon	description	External morphology (Fig. 54 N). Foot sole evenly grey; foot edge and cephalic folds cream, slightly lobed; foot wall, mantle and pneumostomal lobe evenly shaded dark grey, lighter to foot edge; mantle narrower than foot wall, translucent, elongated at anterior, edge thickened, lobed with grey / black banding (over mantle width) aligning with rib interstices and interstice width; pneumostome elongated. Shell (Figs 54 A – C; Table S 9). Small sized (max sl mean = 8.1 mm, SD = 0.6 mm, n = 4), ovate; height low to medium; apex offset to posterior and slightly left; apical sides convex, concave and elongated to posterior; protoconch direction homostrophic (n = 1), shell whorl dextral; growth striae prominent uneven, shell thickness thin; shell edge uneven; rib count (mean = 28.3, SD = 2.3, n = 4), exterior with dual shaded bands, paler apical (white ribs and pale brown interstices) and darker shell edge (white ribs and black / dark brown interstices); primary ribs white, crooked, broaden to shell lip, weakly protrude beyond shell lip to unevenly scallop and corrugate the edge, 0 – 1 interspersed pale white finer secondary ribs; loosely paired primary ribs form siphonal ridge, no more prominent than other primary ribs; interior shell margin very dark brown to black, white rays on shell margin to lip align under primary / secondary ribs, siphonal groove indistinct, spatula very dark chocolate to black with some underlying white; ADM scar distinct, CMS straight; thickening of shell lip not apparent. Reproductive system (Fig. 53 C; n = 2). Positioned within entire right side of coelom, against foot wall on foot muscle, under the respiratory cavity occupying large proportion of animal body volume; epiphallic parts positioned over BM. GA small, with singular GP through foot wall; AO very small, narrow, bluntly pointed, joined to lower ED and upper GA; ED very short, very broad, centrally bent, joins to side of GA; GA, AO, ED all white muscular fibrous tissue; EG very large, soft whitish tissue, slightly folded, joins ED; extension joins in parallel to single very broad flagellum (F 1), similar width to ED, appears as an extension of ED; BD and CD connect side-by-side into GA between ED join and GP, both ducts short, slightly bent, smooth, thickened, whitish, featureless, pass closely together inside outer RAM (BD over CD) into soft white folded tissues of MG; MG / AG complex medium; CD connecting to ducts, BC embedded in folds close to embedded purplish SV; BD without distal loop and MA, with loop immediately prior to BC; BC relatively large, spherical, thin whitish translucent test; HD short, narrow, coiled, links ducts in soft white folded tissues of AG to yellowish granulated HG; outer edge of MG lobbed; AG larger than HG, outer sides match curvature of inner foot wall. Spermatophore (Fig. 53 D). Broad head with short flagellum (length = 2.38 mm, n = 1); head section cylindrical, bulbous, centrally twisted, rounded tip; test thin, smooth, featureless, translucent encasing a white opaque central core; short tapering section merges head to filamentous flagellum; head shorter, wider than translucent flagellum (head length = 0.95 mm, ~ 40 % of SPM length, head width = 103 μm; flagellum width = 17 μm, n = 1); 1 SPM found coiled in two BCs (Fig. 53 D).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF118291FCCAF982FA08FD56.taxon	diagnosis	Comparative remarks. Siphonaria campestra sp. nov. (normalis group, unit 82) is most closely related to S. normalis, S. madangensis, S. fuliginata, and S. costellata, with which it forms a sub-clade in the normalis group (Figs 1, 4). However, it is well-differentiated from other species by COI distances of ≥ 18 % (Table S 2). This species has been found in sympatry with five congeners in TL: For comparisons with S. alba, S. javanica, and S. viridis refer to comparative remarks under these species. Siphonaria forticosta sp. nov. has a larger, darker shell with a more distinct siphonal ridge, a larger BC and narrower ED. Siphonaria planucosta sp. nov. has a larger shell with more raised ribbing, a less prominent siphonal ridge, a narrower AO, BD with distal loop, and a larger BC.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF118291FCCAF982FA08FD56.taxon	distribution	Distribution and habitat. Recorded from Dolokoan Beach, Timor-Leste (Fig. 51). In this study, found in sheltered positions (mainly crevices) on moderately exposed rocky shores, upper and lower littoral levels (Fig. 54 M).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF118291FCCAF982FA08FD56.taxon	etymology	Etymology. From ‘ campestris’ (Latin = level, even), referring to the level or even primary ribs on the shell.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF108293FCCAFD62FBF7FED6.taxon	description	(Figs 53 E – F, 54 D – E, O – P)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF108293FCCAFD62FBF7FED6.taxon	materials_examined	Material examined. Type material. Holotype, from Tancha Bay, 26 ° 27.897 ’ N, 127 ° 49.131 ’ E, Okinawa, Japan; coll. B. W. Jenkins, JP 01 - 5, 20 March 2020 (AM C. 585614 [M 491, SK 310]). Paratype, same data as holotype (AM C. 585195 p [SK 331]); paratype from Tancha Bay point, 26 ° 27.941 ’ N, 127 ° 49.194 ’ E, Okinawa, Japan; coll. B. W. Jenkins, JP 01 - 6, 20 March 2020 (AM C. 584920 p [M 490, SK 309]); 5 paratypes from Moon Bay, Onna, Okinawa; coll. B. W. Jenkins, JP 01 - 4, 18 March, 2020 (AM C. 585621 5 p). Other, non-type material. Japan, Okinawa: Sun Marina Beach, seawall, 26 ° 27.842 ’ N, 127 ° 48.755 ’ E, JP 01 - 1 (AM C. 585617 8 p); Tancha Bay 1, 26 ° 27.897 ’ N, 127 ° 49.131 ’ E, JP 01 - 5 (AM C. 585951 5 p, C. 585195 p [SK 331]); Tancha Bay 2, 26 ° 27.941 ’ N, 127 ° 49.194 ’ E, JP 01 - 6 (AM C. 585950, 4 p, C. 585613 p [SK 510]). China, Hong Kong: Repulse Bay 22 ° 14.1 ’ N, 114 ° 11.71 ’ E (ZRC. MOL. 24899 p).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF108293FCCAFD62FBF7FED6.taxon	description	External morphology (Fig. 54 P). Foot sole dark yellowish / grey, paler to foot edge; foot wall, mantle, cephalic folds and pneumostomal lobe all evenly dark yellowish / green in colour; mantle thin, translucent, narrower than foot wall, weakly lobed with a thickened yellow banded edge; faint irregular black blotches of pigmentation on foot wall and cephalic lobes; pneumostome fold long between right ADMs and within mantle. Shell (Figs 54 D – E; Table S 9). Small sized (max sl mean = 11.6 mm, SD = 0.9 mm, n = 8), ovate, height tall; shell thin, apex offset strongly posterior and central; apical sides convex, strongly concave at posterior; protoconch direction homostrophic (n = 2; Fig. 54 P), peaked, hooked to posterior, shell whorl dextral; growth striae uneven; rib count (mean = 35, SD = 0.7, n = 8), exterior unevenly brown, weak radial banding, protoconch area dark brown, paler to shell edge; ribs uneven, increasingly broader and align to uneven fragile and corrugated shell edge, periostracum freely extending; rib interstices darker; siphonal ridge prominent formed by paired primary ribs. Interior shell margin and spatula evenly dark chocolate brown, white rays on shell lip align under rib ends, fade to margin; siphonal groove distinct, same colour as shell margin; ADM scar distinct, CMS straight, indistinct; thickening of shell lip not observed. Reproductive system (Fig. 53 E; n = 3): positioned within coelom under the respiratory cavity, hermaphroditic glands positioned to posterior against right foot wall and over foot sole, epiphallic parts positioned over BM, BC directly lodged in soft white folds of MG, BD not through RAM as with CD (n = 2, Fig. 53 E); AO small, elongated, narrow, bluntly pointed; ED broad, elongated, centrally twisted, longer than BD; ED (with outer MA) and AO each enter into top of small GA; EG relatively large with folds, often elongated; single, short, broad flagellum F 1 with curled end, appears as a right-angled protrusion from end of ED at join with EG; AO, GA and ED all muscular white tissue; BD and CD join GA with swollen opposing connections close to entry of ED and AO; CD passes between inner foot wall and outside RAM connecting into large MG; BD without distal loop, longer and narrower than CD, enters MA alongside or behind ED, both ducts broad smooth; BC large, club shaped, translucent test; HD large, coiled, links soft white folds of AG to similar sized brownish finely granulated HG; SV embedded within AG, outer sides of AG and HG match curvature of inner foot wall at right posterior quarter of coelom. Spermatophore (Fig. 53 F). Body cylindrical, thread-like (length = 10.9 ± 3.0 mm, n = 2), test thin, translucent, smooth; head tip bluntly rounded, section containing a white gelatinous core tapers to a thinner flagellum; head section longer thicker than flagellum (head length = 6.0 ± 0.96 mm, n = 2; ~ 54 % of SPM length, flagellum length = 4.8 ± 2.1 mm, head width = 120 ± 0 μm, flagellum width = 50 ± 0 μm, n = 2), outer ridge of mid flagellum possesses 35 - 17 + evenly spaced barbs pointing towards head; 2 SPM tightly coiled in white gelatinous mass in each BC of two specimens (AM C. 584920, AM C. 585614).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF108293FCCAFD62FBF7FED6.taxon	diagnosis	Comparative remarks. In our mitochondrial phylogeny, S. camura sp. nov. (unit 84) is the sister species of S. japonica (unit 2); both species together forming a distinctive subclade in the Siphonaria tree (Figs 1, 4). Both species differ form each other by COI distances of ≥ 11 % (Table S 2). Throughout its range, this species has been found in sympatry with five congeners: For comparisons with S. japonica, S. subatra, S. sipho, and S. rucuana refer to comparative remarks under these species. Siphonaria tanchaensis sp. nov. has a larger, taller, paler shell with greater edge scalloping, a white / brown interior and golden-brown spatula, a larger AO, longer narrower BD distal loop, smaller BC, and a longer F 1.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF108293FCCAFD62FBF7FED6.taxon	distribution	Distribution and habitat. Recorded from Okinawa, Japan and Repulse Bay, Hong Kong (Fig. 55). In this study, found on exposed rocky shores in sheltered positions, such as rock crevices, hollows and amongst oysters, mid to upper littoral levels (Fig. 54 O).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF108293FCCAFD62FBF7FED6.taxon	etymology	Etymology. From ‘ camur’ (Latin = curved, bent), referring to the slightly curved, upturned end of the siphonal ridge at the shell lip.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF12828DFCCAFEE2FE8CFB96.taxon	description	(Figs 53 G – H, 54 F – G)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF12828DFCCAFEE2FE8CFB96.taxon	materials_examined	Material examined. Type material. Holotype, from Caubian Island, 10 ° 17.22 ’ N, 124 ° 10.53 ’ E, Bohol, Philippines (AM C. 595933, Fig. 53 F). Paratypes. Same data as holotype (AM C. 595934 6 d, Fig. 53 G). Other, non-type material. Philippines: Caubian Island, Bohol, 10 ° 17.22 ’ N, 124 ° 10.53 ’ E (AM C. 595932 20 + d, C. 595935 p [SK 559]); NW Polillo Is, E Quezon, Bolunga District, nr Panukalan, 14 ° 59 ’ N, 121 ° 49 ’ E (WAM S 72342 10 p, WAM S 113803 p [SK 557]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF12828DFCCAFEE2FE8CFB96.taxon	description	External morphology (preserved). Foot sole and foot wall evenly cream, paler to foot edge; mantle pale, translucent, edge thick and lobed band; irregular widely spaced dark pigmentation spots on foot wall, concentrated over posterior and faintly over cephalic lobes. Shell (Figs 54 F – G; Table S 9). Medium sized (max sl mean = 20.6 mm, SD = 2.1 mm, n = 20), circular ovate; height medium; apex offset weakly left and posterior, apical sides convex; protoconch direction homostrophic (n = 3), shell whorl dextral; exterior uneven, dark brown, weak radial colour banding; rib count (mean = 30, SD = 3.1, n = 20), ~ 14 – 16 primary ribs, straight to slightly bent towards shell edge, primary ribs white or dark brown, number (0 – 14) and position on shell extremely variable, ridges raised, rounded, broaden slightly and increasingly raised to shell edge, variably and strongly protrude beyond shell lip (often> 3 mm) to strongly and unevenly scallop shell edge; 0 – 4 finer secondary ribs between primary ribs, rib interstices darker; siphonal ridge formed by closely paired, often end flared, primary ribs. Interior shell dark chocolate brown, spatula white to mottled dark brown; white rays on shell margin aligned in furrows under primary ribs, may extend to spatula, siphonal groove prominent, pale white to dark chocolate brown; ADM scar indistinct, darker than margin and shell edge, CMS straight; thickening and whitening of shell lip not observed. Reproductive system (Fig. 53 G; n = 1). Positioned within right side of coelom, against foot wall on foot muscle, under the respiratory cavity; epiphallic parts positioned over BM; GA small, with singular GP through foot wall; AO large, broad, blunt, joined to upper GA; ED long, wide, centrally twisted, joins to side of AO and GA; GA, AO, ED all white muscular fibrous tissue; EG large, soft whitish tissue, folded, joins ED with single long narrow bent flagellum (F 1), appears as a narrower extension of ED; BD and CD connect juxtapose into GA / AO, opposing ED join, both ducts long, narrow, slightly bent, whitish, pass closely together through outer side of RAM (smooth featureless BD over wrinkled CD) into soft white folded tissues of MG; MG / AG complex large; CD connecting to ducts, BC embedded in folds close to embedded SV; BD with bursal loop; BC relatively small, elongated, thin whitish translucent test; HD long, broad, coiled, links ducts in soft white folded tissues of AG to granulated HG; outer edge of MG unlobed; AG larger than HG, sides match curvature of inner foot wall. Spermatophore (Fig. 53 H). Thread-like (length = 11.1 mm, 74 % of AL, n = 1); translucent, test thin; head section bluntly rounded, cylindrical, containing a prominent coiled white core, tapers to a thin tip; both sections smooth, featureless. Head section longer wider than flagellum (head length = 9.7 mm, mean ~ 88 % of SPM length, head width = 155 μm, flagellum width = 34 μm); 1 SPM tightly folded in BC (WAM S 113803).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF12828DFCCAFEE2FE8CFB96.taxon	diagnosis	Comparative remarks. We found this species in sympatry with four congeners on Cebu, Philippines. For comparisons with S. sirius, S. bifurcata, S. alba, and S. sipho refer to comparative remarks under these species. We have not been able to sequence material of this species.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF12828DFCCAFEE2FE8CFB96.taxon	distribution	Distribution and habitat. Recorded from Cebu and Bohol Islands, Philippines (Fig. 55). Found on rocky intertidal shores.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF12828DFCCAFEE2FE8CFB96.taxon	etymology	Etymology. For the type locality, Caubian Island, Philippines.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF0C828EFF68FB22FEA4FBF6.taxon	description	(Figs 53 I – J, 54 H – J, Q – R)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF0C828EFF68FB22FEA4FBF6.taxon	materials_examined	Material examined. Type material. Holotype, from E side Smith Point, Flying Fish Cove, 10 ° 25.749 ’ S, 105 ° 39.957 ’ E, CI; coll. B. W. Jenkins, CI 01 - 1 / 2, 11 Sept 2018 (AM C. 595957 [M 298], Fig. 54 H), 23 paratypes, same data as holotype (WAM S 74040 p [SK 069], Fig. 54 I, WAM S 74075 9 p; AM C. 585992 9 p, AM C. 584837 p [M 299], AM C. 584838 p [M 300], AM C. 584839 p [M 301]); Ethel Beach, 10 ° 27.827 ’ S, 105 ° 42.497 ’ E; coll. B. W. Jenkins, CI 02 - 1, 11 Sept 2018 (AM C. 584846 p [M 308], Fig. 54 J). Other, non-type material. Australia, CI: Off Dale 3, 10 ° 28.52 ’ S 105 ° 33.503 ’ E (AM C. 595955 p [SK 566]); E side Smith Point Flying Fish Cove, 10 ° 25.749 ’ S, 105 ° 39.957 ’ E CI 01 - 1 (AM C. 584836 p [SK 021 protoconch A 11], C. 585319 p [SK 071 protoconch C 1], C. 585321 p [SK 083], C. 585954 p [SK 568]), CI 01 - 2 (AM C. 584729 10 p); West White Beach, 10 ° 27.748 ’ S, 105 ° 34.934 ’ E CI 01 - 3 (AM C. 585922 p [SK 567]); Ethel Beach, 10 ° 27.827 ’ S, 105 ° 42.497 ’ E CI 02 - 1 (AM C. 585695 8 p, C. 584846 p [M 308]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF0C828EFF68FB22FEA4FBF6.taxon	discussion	Taxonomic remarks. This species has previously been identified as ‘ Siphonaria sp. 1 ’ by Wells & Slack-Smith (2000: 113).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF0C828EFF68FB22FEA4FBF6.taxon	description	External morphology (Fig. 54 R). Foot sole grey, foot wall grey to yellowish with black flecked pigmentation; edge of foot sole and foot wall yellowish to white, fading to grey at mantle join; mantle lobed, translucent, narrower than foot wall, patches of black pigmentation occur on lobes, aligning with dark shell rib interstices; cephalic folds grey with black flecks concentrating around lobe join; foot wall and pneumostome pustulose, pneumostome yellowish, wide, within mantle, between right ADMs. Shell (Figs 54 H – I, Q; Table S 9). Small (max sl mean = 10.1 mm, SD = 1.4 mm, n = 11); height low to medium; ovate; apex offset posterior and left, apex cap brown, often eroded to appear white; apical sides convex, posterior concave; protoconch direction central to weakly homostrophic (n = 2; Fig. 54 Q), shell whorl dextral; growth lines distinct, undulating; rib count (mean = 30.6, SD = 6.3, n = 11), 20 – 30 primary ribs prominent, often very broad, weakly raised, whitish, straight to weakly bent, width increases strongly to shell lip often touching to replace rib interstices and any infill of 0 – 1 secondary ribs, extend slightly beyond shell edge creating corrugations and weak scalloping in the uneven shell lip; rib interstices black to dark brown, siphonal ridge slightly raised, indicated by abutting paired primary ribs; segments either side of siphonal ridge often clear of primary ribs, black or brown coloured, show finer secondary ribs; interior shell margin yellowish, white under primary ribs, dark black / brown markings under rib interstices span from shell lip to spatula across the shell margin; spatula brown to dark brown; ADM scar impression distinct, same as shell margin colouring, cephalic scar convex; thickening of shell lip commonly occurs, translucent, not covering black / brown markings of rib interstices, CMS curved. Reproductive system (Fig. 53 I; n = 2). Positioned within coelom under the respiratory cavity, hermaphroditic glands positioned to posterior against right foot wall and over foot sole, epiphallic parts positioned to anterior between BM and RAM; GA small with singular GP through foot wall; AO large elongated centrally twisted, tapers to point, joined to upper GA alongside ED; ED long broad slightly centrally bent, joins to side of GA; GA, AO, ED all white muscular fibrous tissue; EG large pointed soft whitish tissue, slightly folded, joins ED; single short flagellum (F 1) on side. BD and CD join side-by-side into GA between AO join and GP; CD short curved smooth thickened whitish featureless; BD long, narrower than CD, with distal loop and prominent MA; both pass closely together through RAM (BD over CD) into soft white folded tissues of MG; MG / AG complex relatively large; BC embedded in folds of AG / MG close to embedded SV; BD similar thickness to CD; BC small bulbous thin whitish translucent test; HD short narrow coiled, links ducts in soft white folded tissues of AG to yellowish granulated HG; AG larger than HG, both outer sides curved to curvature of inner foot wall. Spermatophore (Fig. 53 J). Test thin, translucent (length = 11.27 mm, n = 1, AL = 7.21 mm; possibly longer as flagellum appeared incomplete); head bulbous, tip bluntly rounded, containing a white gelatinous mass; very short taper region into the filamentous transparent flagellum; both sections smooth, featureless; head shorter and much thicker than flagellum (head length = 4.94 mm, ~ 44 % of SPM length, head width = 126 μm, flagellum width = 32 μm); single SPM found in one BC (AM C. 585321).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF0C828EFF68FB22FEA4FBF6.taxon	diagnosis	Comparative remarks. In our mitochondrial phylogeny (Figs 1, 3), S. christmasensis sp. nov. (plicata group, unit 63) is the sister species of S. delicata sp. nov. (unit 62). Both species occur on CI forming a well-differentiated lineage. They differ from each other by COI distances of ≥ 10.7 % (Table S 7). We found this species in sympatry with five congeners on CI: For comparisons with S. alba, and S. incerta refer to comparative remarks under these species. Siphonaria tenebrae sp. nov. has a slightly larger, lower, darker shell with greater edge scalloping, a single primary rib forming the siphonal ridge, darker interior, a more pointed AO and a longer F 1. Siphonaria umbra sp. nov. has a slightly larger, lower and darker shell, a larger AO and BC, and a longer F 1. Siphonaria delicata sp. nov. has a smaller, taller, darker shell with less prominent ribs, darker interior, a larger AO and BC, and longer ED and F 1.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF0C828EFF68FB22FEA4FBF6.taxon	distribution	Distribution and habitat. Known only from CI, Indian Ocean (Fig. 55). In this study, found in sheltered positions on moderately exposed and exposed limestone rocky shores, upper and mid littoral levels (Fig. 54 R).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF0C828EFF68FB22FEA4FBF6.taxon	etymology	Etymology. For the type locality, Christmas Island, Indian Ocean.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF0F828FFF68FBC2FDFCFA76.taxon	description	(Figs 54 K – L, S – T, 58 A – B)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF0F828FFF68FBC2FDFCFA76.taxon	materials_examined	Material examined. Type material. Holotype, from Lazarus Island, causeway 01 ° 13.288 ’ N, 103 ° 51.195 ’ E, Singapore; coll. Chim C. K. and B. W. Jenkins, SI 04 - 3, 21 Nov 2018 (ZRC. MOL. 24890 [M 416, SK 100], Fig. 54 K). 18 paratypes, same data as holotype (AM C. 585234 p [M 326], AM C. 585427 10 p, AM C. 585677 6 p), paratype from Lazarus Island, 01 ° 13.355 ’ N, 103 ° 51.148 ’ E, Singapore; coll. B. W. Jenkins, SI 04 - 2, 24 Nov 2018 (AM C. 585226 p [M 338], Fig. 54 L). Other, non-type material. Australia, Qld: Weipa, 12 ° 37.795 ’ S, 141 ° 51.853 ’ E, Q 52 - 1 (AM C. 585186 p [M 075]; C. 585187 p [M 178]); Inspection Pt, Sweers Is, 17 ° 08.471 ’ S, 139 ° 36.868 ’ E, Q 56 - 3 (AM C. 585189 p [M 147]). NT: Nhulunbuy, Cape Wirawawoi, 12 ° 09.513 ’ S, 136 ° 46.904 ’ E, NT 05 - 1 A (AM C. 585529 20 + p, C. 585075 p [SK 044], C. 585076 p [M 028], C. 585077 p [M 092]); East Woody Islet, 12 ° 09.695 ’ S, 136 ° 45.075 ’ E, NT 05 - 2 (AM C. 585684, 7 p); N Turtle Beach, 12 ° 18.816 ’ S, 136 ° 55.930 ’ E, NT 04 - 1 (AM C. 585477 17 p; C. 585990 10 + p); Smith Pt 2, 11 ° 07.466 ’ S, 132 ° 08.538 ’ E, NT 21 - 3 (AM C. 585989 10 + p). Singapore: Pasir Ris, seawall, 01 ° 22.981 ’ N, 103 ° 56.956 ’ E, SI 01 - 1 (AM C. 585986 10 p); Fort Road, drain seawall, 01 ° 17.605 ’ N, 103 ° 53.809 ’ E, SI 01 - 3 (AM C. 585603 3 p); Lazarus Island, 01 ° 13.355 ’ N, 103 ° 51.148 ’ E, SI 04 - 2 (AM C. 585988 16 p); Lazarus Island causeway, 01 ° 13.288 ’ N, 103 ° 51.195 ’ E, SI 04 - 3 (AM C. 585226 p [M 338], C. 585233 p [SK 099], C. 585234 p [M 326], C. 585236 p [M 457, SK 183], C. 595973 p [M 327]); Lazarus Island channel headland, 01 ° 13.085 ’ N, 103 ° 51.429 ’ E, SI 04 - 4 (AM C. 585985 10 p).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF0F828FFF68FBC2FDFCFA76.taxon	description	External morphology. Foot sole, foot wall and cephalic folds evenly dark yellowish grey, paler at foot / wall edge; foot wall and mantle more yellowish; mantle lobed with small black pigmentation at thickened edge aligning with rib interstices; translucent, as wide as foot wall, covers exposed inner shell lip; genital pore inconspicuous, located on foot wall to right anterior of right cephalic fold; two small black epithelial spots centralised on two centrally touching unpigmented cephalic folds; pneumostomal lobe relatively long, under the mantle, unpigmented, between the right ADMs. Shell (Figs 54 K – L, S; Table S 9). Medium to large sized (max sl mean = 11.6 mm, SD = 1.7 mm, n = 7), ovate; medium to tall; apex offset central sightly posterior (usually eroded), apical sides strongly convex, protoconch direction heterostrophic (n = 2; Fig. 54 S), shell whorl dextral; growth striae prominent in bands, shell thickness thick; rib count (mean = 45.7, SD = 5.5, n = 7), primary ribs pale white, fairly straight, increasingly raised and protrude beyond shell lip to unevenly scallop and corrugate the edge; 1 – 2 interspersed pale white finer secondary ribs, rib interstices darker; paired primary ribs on siphonal ridge, no more prominent than other primary ribs. Interior shell margin dark brown to tan, white rays align on shell margin under primary / secondary ribs, siphonal groove distinct, same colour as shell edge, points to right anterior; spatula dark chocolate brown to mottled tan; ADM scar distinct, CMS straight, paler than shell lip; thickening of shell lip translucent, infills and reduces lip scalloping, spatula becomes whitened. Reproductive system (Fig. 58 A; n = 1). Positioned within right side of coelom, against foot wall on foot muscle, under the respiratory cavity; epiphallic parts positioned between RAM and BM. GA very small indistinct with singular GP through foot wall; AO small broad bluntly pointed, joined to GA; ED elongated narrow thickened, slightly centrally bent, joins to GA; GA, AO, ED all white muscular fibrous tissue; EG large, soft whitish tissue, folded, joins ED; single wide flagellum (F 1) lays over BM, shorter but similar width to ED, appears as an extension of ED. BD and CD connect side-by-side to GA between AO and GP, both ducts thick short bent smooth whitish, pass together through RAM (BD over thicker CD) into soft white folded tissues of MG, BC embedded in folds close to embedded blackish SV; BD short narrow bent before BC, without distal loop or MA to inner anterior foot wall; CD shorter wider than BD; BC relatively large bulbous, thin whitish translucent test (2 SPM in brownish gelatinous mass of BC); MG / AG complex relatively large; HD short narrow coiled, links ducts in soft white folded tissues of AG to yellowish granulated HG; AG / MG larger than HG, both outer sides curved to curvature of inner foot wall. Spermatophore (Fig. 58 B). Relatively short (length = 4.25 ± 0.196 mm, n = 2, flagellum incomplete), test thin, translucent, containing a white gelatinous core mass; over half-length comprises a translucent bulbous cylindrical head section (head length = 1.646 mm, SD = 0.213 mm, n = 2; mean ~ 39 % of SPM length, SD = 3 %); tip bluntly rounded, with a prominent twist before tapering into a filamentous transparent flagellum; head section much thicker than flagellum (head width = 116 ± 16 μm, flagellum width = 12 ± 0 μm, n = 2); both sections smooth, featureless; 7 – 8 SPM tightly coiled in one BC (AM C. 585236).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF0F828FFF68FBC2FDFCFA76.taxon	diagnosis	Comparative remarks. In our mitochondrial tree (Figs 1, 4, Clade E), S. costellata sp. nov. (normalis group, unit 13) is more closely related to four species, S. normalis, S. madangensis, S. fuliginata, and S. campestra. It differs from any of these by COI distances of ≥ 7.3 %. Throughout its range, this species has been found in sympatry with four congeners. Three species are sympatric in northern Australia, from N Qld to Nhulunbuy, NT: For comparisons with S. viridis, S. atra, and S. normalis refer to comparative remarks under these species. In Singapore the species is sympatric with S. viridis and S. alba; refer to comparative remarks under these species. Siphonaria madangensis sp. nov. differs in having a smaller, lower shell, a smaller AO, larger BC, and a more bulbous SPM. The specimen identified as ‘ normalis group, unit 13 ’ in Dayrat et al. (2014: 266, fig. 3 O) is a member of S. costellata.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF0F828FFF68FBC2FDFCFA76.taxon	distribution	Distribution and habitat. Recorded from Thailand, Singapore, Malaysia and NT, Australia (Fig. 55). In this study, found in sheltered and vertical positions on moderately exposed rocky shores across upper littoral levels (Fig. 54 T).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF0F828FFF68FBC2FDFCFA76.taxon	etymology	Etymology. From ‘ costellata’ (Latin = ribbed), referring to the finely ribbed shell; adjective.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF0E8288FF68FA42FE63FD76.taxon	description	(Figs 56 A – C, M, 58 C – D)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF0E8288FF68FA42FE63FD76.taxon	materials_examined	Material examined. Type material. Holotype, from Ethel Beach 10 ° 27.827 ’ S, 105 ° 42.497 ’ E, Christmas Is, Indian Ocean; coll. B. W. Jenkins, CI 02 - 1, 11 Sept 2018 (AM C. 585322 [M 421, SK 108], Fig. 56 A). Five paratypes, same data as holotype (AM C. 585585 2 p, WAM S 74043 p, WAM S 74041 p [M 307], Fig. 56 C, WAM S 74042 p [M 410], AM C. 608199 p [SK 093], Fig. 56 B).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF0E8288FF68FA42FE63FD76.taxon	description	External morphology. Animal evenly pale cream without any dark / black pigmentation; paler at foot / wall edge; mantle wider than foot wall, weakly lobed, translucent, covers exposed inner shell lip, outer edge thickened; genital pore indistinct, located on foot wall to right anterior of right cephalic fold; two small black epithelial eye spots centralised on two centrally touching cephalic folds; pneumostomal lobe long, under the mantle, unpigmented, between the right ADMs. Shell (Figs 56 A – C; Table S 9). size small (max sl mean = 8.5 mm SD = 1.0 mm, n = 4), ovate, shell thin translucent; tall, apex offset to posterior; protoconch below height of apex, direction heterostrophic (n = 2), protoconch area distinctly dark brown to black; anterior apical side convex, posterior side concave, lateral apical sides straight to weakly concave; growth lines prominent; rib count (mean = 29.3, SD = 4.8, n = 3) primary ribs pale to dark grey, weakly raised, secondary ribs not prominent, 1 – 2 between primary ribs, rib interstices dark brown to black; siphonal ridge not prominent, shell lip even, weakly corrugated with some primary rib ridges; interior polished, shell lip apically banded with white rays aligning with primary ribs extending and narrowing over shell margin and ADM scar to the start of dark chocolate brown spatula; CMS weakly concave, similar but darker colouration to spatula and shell margin; thickening of inner shell lip occurs in larger specimens, resulting in white coating covering the brown / black colouration of inner shell lip. Reproductive system (Fig. 58 C; n = 2). Positioned within entire right side of coelom, against foot wall on foot muscle, under the respiratory cavity occupying large proportion of animal body volume. GA, EG and ED positioned between BM and RAM. GA small, with singular GP through foot wall; AO distinct, bluntly pointed, maybe elongated, joined to upper GA alongside ED; ED long, broad, centrally bent, joins to side of GA; GA, AO, ED all white muscular fibrous tissue; EG medium, soft whitish tissue, slightly folded, joins ED; single very long flagellum (F 1), similar length and width to ED, appears as an extension of ED, possible F 2. BD and CD with opposing connections into GA between ED join and GP, CD short,; BD long, both ducts smooth, thickened, whitish, featureless, pass closely together through RAM (BD over CD) into soft white folded tissues of MG; MG / AG complex relatively large; BC embedded in folds close to embedded SV; BD with distal loop (s) and MA, longer thinner than CD; BC relatively large, spherical, thin whitish translucent test; HD short, wide, coiled, links ducts in soft white folded tissues of AG to small, yellowish granulated HG; outer edge of MG lobbed; AG larger than HG, both left outer sides curved to curvature of inner foot wall. Spermatophore (Fig. 58 D). Test thin, translucent (length = 5.77 ± 1.23 mm, n = 2); over half-length comprises a translucent cylindrical head section (head length = 4.73 ± 0.79 mm, ~ 49 % of SPM length, n = 3); tip bluntly rounded, containing a white gelatinous core mass; tapers into a filamentous transparent flagellum; head section much thicker than flagellum (head width = 59 ± 1.2 μm, flagellum width = 11 ± 0 μm, n = 3), both sections smooth, featureless; 6 and 3 SPM each in holotype and paratype.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF0E8288FF68FA42FE63FD76.taxon	diagnosis	Comparative remarks. In our mitochondrial phylogeny, S. delicata sp. nov. (plicata group, unit 62) is the sister species of S. christmasensis sp. nov. (Figs 1, 3). Both differ from each other by COI distances of ≥ 10.7 % (Table S 7). For comparison with S. christmasensis sp. nov. refer to comparative remarks under that species. We found this species in sympatry with four other congeners on CI: For comparisons with S. alba, and S. incerta refer to comparative remarks under these species. Siphonaria tenebrae sp. nov. has a lower, darker shell with greater edge scalloping and raised ribbing, a smaller AO and BC, and a shorter ED. Siphonaria umbra sp. nov. has a slightly larger, taller shell with a dual-ribbed siphonal ridge, a shorter ED, and a slightly shorter SPM.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF0E8288FF68FA42FE63FD76.taxon	distribution	Distribution and habitat. Known only from CI, Australia, and likely endemic to this island (Fig. 55). In this study, found in sheltered positions on exposed rocky shores, at upper littoral level (Fig. 56 M).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF0E8288FF68FA42FE63FD76.taxon	etymology	Etymology. From ‘ delicata’ (Latin = delicate) for the small, delicate shell; adjective.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF098288FF68FD42FAB4F956.taxon	description	(Figs 56 D – F, P, 58 E – F)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF098288FF68FD42FAB4F956.taxon	materials_examined	Material examined. Type material. Holotype, from Christi Rea Beach, 8 ° 32.072 ’ S, 125 ° 36.868 ’ E, Timor-Leste; coll. B. W. Jenkins, TL 01 - 2, 14 July 2019 (AM C. 584829 p [M 441, SK 227], Fig. 56 P). Paratype, same data as holotype (AM C. 585318 p [SK 546], Fig. 56 E); two paratypes from Dolokoan Beach, 8 ° 31.424 ’ S, 125 ° 37.091 ’ E, N of Dili, Timor-Leste; coll. B. W. Jenkins, TL 01 - 1, 18 July 2019 (AM C. 584825 p [M 445], Fig. 56 F, AM C. 584830 p [SK 228], Fig. 56 P). Other, non-type material. Dolokoan Beach, 8 ° 31.424 ’ S, 125 ° 37.091 ’ E, N of Dili, Timor-Leste (AM C. 585981 8 p, C. 585991 p [SK 547]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF098288FF68FD42FAB4F956.taxon	description	External morphology. Foot sole evenly cream; foot wall, foot edge, cephalic folds and pneumostomal lobe evenly grey cream, darker to foot edge, faint black pigmentation on foot wall; mantle thin, translucent, narrower than foot wall, edge weakly lobed, without pigmentation; cephalic folds small, pneumostome wide. Shell (Figs 56 D – F, P; Table S 9). medium to large sized (max sl mean = 15.6 mm, SD = 6.1 mm, n = 2), ovate; height medium to low; apex offset sightly posterior and central (usually eroded), apical sides convex, protoconch direction homostrophic (n = 1, Fig. 56 P), shell whorl dextral; growth striae indistinct, shell thick; exterior evenly brown maybe mottled, juveniles display a mix of bluish colouration; rib count (mean = 36, SD = 1.0, n = 2), primary ribs weakly bent, raised and protrude beyond shell lip (up to 1 mm) to unevenly scallop the edge; interspersed with 1 – 2 finer secondary ribs; single or paired primary ribs form siphonal ridge. Interior shell margin dark brown to tan, spatula dark chocolate brown, white rays aligning under primary / secondary ribs extend irregularly from shell lip over margin, siphonal groove prominent, same colour as shell margin; ADM scar distinct, CMS straight, paler than shell lip; thickening of shell margin noted forming a translucent layer. Reproductive system (Fig. 58 E; n = 2). Positioned within coelom under the respiratory cavity, hermaphroditic complex (HG, AG and MG) to posterior against right foot wall and over foot sole, epiphallic parts positioned to anterior between RAM and BM; AO medium, elongated, blunt, weakly centrally bent, merges to upper part of indistinct GA, singular GP; ED short, wide, compressed in coil, longer than AO, joins to lower side of GA; GA, AO, ED all white muscular fibrous tissue; EG soft whitish, folded, smaller than AO; single very long twisted uneven thin flagellum (F 1), tip of F 1 tucks into gap between start of FI and EG; BD and CD jointly but opposing connections to side of GA between AO and GP; both ducts smooth and pass closely together through RAM (BD over CD); BD long narrow with prominent short distal loop without MA to inner body wall, twist immediately prior to connection to medium sized bulbous BC with thin translucent test, embedded in lower folds of MG; CD short, wider than BD; CD connects into MG; HD short, thick, brown markings, coiled, under AG, links AG to much smaller yellowish granulated HG; MG and AG folded, soft white tissue. Spermatophore (Fig. 58 F, J). Thread-like (length = 5.21 mm, 52 % of AL, n = 1); translucent, test thin; head section narrow, bluntly rounded, cylindrical, flagellum tapers to a thin tip, tip attached to inner test of BC; both sections smooth, featureless. Head section longer wider than flagellum (head length = 3.16 mm; ~ 61 % of SPM length, head width = 70 μm, flagellum width = 17 μm, n = 1); 3 SPM tightly folded in yellowish gel in BC (AM C. 585991).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF098288FF68FD42FAB4F956.taxon	diagnosis	Comparative remarks. In our mitochondrial phylogeny (Figs 1, 2), S. forticosta sp. nov. (atra group, unit 71, Clade G) is the sister species of a sub-clade containing S. sirius, S. recurva sp. nov., S. incerta sp. nov. and an unidentified species from Tonga. It differs from these species by COI distances of ≥ 20 % (Table S 3). Siphonaria forticosta has been found in sympatry with six congeners in TL: For comparisons with S. javanica, S. viridis, S. campestra sp. nov., S. alba, and S. atra, refer to comparative remarks under these species. Siphonaria planucosta sp. nov. has a smaller, slightly darker shell with more prominent ribbing, weaker edge scalloping, and a smaller, narrower AO.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF098288FF68FD42FAB4F956.taxon	distribution	Distribution and habitat. Recorded from Dolokoan Beach, Timor-Leste (Fig. 55). In this study, found at mid littoral level in sheltered positions (mainly in crevices) on exposed shore boulders.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF098288FF68FD42FAB4F956.taxon	etymology	Etymology. From contraction of ‘ fortis’ (Latin = strong, sturdy) and ‘ costa’ (Latin = rib), referring to the sturdy primary ribs on the shell of this species.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF098284FCCAF962FE06F7D6.taxon	description	(Figs 56 G – T, 57 A – B, 58 G – J, 59 A – D)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF098284FCCAF962FE06F7D6.taxon	materials_examined	Material examined. Type material. Holotype, from Nightcliff 12 ° 22.836 ’ S, 130 ° 50.402 ’ E, Darwin, NT; coll. B. W. Jenkins, NT 23 - 1, 8 Sept 2017 (AM C. 585485 [M 088, SK 089], Fig. 56 G). Paratype, from Cox Peninsula 2, 12 ° 25.558 ’ S, 130 ° 44.593 ’ E; coll. B. W. Jenkins, NT 25 - 2, 10 Sept 2017 (AM C. 585487 p [M 089], Fig. 56 H), 2 paratypes, from Native Point oyster reef, Dundee Beach 12 ° 42.906 ’ S, 130 ° 20.653 ’ E, NT; coll. B. W. Jenkins, NT 26 - 6, 11 Sept 2017 (AM C. 585096 p [M 140], AM C. 585099 p [M 141]). Other, non-type material. Australia, Qld: Sweers Is, 12 ° 03.456 ’ S, 96 ° 52.329 ’ E, Q 56 - 2 (AM C. 585188 p [M 091]). NT: Luxmore Hd, Melville Is, 11 ° 20.639 ’ S, 130 ° 23.149 ’ E, NT 24 - 1 (AM C. 585336 p); Nightcliff, Darwin, 12 ° 22.836 ’ S, 130 ° 50.402 ’ E NT 23 - 1 (AM C. 585090 p [SK 078 protoconch C 5], C. 585091 p [SK 089]); Cox Peninsula 2, 12 ° 25.558 ’ S, 130 ° 44.593 ’ E NT 25 - 2 (AM C. 585092 p [M 144], C. 585093 p [M 145], C. 585094 p [M 090, SK 091], C. 585098 p [M 143]); N of Native Point, Dundee Beach, 12 ° 42.182 ’ S, 130 ° 20.881 ’ E NT 26 - 1 (AM C. 585095 p [SK 103], C. 585337 p [SK 088]); Native Point oyster reef, Dundee Beach, 12 ° 42.906 ’ S, 130 ° 20.653 ’ E NT 26 - 6 (AM C. 585097 p [M 094]); Native Point reef, Dundee Beach, 12 ° 42.981 ’ S, 130 ° 20.807 ’ E NT 26 - 3 (AM C. 585488 p [M 146]). WA: Tait Point, Mission Bay, 14 ° 05.442 ’ S, 126 ° 41.143 ’ E, WA 04 - 1 (AM C. 584669 p [SK 246]; Raft Point Collier Bay, 16 ° 04.045 ’ S, 124 ° 26.814 ’ E, WA 17 - 2 (AM C. 584668 p [M 464, SK 199]); CatamaranBay, 16 ° 27.622 ’ S, 123 ° 00.242 ’ E, WA 22 - 3 (AM C. 585301 p [M 071, SK 090], C. 585302 p [M 072], C. 585303 p [M 043]); Cape Keraudren, 19 ° 57.393 ’ S, 119 ° 46.358 ’ E, WA 29 - 2 (AM C. 585308 p [M 032]). Mauritius: Nth Albion Is, Mauritius, 20 ° 12.258 ’ S, 57 ° 24.252 ’ E, MRU 2 - 2 (AM C. 584971 [M 249], Fig. 56 I, C. 585976 20 p, AM C. 584972 p [M 250], Fig. 56 J). Australia, CKI: E coast of Home Is, 12 ° 06.917 ’ S, 96 ° 53.835 ’ E, CKI 04 - 1 (WAM S 74044 [M 317, SK 255], Fig. 56 K, AM C. 608183 15 p, C. 584666 p [M 318], C. 584667 p [M 319]); S coast of West Is, 12 ° 12.210 ’ S, 96 ° 50.372 ’ E, CKI 02 - 2 (AM C. 584849 p [M 315], Fig. 56 L, C. 584850 p [M 316]); West Island, N point (Trannies), 12 ° 08.540 ’ S, 96 ° 49.013 ’ E, CKI 01 - 1 (AM C. 585372 10 p); S coast of West Is, 12 ° 12.210 ’ S, 96 ° 50.372 ’ E, CKI 02 - 2 (AM C. 585373 10 p, C. 584665 p [M 314], C. 585933 p [SK 092], C. 585942 p [SK 086]); N coast Direction Is, 12 ° 03.456 ’ S, 96 ° 52.329 ’ E, CKI 03 - 1 (AM C. 585374 10 p, C. 585941 p [SK 535]). Marquesas Islands: Nuku Hiva, Baie des Controleurs, 08 ° 53.92 ’ S, 140 ° 02.92 ’ W, MQ 7 - M (MNHN IM- 2013 - 74895 [M 574], Fig. 57 A), Baie d’Hakatea, 08 ° 56.53 ’ S, 140 ° 09.69 ’ W, MQ 25 - M, (MNHN IM- 2013 - 74904 p [M 566], Fig. 57 B; IM- 2013 - 74902 p [M 571], IM- 2013 - 74900 p [M 576], IM- 2013 - 74903 p [M 572], MNHN IM- 2013 - 74901 p [M 573]), Baie des Controleurs, 08 ° 53.92 ’ S, 140 ° 02.92 ’ W, MQ 7 - M (MNHN IM- 2013 - 67892 p [M 578], IM- 2013 - 74894 p [M 575], IM- 2013 - 74896 p [M 564]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF098284FCCAF962FE06F7D6.taxon	description	External morphology (Fig. 56 M). Foot sole and foot wall evenly pale yellowish grey, paler at foot / wall edge; foot wall and mantle more yellowish, without black pigmentation; mantle strongly lobed, translucent, as wide as foot wall, covers exposed inner shell lip; genital pore inconspicuous, located on foot wall to right anterior of right cephalic fold; two small black epithelial eye spots centralised on two centrally touching unpigmented cephalic folds; pneumostomal lobe long, under the mantle, unpigmented, between the right ADMs. Shell (Fig. 56 G – T, 57 A – B; Table S 9). Shell ribbing and colouration variable; small sizing (max sl mean = 9.3 mm, SD = 1.7 mm, n = 8); ovate to elongate, height medium; apex often tall, pointed (protoconch very tall when intact), slightly offset posteriorly and left; protoconch direction heterostrophic (n = 2; Fig. 56 T) to upright (n = 2; Fig. 56 Q), shell whorl dextral; apical sides weakly convex, posterior concave; protoconch area distinctly darker brown, growth lines usually distinct undulating; rib count (mean = 28, SD = 5.3, n = 8), primary ribs white, curved, wavy, moderately flare at and extend beyond shell edge, rib ridges rounded, rib interstices dark brown / black; paired primary ribs form siphonal ridge, extend well beyond line of shell lip, strongly to weakly flared upwardly at shell edge; none to one secondary ribs between primary ribs, either side of siphonal ridge the number of secondary ribs greater; interior shell lip uneven, corrugated to no corrugations, shell lip with white blotches or rays under primary ribs, dark brown / black under rib interstices; white rays may extend over shell margin; spatula usually evenly dark brown / black but may be white to dull white; siphonal groove usually white, may be dark brown / black, bounded on either side by prominent dark brown / black axial patches aligning with primary rib interstices. WA and NT specimens often with heavy and strongly raised ribbing with corrugations on shell lip, CKI specimens possess flatter ribs and few to no corrugations on shell lip; thickening of inner shell lip and spatula not observed. Reproductive system (Fig. 58 G, I, 59 A; n = 5). Positioned within coelom under the respiratory cavity, occupies the entire right side of coelom, hermaphroditic glands positioned to posterior against right foot wall and over foot sole, epiphallic parts positioned to anterior between BM and RAM; GA small, with singular GP through foot wall; AO very small, bluntly pointed, joined to upper GA alongside ED; ED short, broad, slightly bent (without prominent MA), joins to side of GA; GA, AO, ED all white muscular fibrous tissue; EG large, soft whitish tissue, slightly folded, joins ED; single narrow flagellum (F 1; possible 2 nd flagellum), similar length and width to ED, appears as a continuous extension of ED to EG, laid over BM, often centrally bent; BD and CD connect together into GA between ED / AO joint and GP, both ducts short, straight, smooth, thickened, whitish, featureless, pass closely together through RAM (BD over CD) into soft white folded tissues of MG; MG / AG complex relatively large; BC embedded in folds of AG / MG close to embedded SV; BD on posterior side only, unlooped, similar thickness to CD; BC relatively large, spherical, thin translucent test; HD short, not thickened, coiled, links ducts in soft white folded tissues of AG to yellowish granulated HG; outer edge of MG lobbed; AG much larger than HG, sides match curvature of inner foot wall. Spermatophore (Fig. 58 H, J, 59 B). Broad head with short flagellum (length = 4.0 ± 2.9 mm, 64 % of AL, n = 4), head section cylindrical, bulbous, centrally bent, rounded tip; test thin, smooth, featureless, translucent encasing a white opaque central core; short tapering section merges head to filamentous flagellum; head shorter, wider than translucent flagellum (head length = 1.8 ± 0.18 mm, ~ 82 % of SPM length, head width = 171 ± 14.8 μm, flagellum width = 24 ± 1.3 μm, n = 4); 1 SPM coiled in one BC (AM C. 585488). The SPM of specimens from Mauritius differs somewhat in having a shorter flagellum (Fig. 58 J, length = 4.34 ± 3.42, n = 3, AM C. 584971). Specimens from the Cocos (Keeling) Islands have a broad head with short flagellum (length = 5.8 ± 3.2 mm, n = 2), head section cylindrical, bulbous, centrally bent, rounded tip; test thin, smooth, featureless, translucent encasing a white opaque central core; short tapering section merges head to filamentous flagellum; head shorter, wider than translucent flagellum (length = 4.0 ± 2.1 mm, ~ 69 % of SPM length, head width = 249 ± 20.2 μm; flagellum width = 29 ± 2.1 μm, n = 4); 1 – 2 SPM in each of two BC (WAM S 74044, AM C. 585942).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF098284FCCAF962FE06F7D6.taxon	diagnosis	Comparative remarks. In our mitochondrial phylogeny (Figs 1, 4), S. gemina populations from northwestern Australia, CKI, Mauritius, and the Marquesas Islands form a clade with little apparent genetic differentiation between them (normalis group, unit 78). COI sequences of this unit differ from each other by genetic distances of up to 6.2 % (Table S 8). We observed minor, yet consistent morphological differences between populations from mainland Australia, Mauritius, the Marquesas Islands, and the Cocos Keeling Islands in features of the shell, BC, and SPM. However, we consider these differences to represent intraspecific morphological variation. Siphonaria gemina sp. nov. is the sister species of an unidentified Siphonaria sp. (unit 11, from NT, Australia and Yap Is). These two clades differ by COI distances of ≥ 14.1 % (Table S 8). Throughout its range in tropical WA, we found S. gemina gemina in sympatry with five congeners: For comparisons with S. alba, S. normalis, S. zelandica, and S. viridis refer to comparative remarks under these species. Siphonaria restis sp. nov. has a larger, lower, sturdier shell with a less prominent siphonal ridge, stronger edge scalloping, a larger AO and BC, longer ED and BD, and a shorter SPM. Specimens figured as ‘ Siphonaria unit 51 ’ by Ossenbrügger et al. (2023) belong to the same MOTU, unit 78 and correspond well with specimens examined here.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF098284FCCAF962FE06F7D6.taxon	distribution	Distribution and habitat. Siphonaria gemina has a wide distribution spanning Mauritius, the Seychelles, the Cocos (Keeling) Islands, Tropical Australia, between Cape Keraudren in northern WA and the Sweers Is, Gulf of Carpentaria, Qld, to the Marquesas Islands (Fig. 55). In this study, found on moderately exposed rocky shores, in upper littoral level (Fig. 56 S).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF098284FCCAF962FE06F7D6.taxon	etymology	Etymology. From ‘ geminus’ (Latin = paired), referring to the prominence of paired siphonal primary ribs accentuated by dark flanking bands on the shell of this species; adjective.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF058286FCCAFF02FE21FC56.taxon	description	(Figs 57 C – E, M – N, T, 59 E – F)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF058286FCCAFF02FE21FC56.taxon	materials_examined	Material examined. Type material. Holotype, from Souillac 20 ° 31.519 ’ S, 57 ° 31.633 ’ E, Mauritius; coll. B. W. Jenkins, MRU 01 - 1, 9 Aug 2018 (AM C. 584963 [M 256], Fig. 57 A). Six paratypes, same data as holotype (AM C. 585910 4 p), Souillac 20 ° 31.467 ’ S, 57 ° 31.582 ’ E, Mauritius; coll. B. W. Jenkins, MRU 01 - 2, 9 Aug 2018 (AM C. 584966 p [M 257], Fig. 57 B; AM C. 585909 p [SK 389], Figs 57 C, T). Other, non-type material. Mauritius: Isle de la Passe 20 ° 24 ’ S, 57 ° 46.133 ’ E (WAM S 72343 7 p).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF058286FCCAFF02FE21FC56.taxon	description	External morphology (Fig. 57 M). Foot sole, foot wall, cephalic folds and pneumostomal lobe evenly cream, foot edge and mantle paler; mantle narrow, edge lobed, thickened, faint grey banding aligning with shell rib interstices; faint grey pigmentation over cephalic lobes. Shell (Figs 57 C – E; Table S 9). small sized (max sl mean = 9.3 mm, SD = 0.5 mm, n = 2), ovate, height medium; apex offset weakly posterior and to left; apical sides convex, posterior concave to straight; protoconch direction weakly heterostrophic to central (n = 1, Fig. 57 T), area black colouration, shell whorl dextral; growth striae weak; shell thickness thin; colouration uneven with some radial banding; rib count (mean = 41.5, SD = 1.5, n = 2); slightly raised, pale white, fairly straight, faintly protrude beyond shell lip; predominantly primary ribs, finer secondary ribs interspersed, develop between primary ribs with shell growth, rib interstices darker; siphonal ridge formed by paired primary ribs, protrudes past shell edge, otherwise indistinct. Interior evenly dark brown to black from margin to spatula, paler on shell lip aligning under rib ends, siphonal groove clear; ADM scar indistinct, CMS weakly convex; No evidence of growth variations in shell thickness or shell margin colouration. Reproductive system (Fig. 59 E; n = 1): Positioned within coelom under the respiratory cavity, hermaphroditic glands positioned to posterior against right foot wall and over foot sole, epiphallic parts lay over BM and to side of RAM, F 1 folded over back of BM. ED distinct entry to top of GA; AO large, elongated, bluntly pointed hook, centrally bent, attached to top of large, bulbous GA; ED thick, elongated, centrally bent, twisted; AO, GA and ED all muscular white tissue; EG large, folded, soft white tissue; single elongated, centrally looped, broad, blunt flagellum F 1 appears as extension of ED at EG join; BD and CD connect into GA in opposing directions close to ED attachment, both ducts smooth, short, thick, slightly bent, pass together between RAM and inner foot wall (BD over CD) connecting into MG, BD with coiled distal loop; BC white opaque test, relatively large, bulbous, embedded along with part of BD in soft white folds of MG, inner edge of MG lobed; HD narrow, coiled, links soft white folded AG to small yellowish granulated HG; AG larger than HG, both with outer sides curved reflecting the close positioning to curvature of inner foot wall at right posterior quarter of coelom; SV embedded in AG close to BC. Spermatophore (Fig. 59 F). Thread-like (length = 8.98 mm, n = 1), translucent, test thin; head section bluntly rounded, evenly cylindrical, containing a core white gelatinous mass, tapers along the transparent flagellum to a thin tip; keel as high as body width occurs along mid ¾ of head section; both sections smooth, featureless. Head section longer wider than flagellum (head length = 6.89 mm, ~ 77 % of SPM length, flagellum length = 2.08 mm, head width = 103 μm, flagellum width = 17 μm, n = 1). 3 SPM tightly coiled embedded in whitish gelatinous mass in BC (AM C. 584966).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF058286FCCAFF02FE21FC56.taxon	diagnosis	Comparative remarks. Siphonaria griffithsorum sp. nov. (plicata group, unit 64) is well-individualised genetic lineage within the pectinate group. It represents the sister lineage of the species pair S. delicata sp. nov. and S. christmasensis sp. nov. (Figs 1, 3). It differs from other species by COI distances of ≥ 23.7 % (Table S 7). We found S. griffithsorum sp. nov. in sympatry with two congeners: For a comparison with S. plana and S. incerta refer to comparative remarks under these species. Siphonaria griffithsorum sp. nov. closely resembles S. lirata from Guam by having a heterostrophic protoconch.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF058286FCCAFF02FE21FC56.taxon	distribution	Distribution and habitat. Recorded as endemic to Mauritius, Indian Ocean (Fig. 60). In this study, found in sheltered positions on exposed rocky shores, upper and mid littoral levels (Fig. 57 N).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF058286FCCAFF02FE21FC56.taxon	etymology	Etymology. For Mary-Ann and Owen Griffiths, Mauritius, in recognition of their hospitality and selfless assistance with collecting provided to the first author whilst visiting Mauritius.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF078286FF68FC62FC0EF7F6.taxon	description	(Figs 57 F – H, O – P, S, 59 G – H)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF078286FF68FC62FC0EF7F6.taxon	materials_examined	Material examined. Type material. Holotype, from Halafuoleva Beach, 21 ° 12.021 ’ S, 175 ° 14.680 ’ W, S coast of Tongatapu, Tonga; coll. B. W. Jenkins, TO 03 - 1 - 2, 23 May 2019 (AM C. 585279 [M 420, SK 107], Fig. 57 F). Paratypes, same data as holotype (AM C. 585540 20 + p; AM C. 585281 p [SK 224], Fig. 57 G; AM C. 585911 p [SK 387], Fig. 57 H). Other, non-type material. Tonga: Halafuoleva Beach, S coast Tongatapu, 21 ° 08.358 ’ S, 175 ° 02.443 ’ W, TO 03 - 1 - 2 (AM C. 586002 3 p, C. 585282 p [SK 222]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF078286FF68FC62FC0EF7F6.taxon	description	External morphology (Fig. 57 P). Foot sole dark grey; foot edge and cephalic folds cream; foot wall, mantle and pneumostomal lobe evenly grey, darker to foot edge, some irregular blotches of black pigmentation on foot wall; mantle thin, translucent, narrower than foot wall, edge thickened, lobed, with weak black banding aligning with rib interstices, pneumostome wide. Shell (Figs 57 F – H, O, P; Table S 9). Small sized (max sl mean = 11.1 mm, SD = 1.5 mm, n = 2), circular ovate; height medium; apex offset central, apical sides convex, protoconch direction central to homostrophic (n = 1, Fig. 57 O); shell whorl dextral; initial growth profile very tall, profile flattens with maturity, exterior pale blue / grey uneven, protoconch area darker, shell thick, growth striae prominent; rib count (mean = 34.5, SD = 7.5, n = 2), primary ribs pale grey, fairly straight, ridges rounded, increasingly raised to and extend beyond uneven shell lip to unevenly scallop the lip; two prominent primary ribs extending strongly beyond shell edge form the siphonal ridge. Interior tan to golden brown, shell lip white with dark chocolate brown rays extending over shell margin and ADM to spatula, aligning under rib interstices; siphonal groove prominent; ADM scar distinct, CMS convex; no thickening of shell lip noted. Reproductive system (Fig. 59 G; n = 1). Positioned within right coelom under the respiratory cavity, hermaphroditic complex (HG, AG and MG) to posterior against right foot wall and over foot sole, epiphallic parts to anterior between RAM and BM; AO medium, broad, blunt, weakly centrally bent, side MA, merges to upper part of indistinct GA, singular GP; ED short, wide, compressed in coil, longer than AO, joins to lower side of GA; GA, AO, ED all white muscular fibrous tissue; EG soft whitish, folded, slightly smaller than AO; single twisted thin flagellum (F 1) lays over MG; BD and CD join closely but with opposing connections to side of GA between AO and GP; both ducts smooth and pass closely together through RAM (BD over CD); BD long very narrow with prominent short distal loop with MA to inner body wall, connected to small sized bulbous BC with thin translucent test, positioned in folds of AG; CD short, wider than BD; CD connects into MG; HD short, thick, brown markings, coiled, under AG, links AG to much smaller yellowish granulated HG; MG and AG folded, soft white tissue. Several parasite eggs were found on the EG and AG of holotype (Halafuoleva Beach, S coast Tongatapu, Tonga). Spermatophore (Fig. 59 H). Thread-like (length = 5.64 mm, n = 1), translucent, test thin; head section bluntly rounded, evenly cylindrical, containing a core white gelatinous mass, tapers along the transparent flagellum to a thin tip; both sections smooth, featureless. Head section longer wider than flagellum (head length = 3.36 mm, ~ 64 % of SPM length, n = 1; flagellum length = 1.98 mm, head width = 45 μm, flagellum width = 11 μm). 2 SPM tightly coiled, embedded in purple / brown gelatinous mass in one BC (AM C. 585281).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF078286FF68FC62FC0EF7F6.taxon	diagnosis	Comparative remarks. In our mitochondrial phylogeny (Figs 1 – 4), S. tongatapuensis sp. nov. (plicata group, unit 69) is a well-individualized lineage that represents the sister group of a clade comprising S. guamensis, an unidentified species from Rarotonga (‘ unit 17 ’ in Dayrat et al. 2014), and S. nusalikensis sp. nov. It differs from other species by COI distances of ≥ 26 % (Table S 7). We found S. tongatapuensis sp. nov. in sympatry with S. plicata; for comparative remarks refer to that species. While both species have a similar RS structure, in particular size and shape of the epiphallic parts and SPM, they clearly differ in shell sculpture, colouration, and absence of a prominent multi-ribbed siphonal ridge. Siphonaria tongatapuensis sp. nov. differs from other species of the plicata group, such as S. nuttallii, S. lirata, and S. monticulus particularly in size and shape of AO, ED, and HD. The SPM shape of these species is generally similar. Siphonaria nuttallii has the most similar RS.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF078286FF68FC62FC0EF7F6.taxon	distribution	Distribution and habitat. Recorded exclusively from Halafuoleva Beach, Tonga (Fig. 60). In this study, found in sheltered positions (mainly in rock hollows) on exposed rocky shores, mid and upper littoral levels (Fig. 57 P).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF078286FF68FC62FC0EF7F6.taxon	etymology	Etymology. For Tongatapu, Tonga, where this species is found.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF068280FF68F802FB1EF816.taxon	description	(Figs 57 I – L, Q – R, 59 I – J)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF068280FF68F802FB1EF816.taxon	materials_examined	Material examined. Type material. Holotype, from Ponerihouen, 21 ° 05.644 ’ S, 165 ° 26.646 ’ E, NC; coll. B. W. Jenkins, NC 03 - 1, 23 Oct 2018 (AM C. 584989 [SK 127], Fig. 57 I). 27 paratypes, same data as holotype (AM C. 585524 20 + p, C. 584985 p [M 355], Fig. 57 J, C. 584986 p [M 357], C. 584987 p [M 358], C. 584988 p [M 378], C. 584990 p [SK 170], Fig. 57 K, C. 584991 p [SK 362]); paratye from Ouassé nr Canala, 21 ° 30.346 ’ S, 166 ° 03.732 ’ E, NC; coll. B. W. Jenkins, NC 02 - 1, 22 Oct 2018 (AM C. 584805 p [M 375], Fig. 57 L). Other, non-type material. NC: Hienghène, 20 ° 41.210 ’ S, 164 ° 59.108 ’ E NC 04 - 1 (AM C. 584808 p); Ouassé nr Canala, 21 ° 30.346 ’ S, 166 ° 03.732 ’ E, NC 02 - 1 (AM C. 585968 10 + p); S of Pouebo NC 04 - 2 (AM C. 585399 7 p); Presqu’ile de Ouano La Foa 20 ° 51.434 ’ S, 165 ° 48.479 ’ E NC 06 - 4 (AM C. 584816 3 p); Presq’ile Ducos Baie des Dames Noumea 22 ° 14.170 ’ S, 166 ° 24.524 ’ E NC 01 - 1 (AM C. 584803 16 p).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF068280FF68F802FB1EF816.taxon	description	External morphology (Fig. 57 Q). Foot sole, foot wall, mantle evenly pale grey, paler to foot edge; no pigmentation; mantle covers shell lip, wide, edge thickened lobed, paler grey; pneumostome lobe large, under mantle. Shell (Figs 57 I – L; Table S 9). Medium sized (max sl mean = 19.8 mm, SD = 1.9 mm, n = 7), ovate to elongate; height medium to low; apex offset central, apical sides strongly convex, protoconch direction undetermined, shell whorl dextral; growth striae prominent in bands, shell thickness thick; 3 radial band layers: whitish protoconch, mid shell dark brown band, and mid shell to edge pale brown / grey band; growth lines indistinct; rib count (mean = 45, SD = 6.5, n = 7), primary ribs pale white, fairly straight, ridges rounded, increasingly raised to and slightly protrude beyond uneven shell lip; shell lip scalloped between primary ribs; paired primary ribs form siphonal ridge which protrudes up to 1 mm beyond shell lip; secondary ribs similar to primary ribs, rib interstices slightly darker, indistinct. Interior margin irregular brown patterns; underside of primary ribs furrowed, white to cream rays, rib interstices marked with dark brown rays, some rays extend from spatula to shell lip; spatula dark chocolate brown, maybe white in smaller / juvenile specimens; siphonal groove distinctly furrowed, fairly straight bounded by dark brown patches; ADM scar distinct white to brown, paler than spatula and margin, CMS straight, paler than shell lip; Internal shell thickening occurs in larger / more mature specimens; shell lip thickens, whitens with yellow tinge, infills and reduces lip scalloping, spatula becomes whitened, distinct dark brown patches remain prominent on both sides of the siphonal groove. Significant colour and sculpture variation exists in the shell of this species. Reproductive system (Fig. 59 I; n = 5). Overall very small; hermaphroditic (HG, AG and MG) complex (AL = 12.52 mm) on right side within coelom against inside of foot muscle and foot wall under the respiratory cavity and intestine; epiphallic parts beside BM and RAM; GA very small; AO very large, elongated, bluntly bulbous, joins to GA, singular GP; ED relatively short, broad, smaller than AO, joins to side of GA; GA, AO, ED all white muscular fibrous tissue; EG soft whitish, slightly folded, smaller than AO, single short blunt flagellum (F 1); BD and CD jointly but opposing connections to GA between ED, AO and GP; BD narrow short with a prominent distal loop without any MA; CD broad short; both ducts smooth and pass closely together through RAM (BD over much broader CD); CD connects into MG; BD connects to small BC with thick white test, embedded in mid folds of MG / AG close to SV; HD short, narrow, coiled, links AG to a larger yellowish granulated HD; MG and AG small, folded, soft white tissue. Spermatophore (Fig. 59 J). Relatively short (length = 5.69 ± 0.99 mm, n = 2) and narrow; test thin, translucent, containing a white gelatinous core mass; over half-length comprises a translucent bulbous cylindrical body section (head length = 3.01 ± 1.18 mm, ~ 63 % of SPM length, n = 3), tip bluntly pointed, tapering into a filamentous transparent flagellum; head section much thicker than flagellum (head width = 72 ± 29 μm, n = 3; flagellum width = 13 ± 3 μm), both sections smooth, featureless; 2 SPM tightly coiled in single BC (AM C. 584989).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF068280FF68F802FB1EF816.taxon	diagnosis	Comparative remarks. In our mitochondrial tree (Figs 1, 2, Clade G), unit 40 (atra group) is the sister lineage of S. atra (unit 41). Unit 40 contains S. hienghenensis sp. nov. and sequences from Qld and PNG that probably represent an undescribed species. These sequences are excluded from S. hienghenensis sp. nov. delimited here. Siphonaria hienghenensis sp. nov. differs from S. atra by COI distances of ≥ 8.3 %. The next more closely related species is S. alba (unit 39), which differs by COI distances of ≥ 13 % (Table S 3). Siphonaria hienghenensis sp. nov., considered to be endemic to New Caledonia, has been found in sympatry with four species in NC: For comparisons with S. monticulus and S. normalis refer to comparative remarks under these species. Siphonaria ouasseensis sp. nov. has a smaller, taller, darker shell with more raised and broader ribbing, BD with a bursal loop and a narrower F 1. Siphonaria bourailensis sp. nov. has a smaller, taller, paler shell with more uneven ribbing, darker interior, a larger pointed AO, and a shorter SPM. Six are sympatric on NC. Siphonaria caledonica sp. nov. has a smaller, taller, darker shell with stronger edge scalloping and darker interior, and a larger pointed AO and ED. Siphonaria poindimiensis sp. nov. has a smaller, taller, darker shell with stronger edge scalloping and darker interior, and a larger pointed AO. Siphonaria namukaensis sp. nov. has a smaller, slightly darker shell with more raised ribbing, weaker edge scalloping, and a smaller, narrower AO, Siphonaria poindimiensis sp. nov. has a taller shell with stronger edge scalloping, wider ribs, fused dual siphonal ridge ribs, patterned interstices, a larger AO, and a longer ED. For comparisons with S. viridis and S. atra refer to comparative remarks under these species.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF068280FF68F802FB1EF816.taxon	distribution	Distribution and habitat. Recorded as endemic to NC and Lifou (Fig. 60). In this study, found on exposed to sheltered rocky boulder and platform shores, mid and upper littoral levels (Fig. 57 R).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF068280FF68F802FB1EF816.taxon	etymology	Etymology. Named after Baie de Hienghène, immediately north of the type location of Ponerihouen, west coast of NC, Pacific Ocean.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF008281FF68FD42FB0EFA96.taxon	description	(Figs 61 F – G, 62 A)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF008281FF68FD42FB0EFA96.taxon	materials_examined	Material examined. Type material. Holotype, from Itampolo, SW Madagascar; coll. O. Griffiths, MA 09 _ 1 b, July 2018 (AM C. 584955 [M 273], Fig. 61 F). Paratype, same data as holotype (AM C. 584956 p [M 274], Fig. 61 G).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF008281FF68FD42FB0EFA96.taxon	description	External morphology. Foot sole, foot wall, cephalic folds and pneumostomal lobe evenly cream in colour, paler to foot edge; mantle thin, translucent, wider than foot wall, strongly lobed with thickened edge, faint brown pigmentation on mantle edge; no black pigmentation; pustules prominent on foot wall; two black ‘ Eye’ spots prominent centrally on thickened cephalic lobes; pneumostome fold prominent. Shell (Figs 61 F – G; Table S 9). Small sized (max sl mean = 9.07 mm, SD = 0.35 mm, n = 2), circular ovate; height tall; apex offset central, apical sides convex, protoconch direction central to weakly homostrophic (n = 1), shell whorl dextral; growth striae distinct, exterior uneven, shell thickness thick; rib count (mean = 25, SD = 1.0, n = 2), predominance of pale white primary ribs, crooked, often discontinuous to shell lip, single secondary ribs may develop between primary ribs, rib interstices dark brown, ridges rounded, increasingly raised and protrude slightly beyond uneven weakly scalloped and strongly corrugated shell edge; siphonal ridge indistinct. Interior shell lip and margin dark chocolate brown with white rays aligning under primary / secondary ribs, fading to prominent thin ADM scar; spatula whitish to golden tan; siphonal groove indistinct, CMS convex; thickening or whitening of shell lip not observed. Reproductive system (Fig. 62 A; n = 2). Positioned within coelom under the respiratory cavity, hermaphroditic glands positioned to posterior against right foot wall and over foot sole, epiphallic parts lay over BM to side of RAM. ED distinct entry to top of GA; AO indistinct, part of GA, GA / AO large, elongated, weakly bent; ED thick, elongated, slightly twisted; AO, GA and ED all muscular white tissue; EG large, folded, soft white tissue; single looped thick blunt flagellum F 1 appears as extension of ED at EG join; BD and CD with bulbous ends connect side by side into GA, both ducts smooth, short, broad, slightly bent, pass together through RAM (BD over CD) connecting into MG; BC white opaque test, relatively small, bulbous, embedded along with part of BD in soft white folds of MG; HD long, narrow, coiled, links soft white folded AG to small yellowish granulated HG; AG larger than HG, both with outer sides curved reflecting the close positioning to curvature of inner foot wall at right posterior quarter of coelom; SV embedded in AG close to BC.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF008281FF68FD42FB0EFA96.taxon	diagnosis	Comparative remarks. In our mitochondrial phylogeny, S. itampoloensis sp. nov. (pectinata group, unit 81) is the sister species of S. carbo (Figs 1, 4) Both species differ from each other by COI distances of ≥ 5.1 % (Table S 8). Morphologically, S. carbo has a darker, more finely ribbed, grey / brown shell, a longer, wider BD and larger BC. We have found S. itampoloensis sp. nov. in sympatry with two congeners in Madagascar: Siphonaria striata sp. nov. has a lower shell with more raised ribs, a prominent siphonal ridge and posteriorly offset apex, a larger AO, a longer BD with distal and bursal loops, and a shorter F 1. For comparison with S. madagascariensis refer to comparative remarks under that species. Siphonaria itampoloensis sp. nov. differs from other members of the pectinata group, such as S. asghar, S. capensis, and S. pectinata (unit 4), by having a taller shell, more prominent shell ribbing, distinct shell interior colouration, and larger and thicker epiphallic parts.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF008281FF68FD42FB0EFA96.taxon	distribution	Distribution and habitat. Recorded exclusively from Madagascar, Indian Ocean (Fig. 60). In this study, found on intertidal rocks, on inner lagoon shores.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF008281FF68FD42FB0EFA96.taxon	etymology	Etymology. Named after the type location of Itampolo, SW Madagascar, Indian Ocean.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF008282FCCAFA22FC2BFCF6.taxon	description	(Figs 61 A – E, 62 B – C)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF008282FCCAFA22FC2BFCF6.taxon	materials_examined	Material examined. Type material. Holotype, from Lizard Island 14 ° 40.908 ’ S, 145 ° 27.007 ’ E, Qld, Australia; coll. B. W. Jenkins, Q 40 - 1, 29 June 2017 (AM C. 584789 [M 423, SK 114 (RS, SPM)], Fig. 61 A). Five paratypes, same data as holotype (AM C. 585641 4 p, C. 585496 p [M 398], Fig. 61 C); two paratypes from Freshwater Beach, Lizard Is, Qld; coll. P. H. Colman, 2 Dec. 1974 (AM C. 585032 p [SK 264], Fig. 61 B, C. 608197 p [SK 400], Fig. 61 E). Other, non-type material. Australia, Qld: Freshwater Beach, Lizard Is, 14 ° 39.922 ’ S, 145 ° 26.854 ’ E (AM C. 585943 20 + p).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF008282FCCAFA22FC2BFCF6.taxon	description	External morphology. Foot sole, foot wall, cephalic folds and pneumostomal lobe evenly light grey; foot sole darker to centre, paler to edge; fringing mantle narrow translucent at foot wall gradually becoming opaque with a thickened paler grey band at the lobed mantle edge, black pigmentation markings on band aligning with rib interstices; genital pore inconspicuous, located on foot wall to right anterior of right cephalic fold; two small black epithelial eye spots centralised on two thick centrally touching dark grey cephalic folds; thin pale grey pneumostomal lobe part of the mantle, between the right ADMs, closes the pneumostome and anus at the mantle edge. Shell (Figs 61 A – C, E; Table S 9). Small sized (max sl mean = 15.3 mm, SD = 2.7 mm, n = 3), elongate ovate; height tall; apex offset central weakly posterior, apical sides weakly convex, protoconch direction homostrophic (n = 1, Fig. 61 E), shell whorl dextral; growth striae prominent, 3 prominent coloured bands — protoconch area whitish, mid dark brown / black, margin and shell lip light brown; shell thick; rib count (mean = 43, SD = 5.0, n = 3), 8 – 10 primary ribs whitish, ridge rounded, fairly straight raised, weakly extend beyond slightly scalloped corrugated uneven shell lip; paired primary ribs on siphonal ridge, no more prominent than other primary ribs; few secondary ribs, rib interstices darker. Interior even white rays align on shell margin under primary / secondary ribs, brown rays under rib interstices, spatula dark brown or whitish; dark brown shell margin dark brown to tan, siphonal groove distinct, same colour as shell edge, points to right anterior; spatula dark chocolate brown to mottled tan even whitish; ADM scar distinct, CMS weakly convex. Degree of intraspecific variability low. Conchologically very similar to S. viridis in northern Qld to WA, Australia. Reproductive system (Fig. 62 B; n = 1). Positioned within coelom under the respiratory cavity, occupies the right side of coelom, hermaphroditic glands positioned to posterior against right foot wall and over foot sole, epiphallic parts positioned to anterior between BM and RAM; GA medium, with singular GP through foot wall; AO large, elongated, broad, rounded point, joins next to ED to upper GA; ED short, broad, coiled; GA, AO, ED all white muscular fibrous tissue; EG reasonably large, soft whitish tissue, slightly folded, joins ED; single elongated narrow flagellum (F 1), appears as an extension of similar length and broader ED. BD and CD connect side-by-side into GA between ED / AO joint and GP, both ducts smooth, thickened, whitish, featureless, pass closely together through RAM (BD over CD) into soft white folded tissues of MG; MG / AG complex relatively large; BC embedded in folds of AG / MG close to embedded SV; BD with distal loop, posteriorly short and without prominent MA, similar thickness to CD; BC relatively large, bulbous, thin whitish translucent test; HD short, narrow, coiled, links ducts in soft white folded tissues of AG to yellowish granulated HG; outer edge of MG lobbed; AG larger than HG, sides match curvature of inner foot wall. Spermatophore (Fig. 62 C). Thread-like (length = 6.73 mm, n = 1), translucent, test thin; head section bluntly rounded, evenly cylindrical, containing a white gelatinous core, tapers along the transparent flagellum to a thin tip; both sections smooth, featureless. Head section longer wider than flagellum (head length = 4.15 ± 0.34 mm, n = 2; ~ 58 % of SPM length, head width = 85 ± 24 μm, n = 2, flagellum width = 34 ± 16 μm); 5 SPM tightly coiled embedded in brown gelatinous mass in single BC (AM C. 584789).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF008282FCCAFA22FC2BFCF6.taxon	diagnosis	Comparative remarks. In our molecular phylogeny (Figs 1, 3), S. jiigurruensis sp. nov. (laciniosa group, unit 21) forms a well-differentiated lineage. The species differs from other species by COI distances of ≥ 23 % (Table S 6). We found S. jiigurruensis sp. nov. in sympatry with four congeners on Lizard Island, Qld: For comparisons with S. normalis, S. oblia, S. atra, and S. viridis refer to comparative remarks under these species. The specimen figured as ‘ laciniosa group, unit 21 ’ in Dayrat et al. (2014: 262, fig. 4 C) corresponds well with S. jiigurruensis sp. nov.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF008282FCCAFA22FC2BFCF6.taxon	distribution	Distribution and habitat. Recorded exclusively from Lizard Island, northern Qld, Australia (Fig. 60). In this study, found on granite boulder / platform exposed rocky shores, mid to upper littoral levels (above barnacle zone) (Fig. 61 D).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF008282FCCAFA22FC2BFCF6.taxon	etymology	Etymology. For Jiigurru, the name of Lizard Island in the language of the Dingaal people, traditional custodians of the land.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF038283FCCAFCC2FB81FA76.taxon	description	(Figs 61 H, 62 D)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF038283FCCAFCC2FB81FA76.taxon	materials_examined	Material examined. Type material. Holotype, from Bak Bak Beach, Kudat, Sabah, 07 ° 00 ′ N, 116 ° 46 ′ E, E Malaysia; coll. P. H. Colman, 31 March, 1984, HWL on rocks on beach (AM C. 585938 [SK 522], Fig. 61 H).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF038283FCCAFCC2FB81FA76.taxon	description	External morphology (preserved). Foot sole, foot wall, cephalic folds and pneumostome evenly cream, paler to foot edge; mantle edge thickened strongly lobed without darker pigmentation on mantle edge or cephalic folds. Shell (Fig. 61 H; Table S 9). Small sized (max sl = 12.8 mm, n = 1), circular ovate, height tall; apex weakly offset to posterior and centrally left, apical sides weakly convex except posterior which is straight; protoconch direction heterostrophic (n = 1), shell whorl dextral; growth striae clear; exterior pale brown, radial colour bands indistinct; rib count (38, n = 1), primary ribs fewer than secondary, very prominent, strongly raised, off white and rounded, broaden to shell edge; paired primary ribs form siphonal ridge, ribs ends protrude weakly at shell lip, shell edge weakly scalloped and corrugated, few secondary ribs, rib interstices indistinct, darker. Interior shell margin to spatula mottled dark chocolate brown; outer shell margin paler with white rays aligning under ribs; siphonal groove distinct, same colour as spatula, darker than margin; ADM scar indistinct, CMS convex; no thickening of shell lip noted. Reproductive system (Fig. 62 D; n = 1). Positioned within right side of coelom, against foot wall on foot muscle, under the respiratory cavity; epiphallic parts positioned between RAM and BM. GA small, with singular GP through foot wall; AO large, elongated, broad, tip bluntly pointed, weakly centrally bent, merges to upper part of GA, singular GP; ED long, wide, upper twist, shorter than AO, joins to lower side of GA; GA, AO, ED all white muscular fibrous tissue; EG soft whitish, folded; twisted narrow flagellum (F 1) lays over MG, possible second shorter flagellum (F 2); BD and CD join closely but with opposing connections to side of GA between AO and GP; both ducts long, featureless, smooth and pass outside RAM (BD over CD); BD with prominent short distal loop and MA attached to inner body wall, connected to small sized bulbous BC with thin translucent test, positioned in folds of AG; CD connects into soft white folded tissues of small MG / AG complex; BC relatively small, elongate, thin whitish translucent test, embedded in MG / AG folds close to large embedded SV; outer edge of MG lobed; HD large, coiled, links ducts in soft white folded tissues of AG to yellowish finely granulated HG; AG larger than HG, sides match curvature of inner foot wall.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF038283FCCAFCC2FB81FA76.taxon	diagnosis	Comparative remarks. We have found S. kudatensis sp. nov. (laciniosa group) in sympatry with two congeners at Kudat, Sabah. Siphonaria radiata has a lower, broader shell with a less offset apex, less prominent ribbing and siphonal ridge, a smaller AO, shorter BD without a distal loop and a larger BC. Siphonaria radians has a lower, broader shell with a less offset apex, finer ribbing and a less prominent siphonal ridge, a smaller AO, AG and HD, larger HG, and a longer F 1. We have been unable to sequence mtDNA markers from specimens of this species.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF038283FCCAFCC2FB81FA76.taxon	distribution	Distribution and habitat. Recorded exclusive from Sabah, Malaysia (Fig. 60). Found on rocky shores, intertidal.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF038283FCCAFCC2FB81FA76.taxon	etymology	Etymology. Named after the type locality of Kadat, Sabah, E Malaysia.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF0282BDFCCAFA42FEACFE96.taxon	description	(Figs 63 A – B, 64 C – D)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF0282BDFCCAFA42FEACFE96.taxon	materials_examined	Material examined. Type material. Holotype, from S Dumduman Is, Rempi Area, Madang, PNG Coll. PNG 2013, PM 12, 9 Nov. 2012 (MNHN IM- 2013 - 12006 [M 550], Fig. 63 A). Paratype, from Sek Island 05 ° 04,7 ’ S, 145 ° 48,9 ’ E, Madang, PNG; coll. PNG 2013, PM 22, 14 Nov. 2012 (MNHN IM- 2013 - 13132 p [M 552], Fig. 63 B).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF0282BDFCCAFA42FEACFE96.taxon	description	External morphology (preserved). Foot sole grey, paler to foot edge; foot wall, mantle and cephalic lobes darker grey; mantle edge band pale grey with darker markings aligning under rib interstices. Shell (Figs 63 A – B; Table S 9). Circular ovate, small sized (max sl mean = 12.1 mm, SD = 1.8 mm, n = 2); shell thin, height low to medium; apex offset central, apical sides straight to weakly convex, protoconch direction undetermined, shell whorl dextral; growth striae indistinct, radial banding apparent, shell edge even thin, weakly scalloped; rib count (mean = 55.5, SD = 2.5, n = 2), ribs white, straight, fairly even, weakly raised to unraised, interstices dark brown; siphonal ridge clear, formed by dual split primary ribs, extends beyond shell edge; 0 to 2 secondary ribs between primary ribs; interior golden brown, dark brown rays aligned under rib interstices, span from shell edge to ADM scar, spatula golden brown to chocolate brown, siphonal groove clear, shallow; CMS weakly convex; no whitening or thickening of shell lip observed. Reproductive system (Fig. 64 C; n = 1). Positioned within coelom under the respiratory cavity, occupies the right side of coelom, hermaphroditic glands positioned to posterior against right foot wall and over foot sole, epiphallic parts positioned to anterior between BM and RAM; GA small, with singular GP through foot wall; AO indistinct; ED short, broad, joins to side of GA; GA, AO, ED all white muscular fibrous tissue; EG small, soft whitish tissue, slightly folded, joins ED; single short broad flagellum (F 1), similar length and width to ED, appears as an extension of ED. BD and broader CD with juxtapose connections into GA between ED / AO joint and GP, both ducts smooth, broad, whitish, featureless, pass closely together through outer side of RAM (BD over CD) into soft white folded tissues of MG; BD without distal loop or prominent MA, centrally twisted; BC embedded in folds of small MG close to small pale embedded SV; BC medium sized, elongated, whitish translucent test; MG / AG complex relatively small; outer edge of MG lobbed; HD elongated, narrow, coiled, links ducts in soft white folded tissues of AG to small granulated HG; AG larger than HG, side reflects curvature of inner foot wall. Spermatophore (Fig. 64 D). Wide thread-like (length = 1.08 mm, n = 3), translucent, test thin; head section looped, pointed to bluntly rounded, bulbous cylindrical, containing a white gelatinous core, tapers along the transparent flagellum to a thin tip; both sections smooth, featureless, head section longer wider than flagellum (head length = 0.82 ± 0.14 mm, ~ 77 % of SPM length, head width = 92 ± 10 μm, flagellum width = 17 ± 0 μm, n = 3). 4 SPM tightly coiled in brown gelatinous mass in single BC (MNHN IM- 2013 - 13132).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF0282BDFCCAFA42FEACFE96.taxon	diagnosis	Comparative remarks. In our mitochondrial tree (Figs 1, 4), S. madangensis sp. nov. (normalis group, Clade E, unit 88) is most closely related to S. fuliginata (unit 80) from Rodrigues. Both species differ by COI distances of ≥ 5.9 %. Siphonaria madangensis differs from S. normalis by COI distances of ≥ 5.6 %. Siphonaria fuliginata has a paler shell with weaker scalloped edge, white to golden brown interior, a larger AO, a wider BD with bursal loop, and a thinner SPM. We found S. madangensis sp. nov. in sympatry with three congeners in northern PNG. For comparisons with S. atra, S. viridis, and S. normalis refer to comparative remarks under these species.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF0282BDFCCAFA42FEACFE96.taxon	distribution	Distribution and habitat. Recorded from S Dumduman and Sek Islands, Madang area, northern PNG (Fig. 67). In this study, found on limestone shores, intertidal.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF0282BDFCCAFA42FEACFE96.taxon	etymology	Etymology. Named after the type locality of Madang, northern PNG.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3C82BDFF68FE22FE6AF7D6.taxon	description	(Figs 63 C – F)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3C82BDFF68FE22FE6AF7D6.taxon	materials_examined	Material examined. Type material. Holotype, from N coast of Malo Is, 15 ° 37.7 ’ S, 167 ° 11 ’ E, Vanuatu; coll. Marine Biodiversity Survey, 18 Sept. 2006 VM 16, sand and coral (MNHN IM- 2006 - 31454 p [M 545] Fig. 63 C). Paratype, same data as holotype (MNHN IM- 2006 - 31358 p [M 546], Fig. 63 D); two paratypes, from Palikulo Peninsula, 15 ° 28.8 ’ S, 167 ° 15.3 ’ E, Vanuatu; coll. Marine Biodiversity Survey, 14 Sept. 2006 VM 11, hard bottom (MNHN IM- 2006 - 31355 p [M 547], Fig. 63 E; IM- 2006 - 31353 p [M 548], Fig. 63 F).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3C82BDFF68FE22FE6AF7D6.taxon	description	External morphology (preserved). Foot sole, foot wall, cephalic folds and pneumostome cream, paler to foot edge; mantle translucent, edge thickened strongly lobed with dark / black pigmentation on mantle edge aligning with rib interstices, faint pigmentation over cephalic folds. Shell (Figs 63 C – F; Table S 9). Ovate, small to medium sized (max sl mean = 17.9 mm, SD = 2.3 mm, n = 4); apex offset central and weakly to left, apical sides strongly convex, height tall; protoconch direction undetermined, shell whorl dextral; exterior uneven without prominent radial colour bands; growth striae distinct, shell thickness thick; rib count (mean = 45.5, SD = 2.3, n = 4), ribs fairly even, primary ribs pale grey, wavy, slightly broaden and increasingly raised to shell edge, ridges rounded narrow; edge finely scalloped and unevenly corrugated; siphonal ridge clear, formed by 2 – 3 primary ribs; few finer secondary ribs, rib interstices darker grey with irregular red / brown markings; interior shell margin white, red brown rays aligned under rib interstices, from shell lip over shell margin to spatula, siphonal groove distinct, same colour as shell margin; spatula dark brown; ADM scar distinct, darker brown, CMS convex; thickening and whitening of shell lip occurs (Fig. 63 C).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3C82BDFF68FE22FE6AF7D6.taxon	diagnosis	Comparative remarks. In our molecular phylogeny (Figs 1, 2), S. maloensis sp. nov. (laciniosa group, unit 87) is the sister species of S. caledonica sp. nov. (unit 23) from New Caledonia. Both species differ from each other by 16 S distances of ≥ 9.7 %. Siphonaria maloensis differs from S. normalis by 16 S distances of ≥ 10 %. Morphologically, S. caledonica sp. nov. has a smaller, thinner, taller, darker shell, with more prominent primary ribbing, and a less scalloped edge. We found S. maloensis sp. nov. in sympatry with S. viridis in Vanuatu; refer to comparative remarks under that species.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3C82BDFF68FE22FE6AF7D6.taxon	distribution	Distribution and habitat. Recorded exclusively from Malo Island, Vanuatu, Pacific Ocean (Fig. 67). In this study, found on hard substrate, sand and coral, intertidal.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3C82BDFF68FE22FE6AF7D6.taxon	etymology	Etymology. Named after the type locality of Malo Island, Vanuatu, Pacific Ocean.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3C82BFFCCAFF02FC49F956.taxon	description	(Figs 63 G – K, 64 A – B)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3C82BFFCCAFF02FC49F956.taxon	materials_examined	Material examined. Type material. Holotype, from Hanakao’o Beach 20 ° 54.586 ’ N, 156 ° 41.338 ’ W, Maui, Hawaii; coll. B. W. Jenkins, HA 03 - 2, 23 June 2018 (AM C. 584888 [M 297], Fig. 63 G). Six paratypes, same data as holotype (AM C. 584890 p [SK 245], Fig. 63 I, C. 584891 p [SK 207], Fig. 63 H, C. 585586 3 p, C. 584921 p [SK 206]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3C82BFFCCAFF02FC49F956.taxon	description	External morphology (Fig. 63 K). Foot sole grey; foot wall, mantle, cephalic folds, pneumostome evenly yellowish / green; irregular blotches of black pigmentation on foot wall, concentrated over cephalic lobes and posterior; mantle translucent narrower than foot wall, covers exposed inner shell lip, wider at anterior, mantle edge thickened white band lobed with even black pigmentation rays aligned with shell rib interstices; genital pore indistinct, located on foot wall to right anterior of right cephalic fold; small black epithelial eye spot centralised on each of centrally touching cephalic folds; pneumostomal lobe under mantle between the right ADMs. Shell (Figs 63 G – I; Table S 9): small sized (max sl mean = 11.93 mm, SD = 1.16 mm, n = 3), circular ovate; height tall; apex offset central, apical sides convex uneven, protoconch direction homostrophic to central (n = 1), shell whorl dextral, apex weakly hooked; growth striae distinct, 2 – 3 discontinuous bands of blue / grey radial shading, protoconch area dark brown, shell thin; rib count (mean = 60, SD = 4.3, n = 3), ribs white / grey, rib interstices narrow brown / black, difference between primary and secondary ribs indistinct, narrow, crooked, weakly raised, slightly broaden to and weakly protrude beyond uneven shell lip; siphonal ridge indistinct formed by 3 – 4 ribs. Interior shell margin mottled brown, shell lip dark brown with uneven white rays extending to shell margin under ribs; siphonal groove distinct, same colour as shell edge but maybe white; spatula dark chocolate brown; ADM scar distinct, CMS straight; thickening of shell lip occurs in mature specimens, infills and reduces scalloping of lip, spatula becomes whitened. Reproductive system (Fig. 64 A; n = 1). Positioned within right side of coelom, against foot wall on foot muscle, under the respiratory cavity; RS very large proportion of animal size compared to other species; epiphallic parts (GA, EG and ED) positioned over back of BM, F 1 long, draped over BM; GA small, with singular GP through foot wall; AO medium, bluntly rounded, joined at base to upper GA, rests against MG; ED long broad, centrally bent, joins to posterior side of GA; GA, AO, ED all white muscular fibrous tissue; EG very large, soft whitish tissue, folded, joins ED; single long broad flagellum (F 1) centrally looped, appears as an extension of wider ED; BD and CD connect closely in opposite directions into GA between ED join and GP, both ducts whitish curved, pass closely together around outside of RAM (BD over CD) into soft white folded tissues of MG / AG complex; outer edge of CD lobed, broadens to connection with AG ducts, BC long with distal loop and MA to inner foot wall in front of BM, initially broader than CD, narrows to embed in folds of AG; BC relatively large bulbous thin whitish translucent test, 2 SPM in BC (n = 1); HD short broad coiled, links ducts in soft white folded tissues of AG to orange granulated small HG; outer edge of MG lobbed; AG much larger than HG, outer sides of both reflect curvature of inner foot wall. Spermatophore (Fig. 64 B). Relatively long and thin; test thin, translucent, comprises a translucent cylindrical body section containing a white gelatinous thread-like core, tapers into a filamentous transparent flagellum (head length = 24.56 mm, n = 1; flagellum incomplete), head section thicker than flagellum (head width = 148 μm, flagellum width = 74.1 μm, n = 1); head tip bluntly rounded; both sections smooth, featureless; 2 SPM tightly coiled, embedded in brown gelatinous mass within BC (AM C. 584888).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3C82BFFCCAFF02FC49F956.taxon	diagnosis	Comparative remarks. Siphonaria mauiensis sp. nov. (plicata group, unit 60) forms a well-differentiated lineage in the mitochondrial tree (Figs 1, 3). It differs from other species by COI distances of ≥ 29.6 % (Table S 7). We found S. mauiensis sp. nov. in sympatry with two congeners in Hawaii. For comparisons with S. normalis and S. nuttallii refer to comparative remarks under these species. Along with the latter, S. undans sp. nov. (unit 61) and S. waikoloaensis sp. nov. (unit 55) are other Hawaiian species of the plicata group, but they are genetically highly distinct. The shell of S. mauiensis sp. nov. resembles that of S. lirata from Guam, but has a consistently darker interior.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3C82BFFCCAFF02FC49F956.taxon	distribution	Distribution and habitat. Recorded exclusively from Hanakao’o Beach, Maui, Hawaii, USA (Fig. 67). In this study, found in sheltered positions (mainly rock crevices) on moderately exposed fine-algal covered rocky basalt shores, upper littoral level (Fig. 63 K).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3C82BFFCCAFF02FC49F956.taxon	etymology	Etymology. Named after the type locality’s island of Maui, Hawaii.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3E82BAFCCAF962FB88FA16.taxon	description	(Figs 65 A – F, M – N, P; 66 A – D)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3E82BAFCCAF962FB88FA16.taxon	materials_examined	Material examined. Type material. Holotype, from Namuka Bay, 18 ° 08.094 ’ S, 177 ° 23.490 ’ E, Viti Levu, Fiji; coll. B. W. Jenkins, FI 02 - 2, 23 Aug 2018 (AM C. 585826 [M 282], Fig. 65 A). 25 + paratypes, same data as holotype (AM C. 585893 20 + p, C. 584857 p [M 278], C. 584858 p [SK 109], Fig. 65 B, C. 585823 p [M 279], C. 585824 p [M 280], Fig. 65 C; C. 585825 p [M 281], Fig. 65 D). Other, non-type material. Fiji, Viti Levu: Vuda Point Marina seawall, 17 ° 40.878 ’ S, 177 ° 23.009 ’ E, FI 03 - 2 (AM C. 585391 10 + p, C. 585696 8 p, C. 585828 p [M 283], C. 585829 p [M 284], C. 585830 p [M 285], C. 585831 p [M 422], C. 584864 p [M 422, SK 110], C. 584865 p [SK 116]); Heal of foot First Landing, 17 ° 40.753 ’ S, 177 ° 23.006 ’ E, FI 03 - 1 (AM C. 585827 p [M 290]); Namuka Bay 18 ° 08.094 ’ S, 177 ° 23.490 ’ E, FI 02 - 2 (AM C. 586006 20 + p, C. 584856 p [M 277], C. 584858 p [SK 109]); nr Tagaqa, S. coast of Viti Levu, 18 ° 11.802 ’ S, 177 ° 38.640 ’ E FI 02 - 1 (AM C. 585822 p [M 276]). NC: Poindimie 21 ° 55.901 ’ S, 165 ° 19.672 ’ E, NC 03 - 2 (AM C. 584994 p [M 455, SK 193], C. 585000 p [M 461, SK 194], Fig. 65 F); Poum 2, 20 ° 13.754 ’ S, 164 ° 01.699 ’ E, NC 05 - 3 (AM C. 585999 5 p).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3E82BAFCCAF962FB88FA16.taxon	description	External morphology (Fig. 65 M). Foot sole evenly dark grey; foot edge, foot wall, cephalic folds and pneumostomal lobe yellow grey; mantle wider than foot wall, thin translucent, edge thickened, weakly lobed, yellow band; irregular blotches of dark pigmentation on foot wall, cephalic folds, fading to mantle join; pneumostome elongated. Shell (Figs 65 A – F, P; Table S 9). Small to medium sized (max sl mean = 14.6 mm, SD = 3.7 mm, n = 7), circular ovate; height medium; apex offset central, apical sides convex, protoconch direction homostrophic (n = 2, Fig. 65 P), shell whorl dextral; growth striae prominent; rib count (mean = 52.4, SD = 6.5, n = 7), ribs fairly straight, rib ridges slightly raised, rounded, align with uneven and weakly scalloped shell edge; interstices narrow, black; primary and secondary ribs white, very similar in size, brown flecks and 1 – 2 secondary ribs in spaces between primary ribs; siphonal ridge sole rib fold, formed by 3 – 4 close primary ribs. Interior shell lip tan to whitish, shell margin golden dark brown to tan, white rays align on shell margin under primary / secondary ribs, prominent siphonal groove and spatula pale tan to whitish; ADM scar prominent, darker than shell lip, margin and shell edge; CMS straight; thickening and whitening of shell lip occurs. Reproductive system (Figs 66 A, C; n = 5). Positioned within right side of coelom, against foot wall on foot muscle, under the respiratory cavity; epiphallic parts positioned over BM. GA small, with singular GP through foot wall; AO large, broad, bent, bluntly pointed, joined to lower ED and upper GA; ED very short, very broad, centrally bent, joins to side of GA; GA, AO, ED all white muscular fibrous tissue; EG very large, soft whitish tissue, slightly folded, joins ED; extension joins in parallel to single narrow flagellum (F 1), similar width to ED, appears as an extension of ED, possible F 2 narrow short; BD and CD connect in opposing directions closely into GA between ED join and GP, both ducts short, slightly bent, smooth, thickened, whitish, featureless, pass closely together inside outer RAM (BD over wider CD) into soft white folded tissues of MG; MG / AG complex large; CD connecting to ducts, BC embedded in folds close to embedded SV; BD with distal loop and MA; BC relatively small, spherical, thin whitish translucent test; HD short, narrow, coiled, links ducts in soft white folded tissues of AG to yellowish granulated HG; outer edge of MG lobbed; AG larger than HG, sides match curvature of inner foot wall. Spermatophore (Figs 66 B, D). Long, thread-like (length = 8.76 mm, n = 1), translucent, test thin; head section bluntly rounded, evenly cylindrical, centrally bent, contains a core white gelatinous mass, tapers along the transparent and narrower flagellum (head length = 3.66 mm 42 % of total length; head width = 103 μm, flagellum width = 17 μm, n = 1); both sections smooth, featureless. SPM tightly coiled embedded in purple / brown gelatinous mass in BC (3 SPM in AM C. 585826, Fig. 66 B, 1 SPM in AM C. 585822, 5 SPM in AM C. 585831, 1 SPM in AM C. 584994, Fig. 66 D).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3E82BAFCCAF962FB88FA16.taxon	diagnosis	Comparative remarks. In our mitochondrial phylogeny (Figs 1, 3), S. namukaensis sp. nov. (laciniosa group, unit 22) is the sister species of S. yagasaensis sp. nov. (unit 67). Both species are well-differentiated from each other by COI distances of ≥ 17 %. From all other congeners S. namukaensis differs by COI distances of ≥ 18 % (Table S 6). Throughout its range, we found S. namukaensis sp. nov. in partial sympatry with six congeners. For comparisions with S. atra and S. normalis refer to comparative remarks under these species. Siphonaria vudaensis sp. nov. has a larger, lower, darker shell with stronger edge scalloping, and darker interior, a blunter AO. Siphonaria poindimiensis sp. nov. has a smaller taller darker shell with stronger edge scalloping, a larger AO and ED. Siphonaria tagaquensis sp. nov. has a smaller taller darker shell, a longer ED. Siphonaria poindimiensis sp. nov. has a taller, darker shell with broader ribs and stronger edge scalloping, and wider ribs, a larger BC and a longer, wider ED.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3E82BAFCCAF962FB88FA16.taxon	distribution	Distribution and habitat. Recorded from Viti Levu, Fiji and E coast NC, Pacific Ocean (Fig. 67). In this study found at sheltered positions on exposed and moderately exposed rocky shores, upper and mid littoral (Fig. 65 N).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3E82BAFCCAF962FB88FA16.taxon	etymology	Etymology. Named after the type locality of Namuka Bay, Viti Levu, Fiji.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3A82BBFF68FF02FB75FDD6.taxon	description	(Figs 63 L, 64 E)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3A82BBFF68FF02FB75FDD6.taxon	materials_examined	Material examined. Type material. Holotype, from NW side of Big Nusa Island, 02 ° 34,1 ’ S, 150 ° 46,7 ’ E, New Ireland, PNG; coll. KAVIENG 2014 expedition, KM 21, 28 Jun. 2014 (MNHN IM- 2013 - 55334 [M 532], Fig. 63 L).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3A82BBFF68FF02FB75FDD6.taxon	description	External morphology (preserved). Foot sole, foot wall, cephalic folds and pneumostome cream, paler to foot edge; mantle translucent, edge thickened strongly lobed with dark / black pigmentation on mantle edge aligning with rib interstices, faint pigmentation over cephalic folds. Shell (Fig. 63 L; Table S 9). Small sized (max sl = 4.76 mm, n = 1), circular ovate, height tall; apex weakly offset to posterior and left, apical sides convex except posterior which is concave; protoconch hooked to posterior, direction heterostrophic (n = 1), shell whorl dextral, shell thin; growth striae indistinct; multiple radial colour bands, protoconch area dark brown, central bands pale to dark, shell fringe pale brown; rib count (29, n = 1), predominance of primary ribs, few secondary ribs, weakly raised, off white and rounded, broaden to shell edge; paired primary ribs form siphonal ridge, ribs ends protrude weakly at shell lip, shell edge weakly scalloped and corrugated; rib interstices indistinct, darker. Interior shell margin to spatula mottled dark chocolate brown; outer shell margin paler with light brown rays aligning under ribs; siphonal groove distinct, same colour as spatula, darker than margin; ADM scar indistinct, CMS convex; no thickening of shell lip noted. Reproductive system (Fig. 64 E; n = 2). Positioned within right side of coelom, against foot wall on foot muscle, under the respiratory cavity; epiphallic parts positioned between RAM and BM. GA small, with singular GP through foot wall; AO small, bulbous, tip blunt, joined to upper GA; ED (incomplete) broad, joins to side of AO and GA; GA, AO, ED all white muscular fibrous tissue; BD and CD connect closely together into GA below AO junction, both long, narrow, slightly bent, pass closely together outside RAM (BD over CD) into soft white folded tissues of MG; BD smooth, featureless; CD with short lobed mid-section; CD connecting to large MG / AG complex; BC relatively small, elongate, thin whitish translucent test, embedded in MG / AG folds close to embedded SV; BD without distal loop or MA; HD small, coiled, links ducts in soft white folded tissues of AG to yellowish granulated HG; outer edge of MG lobed; AG larger than HG, sides match curvature of inner foot wall.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3A82BBFF68FF02FB75FDD6.taxon	diagnosis	Comparative remarks. In our mitochondrial phylogeny (Figs 1, 3), S. nusalikensis sp. nov. (plicata group, unit 89) is the sister species of a clade comprising S. guamensis (unit 70) and an unidentified species from Rarotonga known only from one DNA sequence (unit 17). Siphonaria nusalikensis differs from these species by genetic distances in COI of 11 % (S. guamensis) and 13.3 % (unit 17) (Table S 7). We found S. nusalikensis sp. nov. in sympatry with three congeners at New Ireland, PNG: For comparisons with S. normalis and S. viridis refer to comparative remarks under these species. Siphonaria recurva sp. nov. differs in having a larger shell with a less offset apex, a more prominent siphonal ridge and stronger edge scalloping, a similar BD, but with distal loop and a larger BC.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3A82BBFF68FF02FB75FDD6.taxon	distribution	Distribution and habitat. Recorded as exclusive to Nusalik Island (Big Nusa Island), Kavieng, PNG (Fig. 67). In this study, found on rocky shores, intertidal.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3A82BBFF68FF02FB75FDD6.taxon	etymology	Etymology. Named after the type locality of Nusalik Island (Big Nusa Island), Kavieng, PNG.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3A82B4FCCAFDC2FDF3FCF6.taxon	description	(Figs 63 M – O, 64 F – G)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3A82B4FCCAFDC2FDF3FCF6.taxon	materials_examined	Material examined. Type material. Holotype, from Island of Tubuai, Austral Islands, 23 ° 52 ’ S, 147 ° 41 ’ E, IRD-MNHN Stn. X 02 (MNHN IM- 2007 - 35319 p [SK 506], Fig. 63 M). Two paratypes, same data as holotype (MNHN IM- 2007 - 35317 p [SK 564], Fig. 63 N; IM- 2007 - 35318 p [SK 565], Fig. 63 O). Other, non-type material. Austral Islands: Tubuai, 23 ° 52 ’ S, 147 ° 41 ’ E, X 02 (MNHN IM- 2007 - 35316 p).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3A82B4FCCAFDC2FDF3FCF6.taxon	description	External morphology (preserved). Animal exterior evenly cream, foot sole grey, paler to foot edge; mantle translucent, irregular widely spaced pigmentation spots on foot wall, concentrated over posterior and centre of cephalic folds. Shell (Figs 63 M – O; Table S 9). Small (max sl mean = 15.5 mm, SD = 0.5 mm, n = 3), circular ovate; height low; apex offset weakly left and strongly to posterior, apical sides straight to weakly convex; shell edge uneven; protoconch direction undetermined, shell whorl dextral; growth striae indistinct, exterior grey to pale brown and usually eroded, radial colour banding indistinct; shell thickness medium; rib count (mean = 38, SD = 1.6, n = 3), ~ 12 distinct uneven primary ribs, white to pale, fairly straight, rib growth uneven, ridges rounded, broaden to scallop and protrude beyond shell edge; 2 – 3 interspersed pale white finer secondary ribs, rib interstices darker; paired primary ribs form siphonal ridge, more prominent and extend shell edge further than at other primary ribs. Interior shell dark chocolate brown, shell lip thickened; paler uneven rays extend from the shell lip over the shell margin, align under primary / secondary ribs; spatula dark chocolate brown, maybe mottled blueish; siphonal groove clear, similar colour to margin and spatula; ADM scar distinct, CMS convex. Reproductive system (Fig. 64 F; n = 1). Positioned within coelom under the respiratory cavity, hermaphroditic glands positioned to posterior against right foot wall and over foot sole, epiphallic parts positioned to anterior tightly between RAM and over back of BM. Join of AO, ED and GA together at top of GA, AO larger than GA, elongated, bluntly pointed, centrally bent, thicker than ED; ED wide, centrally bent, shorter and narrower than AO; AO, GA and ED all muscular white tissue; EG folded, block-like, flagellum F 1 small indistinct; BD and CD similar in thickness, connect side-by-side into GA, directly opposite to ED; both ducts smooth and pass together through outer side of RAM connecting into MG (BD over CD), BC bulbous, embedded in MG / AG; hermaphroditic glands inaccessible. Spermatophore (Fig. 64 G). Thread-like, test thin, translucent (incomplete); head section cylindrical (head length = 2.1 mm, n = 1), tip bulbous bluntly rounded, containing a white gelatinous core, smooth, featureless; 1 SPM tightly coiled embedded in white gelatinous mass in single BC.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3A82B4FCCAFDC2FDF3FCF6.taxon	diagnosis	Comparative remarks. Siphonaria cacao sp. nov. (atra group) is the only species found in Tubuai in this study. We were unable to sequence COI or 16 S from the available samples.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3A82B4FCCAFDC2FDF3FCF6.taxon	distribution	Distribution and habitat. Recorded exclusively from Austral Islands, French Polynesia, Pacific Ocean (Fig. 67). In this study, found on sheltered rocky shores (harbour at Island of Tubuai), intertidal.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3A82B4FCCAFDC2FDF3FCF6.taxon	etymology	Etymology. For the dark, chocolate-brown interior of the shell in this species; noun in apposition.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3582B4FF68FCC2FC69F916.taxon	description	(Figs 65 G – I, Q – R, 66 E – H)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3582B4FF68FCC2FC69F916.taxon	materials_examined	Material examined. Type material. Holotype, from Ouassé, nr Canala, 21 ° 30.346 ’ S, 166 ° 03.732 ’ E, NC; coll. B. W. Jenkins, NC 02 - 1, 22 Oct 2018 (AM C. 584786 p [M 377, SK 328], Fig. 65 G). Six paratypes, same data as holotype (AM C. 584800 p [SK 344], Fig. 65 H, C. 584908 p [SK 341], C. 585732 p [SK 343], C. 585923 d [SK 346], Fig. 65 I, C. 585924 p [SK 342], C. 585993 p [SK 347], Fig. 65 Q).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3582B4FF68FCC2FC69F916.taxon	description	External morphology. Foot sole, foot wall, cephalic folds and pneumostome cream, paler to foot edge; mantle translucent narrow, wider anteriorly, edge thickened strongly lobed with cream / white band; faint dark / black pigmentation on mantle edge aligning with rib interstices, dark pigmentation over centre of cephalic folds. Shell (Figs 65 G – I, Q; Table S 9): Small (max sl mean = 13.5 mm, SD = 1.02 mm, n = 6); ovate, often elongated, shape irregular, height medium; apex weakly offset sightly posterior and left, apical sides convex, protoconch direction weakly heterostrophic (n = 1, Fig. 65 Q), shell whorl dextral; growth striae indistinct, shell thick; rib count (mean = 35, SD = 2, n = 6), exterior uneven, radial shading bands faint, protoconch area pale; primary ribs white, secondary ribs darker, variably raised, crooked and bent, narrow and uneven in width, ridge rounded, extend unevenly slightly beyond shell lip to scallop and corrugate the shell edge; ribs unevenly spaced, 2 – 3 primary ribs form raised siphonal ridge; variable number secondary ribs between primary ribs, most rib interstices dark brown. Interior evenly dark, chocolate-brown, sometimes mottled paler in places, short white rays on shell margin align under primary / secondary ribs, siphonal groove distinct; ADM scar distinct, CMS straight, thickening and whitening of shell lip apparent in some specimens. Reproductive system (Figs 66 E, G; n = 3). Positioned within right side of coelom, against foot wall on foot muscle, under the respiratory cavity and digestive glands; epiphallic parts all white muscular fibrous tissue, reduced in size and positioned over BM (F 1 looped over front of BM), singular GP through foot wall; GA small indistinct; AO relatively small broad, bluntly pointed (embedded in folds of MG), joined to upper GA; ED elongated broad thickened, centrally twisted, joins to upper GA; EG small, soft whitish tissue, folded joins ED; single long narrow bent flagellum (F 1) as an extension of ED, of similar length. BD and CD connect in opposing directions into GA between ED join and GP, both ducts very narrow long straight smooth whitish, pass together through RAM (BD over thicker CD) into soft white folded tissues of MG; BC embedded in folds of AG / MG; BD with prominent distal twisted loop with MA attached to inner anterior foot wall above BM, bent immediately before BC; BC small bulbous, thin whitish translucent test; MG / AG complex relatively large, often separated (Fig. 66 E, G); HD thickened, brown markings, small coils, links ducts in soft white folded tissues of AG to yellowish granulated reddish spotted HG; HG larger than AG / MG. Spermatophore (Fig. 66 F, H). Test thin, translucent (length = 9.62 ± 0.08 mm, n = 2), head evenly cylindrical, tip bluntly rounded, containing a white gelatinous core, tapers along the transparent flagellum to a thin tip; both sections smooth, featureless; head wider and longer than flagellum (head length = 6.06 ± 1.11 mm, ~ 72 % of SPM length, head width = 103 ± 0 μm, flagellum width = 17 ± 0 μm, n = 2). 5 SPM tightly coiled and embedded in dark brown gelatinous mass in one BC.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3582B4FF68FCC2FC69F916.taxon	diagnosis	Comparative remarks. In our mitochondrial tree (Figs 1, 2), S. ouasseensis sp. nov. (atra group, Clade H, unit 48) forms a clade with S. bourailensis sp. nov. (unit 49), S. pravitas sp. nov. (unit 51) and S. scabra (unit 50) (Figs 1, 2). It differs from S. bourailensis sp. nov. by COI distances of ≥ 11.8 % and from other species by COI distances of ≥ 24 % (Table S 4). We have found this species in sympatry with three congeners in NC: Siphonaria bourailensis sp. nov. has fewer and broader ribs, a more prominent siphonal ridge, a larger, wider and pointed AO, and a larger ED. For comparisons with S. hienghenensis sp. nov. and S. atra refer to comparative remarks under these species.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3582B4FF68FCC2FC69F916.taxon	distribution	Distribution and habitat. Known only from NC (Fig. 70). In this study, found at Ouassé nr Canala in sheltered positions on exposed and sheltered rocky boulder and platform shores, mid to upper littoral level (Fig. 65 R).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3582B4FF68FCC2FC69F916.taxon	etymology	Etymology. Named after the type locality of Ouassé near Canala, NC.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3582B5FCCAF8A2FBCDFAF6.taxon	description	(Figs 65 J – L, O, S – T, 66 I – J)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3582B5FCCAF8A2FBCDFAF6.taxon	materials_examined	Material examined. Type material. Holotype, from Tiari, 20 ° 15.692 ’ S, 164 ° 24.664 ’ E, NC; coll. B. W. Jenkins, NC 04 - 3, 25 Oct 2018 (AM C. 584788 [M 364, SK 130 (RS + SPM)], Fig. 65 J). Paratype, same data as holotype (AM C. 585838 p [M 362], Fig. 65 L); seven paratypes, from Bonhomme de Bourail, La Roche Percee, 21 ° 36.487 ’ S, 165 ° 27.423 ’ E, NC; coll. B. W. Jenkins, NC 06 - 3, 28 Oct 2018 (AM C. 584799 5 p, C. 585015 p [M 366], Fig. 65 K, C. 585016 p [M 367]). Other, non-type material. NC: Tiari, 20 ° 15.692 ’ S, 164 ° 24.664 ’ E NC 04 - 3 (AM C. 585969 20 p, AM C. 584787 p [SK 060], C. 585839 p [M 363, SK 131], C. 585840 p [M 365], C. 585841 p [M 459]); Bonhomme de Bourail, La Roche Percee, 21 ° 36.487 ’ S, 165 ° 27.423 ’ E NC 06 - 3 (AM C. 584798 15 p); Presqu’ile de Ouano La Foa, 20 ° 51.434 ’ S, 165 ° 48.479 ’ E NC 06 - 4 (AM C. 595910 12 p).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3582B5FCCAF8A2FBCDFAF6.taxon	description	External morphology (Fig. 65 S). Foot sole, foot wall, cephalic folds and pneumostome pale grey, paler to foot edge; mantle translucent narrow, edge thickened, weakly lobed with black pigmentation bands aligning with rib interstices; black pigmentation more intense and darkest over centre of cephalic folds. Shell (Figs 65 J – L, T; Table S 9). medium sized (max sl mean = 17.5 mm, SD = 3.7 mm, n = 8); ovate, often elongated, shape irregular, height tall; apex offset slightly posterior and left, apical sides convex, posterior side straight to slightly concave, protoconch direction homostrophic (n = 1, Fig. 65 T), shell whorl dextral; growth striae prominent in bands, shell thick; rib count (mean = 44.4, SD = 5, n = 8), radial shading bands occur, protoconch area dark; primary ribs white, fairly straight often bent, narrow and uneven in width, ridge rounded, extend slightly beyond shell lip to scallop and corrugate the edge; ribs may be fairly evenly spaced, paired primary ribs forming siphonal ridge bounded with a wide interstice either side; 0 – 4 secondary ribs between primary ribs, most secondary ribs and rib interstices dark brown, smaller ribs occur in growth interstitial areas. Interior shell margin dark brown to tan, white rays align on shell margin under primary / secondary ribs, siphonal groove distinct, same colour as shell edge, points to right anterior; spatula dark chocolate brown to mottled tan even whitish; ADM scar distinct, CMS straight, paler than shell lip; thickening of shell lip translucent, infills and reduces lip scalloping, spatula becomes whitened. Reproductive system (Fig. 66 I; n = 2). Positioned within right side of coelom, against foot wall on foot muscle, under the respiratory cavity and digestive glands; epiphallic parts positioned over BM, with singular GP through foot wall; GA very small indistinct; AO relatively large elongated broad centrally bent bluntly pointed (embedded in folds of MG), joined to upper GA; ED elongated broad thickened, centrally bent, joins to lower side of GA; GA, AO, ED all white muscular fibrous tissue; EG large, soft whitish tissue, folded, joins ED and single narrow bent flagellum (F 1) as an extension of ED, shorter length. BD and CD connect in opposing directions into GA between ED join and GP, both ducts narrow long straight smooth whitish, pass together through RAM to outerside (BD over thicker CD) into soft white folded tissues of MG; BC embedded in folds of AG / MG; BD long narrow with prominent distal loop with MA connected to inner anterior foot wall above BM; BC medium bulbous, thin whitish translucent test; MG / AG complex relatively large; HD thickened coiled, links ducts in soft white folded tissues of AG to yellowish granulated HG; AG / MG larger than HG. Spermatophore (Fig. 66 J). Test thin, translucent (length = 8.93 mm, n = 1); head evenly cylindrical, bulbous, tip bluntly rounded, containing a white gelatinous core, tapers along the transparent flagellum to a thin tip; both sections smooth, featureless; head much wider and similar length to flagellum (head length = 4.65 mm, 52 % of SPM length, head width = 124 μm, flagellum width = 22 μm). 2 SPM tightly coiled embedded in brown gelatinous mass in one BC (AM C. 584788).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3582B5FCCAF8A2FBCDFAF6.taxon	diagnosis	Comparative remarks. In our mitochondrial phylogeny, S. caledonica sp. nov. (laciniosa group, unit 23) is the sister species of S. maloensis (see comparative remarks under this species) (Figs 1, 3). It differs from other species by COI distances of ≥ 19 % (Table S 6). We found S. caledonica sp. nov. in sympatry with five congeners on the E coast of NC: For comparisons with S. atra, S. hienghenensis sp. nov., and S. normalis refer to comparative remarks under these species. Siphonaria bourailensis sp. nov. has a shell with slightly more raised ribbing, greater edge scalloping, a paler spatula, and a shorter ED. Siphonaria poindimiensis has a shell with a more central apex, greater edge scalloping, paler spatula, a blunt AO, and a longer ED. The RS resembles that of S. atra and S. viridis. Specimens figured as ‘ laciniosa group, unit 23 ’ in Dayrat et al. (2014: 262) belong to this species.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3582B5FCCAF8A2FBCDFAF6.taxon	distribution	Distribution and habitat. Recorded from E and W coasts of NC (Fig. 70). In this study, found on exposed to sheltered rocky boulder and platform shores, mid to upper littoral level (Fig. 65 O).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3582B5FCCAF8A2FBCDFAF6.taxon	etymology	Etymology. For New Caledonia, where this species has been found; adjective.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3482B7FCCAFAC2FA02FC36.taxon	description	(Figs 68 A – C, K, 69 A – B)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3482B7FCCAFAC2FA02FC36.taxon	materials_examined	Material examined. Type material. Holotype, from Clifton Beach, Karachi, 24 ° 45.500 ’ N, 67 ° 05.968 ’ E, Pakistan; coll. S. Mushtaq, S. Amir and B. W. Jenkins, PA 02 - 1, 4 July 2018 (AM C. 585859 [M 233], Fig. 68 A). Paratypes: same data as holotype (AM C. 585891 17 p, C. 585104 p [M 232], Fig. 68 C, C. 585105 p [M 234], Fig. 68 B); French Beach Karachi, Pakistan; coll. S. Mushtaq, S. Amir and B. W. Jenkins, 24 ° 50.367 ’ N, 66 ° 49.387 ’ E PA 01 - 1 4 July 2018, (AM C. 585183 p [SK 187]). Other, non-type material. Pakistan: Karachi, Clifton Beach 24 ° 45.500 ’ N, 67 ° 05.968 ’ E PA 02 - 1, 4 July 2018 (AM C. 595985 [SK 569]); French Beach 24 ° 50.367 ’ N, 66 ° 49.387 ’ E PA 01 - 1 (AM C. 585894 3 p, C. 595949 p [SK 376]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3482B7FCCAFAC2FA02FC36.taxon	discussion	Taxonomic remarks. Sequences of a specimen of ‘ unit 10 ’ from Hikman Peninsula, Oman first published in Dayrat et al. (2014) have subsequently been referenced by Gonzàlez-Wevar et al. (2018) as ‘ Siphonaria sp. (Caroline Island) ’ under an identical GenBank registration number.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3482B7FCCAFAC2FA02FC36.taxon	description	External morphology. Foot sole, foot wall, cephalic folds and pneumostome evenly cream, without dark pigmentation; mantle narrower than foot wall, edge thickened, weakly lobed. Shell (Figs 68 A – C; Table S 9). Small sized (max sl mean = 4.6 mm, SD = 0.58 mm, n = 4), elongate ovate; height tall, medium thickness; apex offset strongly posterior and left, apical sides very flat to weakly convex, protoconch direction homostrophic (n = 2), apex curled to posterior, shell whorl dextral; growth striae indistinct; rib count (mean = 21, SD = 0.43, n = 4), primary ribs indistinct, pale white, slightly bent, broaden to shell lip; shell lip weakly corrugated, uneven; no distinct secondary ribs, rib interstices darker brown; siphonal ridge indistinct, radial shading bands absent. Interior shell lip white, pale to dark brown rays on shell margin to spatula align under primary / secondary ribs, siphonal groove indistinct, same colour as shell edge, points to right anterior; spatula golden brown to mottled brown; ADM scar distinct, CMS concave, paler than shell lip; thickening of shell lip occurs in some specimens; infills and reduces lip corrugations. Reproductive system (Fig. 69 A; n = 3). Positioned within right side of coelom, against foot wall on foot muscle, under the respiratory cavity and intestine; epiphallic parts positioned between BM and RAM. GA very small with singular GP through foot wall; AO small, narrow, bluntly pointed, joined to inner side of GA; ED short, elongated, centrally bent, joins to outer side of GA; GA, AO, ED all white muscular fibrous tissue; EG medium, soft whitish tissue, slightly folded, joins ED; single centrally folded broad flagellum (F 1), shorter but similar width to ED, appears as an extension of ED; BD and CD connect closely but in opposing directions into GA between ED join and GP; CD short, broad, slightly bent, smooth, whitish, featureless; BD long, narrow, featureless; both ducts pass closely together through outer side of RAM (BD over CD); MG / AG complex medium; CD connecting to ducts, BC embedded in folds of AG / MG; BD without distal loop and MA; BC relatively large, spherical, thin whitish translucent test; HD short, thickened, coiled, links ducts in soft white folded tissues of AG to yellowish granulated HG; outer edge of MG lobbed; AG / MG larger than HG, sides match curvature of inner foot wall. Spermatophore (Fig. 69 B). Thread-like (length = 3.07 ± 0.03 mm, n = 2), translucent, test thin; head section bluntly rounded, evenly cylindrical, containing a core white gelatinous mass, tapers along the transparent flagellum to a thin tip; both sections smooth, featureless. Head section longer wider than flagellum (head length = 1.47 ± 0.06 mm, ~ 48 % of SPM length, head width = 69 ± 0 μm, flagellum width = 17 ± 0 μm, n = 2); 2 SPM tightly coiled embedded in whitish gelatinous mass in BC [SK 562].	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3482B7FCCAFAC2FA02FC36.taxon	diagnosis	Comparative remarks. Siphonaria perexigua sp. nov. (normalis group, unit 10) forms a well-differentiated lineage in the mitochondrial tree (Clade E, Figs 1, 4). It differs from other species by COI distances of ≥ 18 % (Table S 8). Specimens from Pakistan examined and sequenced herein form a sub-clade with an unidentified species from Oman (Siphonaria sp. ‘ unit 10 ’) published by Dayrat et al. (2014). We have not examined the specimen from Oman but presume that it is conspecific based on the close genetic relationships with the samples from Pakistan. In Pakistan, we found S. perexigua sp. nov. in sympatry with four other congeners: For comparisons refer to comparative remarks under these species (S. asghar, S. belcheri, S. kurracheensis, and S. crenata). Bosch et al. (1995: 185) identified individuals of this species as S. compressa Allanson, 1958. However, S. compressa (not reviewed herein) differs in shell geometry (fragile shell, apex weakly offset, not overlapping posterior shell edge; see Chambers & McQuaid, 1994 a: 266, fig. 1 A) and distribution (west coast of S. Africa; Chambers & McQuaid, 1994 a) from the the specimens figured in Bosch et al. (1995: fig. 861).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3482B7FCCAFAC2FA02FC36.taxon	distribution	Distribution and habitat. Recorded from Karachi, Pakistan and Hikman Peninsula, Oman (Fig. 70). In this study, found in sheltered positions on moderately exposed rocky shores, mid littoral level, amongst small barnacles (Fig. 68 K).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3482B7FCCAFAC2FA02FC36.taxon	etymology	Etymology. From ‘ perexiguus’ (Latin = very small), for the very small shell size of this species; adjective.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3682B1FCCAFB82FF77F7F6.taxon	description	(Figs 68 D – E, L – N, 69 C – D)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3682B1FCCAFB82FF77F7F6.taxon	materials_examined	Material examined. Type material. Holotype, from Dolokoan Beach, 8 ° 31.424 ’ S, 125 ° 37.091 ’ E, N of Dili, Timor-Leste; coll. B. W. Jenkins, TL 01 - 1, 14 July 2019 (AM C. 584826 [M 446, SK 229 (RS + SPM)], Fig. 68 D). Paratypes: same data as holotype (AM C. 585443 10 + p, C. 585442 p [M 451, SK 165], Fig. 68 E). Other, non-type material. Timor-Leste: Dolokoan Beach N of Dili, 8 ° 31.424 ’ S, 125 ° 37.091 ’ E TL 01 - 1, (AM C. 585925 p [SK 231]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3682B1FCCAFB82FF77F7F6.taxon	description	External morphology. Foot sole dark grey, foot edge cream; foot wall, cephalic folds and pneumostomal lobe evenly grey; black pigment shading at join of foot wall and mantle, concentrated over cephalic folds; mantle thin, narrower than foot wall, edge lobed with black bands aligning with shell rib interstices and interstice width; pneumostome under mantle between right adductor muscles. Shell (Figs 68 D – E, L; Table S 9). Small sized (max sl mean = 8.3 mm, SD = 0.47 mm, n = 2), ovate; low, fragile, thin; apex offset strongly posterior and to left, apical sides strongly convex, protoconch direction weakly homostrophic to central (n = 1, Fig. 68 L), below apex, curls to posterior, shell whorl dextral; growth striae indistinct, shell edge uneven, faintly scalloped; rib count (mean = 33, SD = 4, n = 2), primary ribs flat, pale white, fairly straight bent in places, broaden to shell lip, 2 – 3 secondary ribs between primary ribs, interstices dark brown; paired primary ribs form siphonal ridge, usually not prominent, may appear raised, protoconch area darker; Interior shell margin white, dark chocolate brown rays extend from shell lip to spatula appearing paired, aligning under rib interstices; spatula and indistinct siphonal groove evenly dark chocolate brown; ADM scar indistinct, CMS convex; thickening of shell lip occurs, infills and reduces lip scalloping. Shell resembles other small dark normalis group. Reproductive system (Fig. 69 C; n = 2). Positioned within entire right side of coelom, against foot wall on foot muscle, under the respiratory cavity and intestine occupying large proportion of animal body volume; epiphallic parts positioned over BM. GA medium, with singular GP through foot wall; AO very small, narrow, bluntly pointed, joined to lower ED and upper GA; ED very short, broad, centrally bent, joins to side of GA; GA, AO, ED all white muscular fibrous tissue; EG very large, soft whitish tissue, slightly folded, joins ED; single very broad flagellum (F 1), similar length and width to ED, appears as an extension of ED, EG and F 1 joined in parallel; BD and CD connect side-by-side into GA between ED join and GP, both ducts short, slightly bent, smooth, thickened, whitish, featureless, pass closely together between outside RAM and inner foot wall (BD over CD) into soft white folded tissues of MG; MG / AG complex medium in size; CD connecting to ducts, BC embedded in folds close to embedded blackish SV; BD without distal loop and MA, with loop immediately prior to BC; BC relatively large, spherical, thin whitish translucent test, (4 SPM in holotype AM C. 584826 TL 01 - 1 [M 446, SK 229]); HD short, narrow, coiled, links ducts in soft white folded tissues of AG to yellowish granulated HG; outer edge of MG lobbed; AG larger than HG, sides match curvature of inner foot wall. Spermatophore (Fig. 69 D). Broad head with short flagellum (length = 1.71 ± 0.08 mm, n = 2); head section cylindrical, bulbous, rounded tip; test thin, smooth, featureless, translucent encasing a white opaque central core; short looped tapering section merges head to filamentous flagellum; head slightly shorter, wider than translucent flagellum (head length = 1.16 ± 0.17 mm, flagellum length = 0.55 ± 0.088 mm, ~ 68 % of SPM length, head width = 119 ± 8 μm, flagellum width = 11 ± 0 μm, n = 2); 4 SPMs tightly coiled in BC (AM C. 584826 holotype [M 446, SK 229]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3682B1FCCAFB82FF77F7F6.taxon	diagnosis	Comparative remarks. In our mitochondrial phylogeny (Figs 1, 4), S. planucosta sp. nov. (normalis group, unit 79) is the sister species of S. radiata (unit 12). Siphonaria planucosta sp. nov. differs from S. radiata by COI distances of ≥ 18.4 % (Table S 8). We found S. planucosta sp. nov. in sympatry with five congeners in TL. For comparisons with S. forticosta sp. nov., S. alba, S. javanica, S. viridis, and S. campestra sp. nov. refer to comparative remarks under these species.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3682B1FCCAFB82FF77F7F6.taxon	distribution	Distribution and habitat. Recorded from Dolokoan Beach, N of Dili, Timor-Leste (Fig. 70). In this study, found at sheltered positions (mainly in crevices) on exposed shore boulders, at mid littoral levels (Fig. 68 L).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3682B1FCCAFB82FF77F7F6.taxon	etymology	Etymology. From ‘ planus’ (Latin = level, even), for the level, even primary ribs on the shell of this species; adjective.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3082B2FCCAFD62FC8AF816.taxon	description	(Figs 68 F – J, O – P, 69 E – F)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3082B2FCCAFD62FC8AF816.taxon	materials_examined	Material examined. Type material. Holotype, from Bonhomme de Bourail, La Roche Percee, 21 ° 36.487 ’ S, 165 ° 27.423 ’ E, NC; coll. B. W. Jenkins, NC 06 - 3, 28 Oct 2018 (AM C. 585017 [M 368], Fig. 68 F). Paratypes: same data as holotype (AM C. 608182 p [SK 126], Fig. 68 J, C. 585466 p [M 372] Fig. 68 H); We Baie de Chateaubriand, East coast Lifou, Loyalty Islands, NC, 20 ° 54.779 ’ S, 167 ° 15.636 ’ E; coll. B. W. Jenkins, LFU 01 - 1, 22 Oct 2018 (AM C. 585438 p [M 390], Fig. 68 I). Other, non-type material. NC: Poum 2, 20 ° 13.754 ’ S, 164 ° 01.699 ’ E, NC 05 - 3 (AM C. 586000 4 p); Ponerihouen, 21 ° 05.644 ’ S, 165 ° 26.646 ’ E, NC 03 - 1 (AM C. 584807 p [M 360]); Bonhomme de Bourail, La Roche Percee, 21 ° 36.487 ’ S, 165 ° 27.423 ’ E, NC 06 - 3 (AM C. 585331 p [SK 125 protoconch I 2]); Presq’ile Ducos Baie des Dames, Noumea, 22 ° 14.170 ’ S, 166 ° 24.524 ’ E, NC 01 - 1 (AM C. 584804 4 p).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3082B2FCCAFD62FC8AF816.taxon	description	External morphology (Fig. 68 O). Foot sole and foot wall evenly cream, without black pigmentation, paler to foot edge and mantle edge; even blackpigmented shading along mantle and foot wall join, black pigmentation darkest over centre of cephalic folds; mantle narrow, edge thickened, weakly lobed without black pigmentation; pneumostomal lobe under mantle, narrow and unpigmented. Shell (Figs 68 F – J; Table S 9). Small to medium sized (max sl mean = 15.1 mm, SD = 2.4 mm, n = 5), ovate; height low to medium; apex offset strongly posterior and left pointing to posterior, apical sides convex, protoconch direction homostrophic (n = 3, Fig. 68 J), shell whorl dextral; growth striae notable uneven, shell thin, rib count (mean = 41, SD = 5.7, n = 5), primary ribs white, fairly straight, increasingly widen to shell lip, weakly extend beyond shell lip; interstices pale brown to black,; secondary ribs indistinct and fewer than primary; paired primary ribs on siphonal ridge, more prominent than other primary ribs; dark patches with 3 – 4 secondary ribs either side of siphonal ridge. Interior shell margin and spatula dark chocolate brown to black, primary and secondary ribs marked by short white rays often extending over shell margin, siphonal groove distinct and weakly bent; shell edge scalloped, not corrugated, swollen rounded; ADM scar distinct, CMS straight, paler than shell lip. Thickening and dark brown / black of shell lip common. Reproductive system (Fig. 69 E; n = 1). Positioned within right side of coelom, against foot wall on foot muscle, under the respiratory cavity; epiphallic parts lie over back of BM; GA very small, singular GP through foot wall; AO very small elongate, tip blunt; ED relatively large, broad, bent, joins to side of GA; single flagellum (F 1) long broad, as long as ED, appears as an extension of ED; GA, AO, ED all white muscular fibrous tissue; EG large, soft whitish tissue, slightly folded; BD and CD short, broad, jointly connect into upper end of GA; BD without any distal loop, longer slightly narrower than CD; both ducts smooth featureless, pass closely together through outer side of RAM (BD over CD) into folded, soft white tissues of large MG complex; BC very large, bulbous, thin translucent test, embedded in MG folds close to embedded SV; HD large, coiled, links AG to small yellowish granulated HG; AG larger than HG, sides match curvature of inner foot wall. Spermatophore (Fig. 69 F). Body cylindrical, relatively short (length = 3.68 ± 0.87 mm, n = 2); test thin, translucent; head bulbous, tip bluntly rounded, wide, evenly cylindrical, containing a core white gelatinous mass, tapers along the transparent flagellum to a thin tip; both sections smooth, featureless. Head section slightly shorter but much wider than flagellum (head length = 1.95 ± 0.37 mm, ~ 55 % of SPM length; head width = 119 ± 8 μm, flagellum width = 13 ± 0.0 μm, n = 2); 6 SPM tightly coiled in one bursa embedded in brown gelatinous mass.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3082B2FCCAFD62FC8AF816.taxon	diagnosis	Comparative remarks. In our mitochondrial phylogeny, S. bourailensis sp. nov. (atra group, unit 49) is the sister species of S. ouassensis sp. nov.; for comparison refer to comparative remarks under that species. We found S. bourailensis sp. nov. in sympatry with seven congeners in NC and on Lifou, Loyalty Islands: For comparison with S. atra (unit 41), S. hienghenensis sp. nov. (unit 40), S. monticulus (unit 57), S. normalis (unit 14), S. ouasseensis sp. nov. (unit 48), S. caledonica sp. nov. (unit 23), and S. viridis (unit 25) refer comparative remarks under these species. Specimens of S. bourailensis sp. nov. on Lifou slightly differ from those on NC in having a smaller shell with wider and fewer primary ribs and a golden-brown internal colouration.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3082B2FCCAFD62FC8AF816.taxon	distribution	Distribution and habitat Recorded as endemic to NC and Lifou (Fig. 70). In this study, found on exposed to sheltered rocky boulder and platform shores, mid to upper littoral levels (amongst and above oysters) (Fig. 68 P).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3082B2FCCAFD62FC8AF816.taxon	etymology	Etymology Named after type locality of Bonhomme du Bourail, NC.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3382B3FCCAFF02FE42F9B6.taxon	description	(Figs 69 G – H, 71 A – B, M – N, S)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3382B3FCCAFF02FE42F9B6.taxon	materials_examined	Material examined. Type material. Holotype, from Laings Point, 33 ° 50.419 ’ S, 151 ° 16.638 ’ E, Sydney Harbour, NSW, Australia; coll. B. W. Jenkins, NSW 06 - 3, 19 April 2018 (AM C. 585040 [M 192], Fig. 71 A [SK 118 (RS and SPM)], Fig. 69 G). Paratypes same data as holotype (AM C. 585334 p; C. 585038 p [M 153], Fig. 71 B; C. 585039 p [M 166]). Other, non-type material. Australia, LHI: Signal Point, 31 ° 31.501 ’ S, 159 ° 03.580 ’ E, LHI 2017 Apr 04 - 099 (AM C. 608190 p [M 010]), LHI 2017 Apr 04 - 104 (AM C. 585939 p). NSW: Sydney, Spit Bridge, 33 ° 48.198 ’ S, 151 ° 14.860 ’ E (AM C. 546769 4 p, C. 546766 p [SK 432 protoconchI 4]); LaingsPoint, 33 ° 50.419 ’ S, 151 ° 16.638 ’ E, NSW 06 - 3 (AM C. 585033 p [M 191]); Spit Bridge 2, 33 ° 48.270 ’ S, 151 ° 14. 520 ’ E (AM C. 546764 p; C. 546767 2 p; C. 608191 p, C. 595942 p [SK 186 protoconch I 3]); 100 m NW of Spit Bridge, 33 ° 48.210 ’ S, 151 ° 14.664 ’ E (AM C. 608194 p).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3382B3FCCAFF02FE42F9B6.taxon	description	External morphology (Fig. 71 N). Foot sole, foot wall and mantle smooth, evenly cream to yellow, paler at foot edge; regular vertical uneven bands / stripes on foot wall, roughly align with rib interstices, fades to none at join of foot wall and mantle; narrow cream pneumostomal lobe under mantle; black pigmentation heavy dark over front of cephalic folds; foot wall shows black blotches; fringing mantle translucent, narrow, covers shell mantle, outer edge thickened, lobed, none to slight black pigmentation aligning with rib interstices; pneumostomal lobe within mantle between the right ADMs, closes the pneumostomal and anal openings at the mantle edge; two small black epithelial eye spots centralised on two thick brownish yellow centrally touching cephalic folds; genital pore inconspicuous, located on foot wall to right anterior of right cephalic fold. Shell (Figs 71 A, B, S; Table S 3). Size medium (max sl mean = 18.03, SD = 0.87, n = 5), elongate ovate; low; apex offset central sightly to left, apical sides straight, posterior concave; protoconch direction homostrophic (n = 3; Fig. 71 S), shell whorl dextral, growth striae uneven, distinct, protoconch area dark brown; shell thick; rib count (mean = 38.8, SD = 3.3, n = 5), 12 – 16 primary ribs pale white, bent, crooked to shell lip, rib ridges rounded, increasingly raised and broaden to shell edge, weakly protrude beyond shell lip to unevenly scallop and corrugate the edge; 3 - 4 interspersed pale white less raised secondary ribs, rib interstices brown; paired primary ribs on siphonal ridge. Interior shell margin dark to golden brown, white / cream rays under primary and secondary ribs, extend to dark chocolate brown blotched white spatula; siphonal groove distinct, whitish; ADM scar distinct, chocolate brown to whitish, CMS straight; no thickening of shell lip noted; pale variety may occur. Reproductive system (Fig. 69 G; n = 2). Positioned within coelom under the respiratory cavity, hermaphroditic glands positioned to posterior against right foot wall and over foot sole, epiphallic parts positioned to anterior between BM and RAM. Join of AO and GA indistinct, AO elongated, bluntly pointed, centrally bent with MA, much larger than GA, much thicker than ED; ED relatively long, slightly twisted, narrow; EG white, folded, elongated; single flagellum F 1 on EG, long, looped, narrow; AO, GA and ED all muscular white tissue; BD and CD with opposing connections to GA between ED, AO and GP; BD slightly longer and thinner than CD with a prominent loop on anterior side, both ducts smooth and pass together through RAM connecting into MG (BD above CD), BC translucent, white test, mid-sized and bulbous; HD short, coiled, links AG to a small elongated narrow brownish finely granulated HG, inner edge firmly moulded; MG and AG small, folded, soft white tissue; purple SV embedded on left side of AG, AG larger than HG, sides match curvature of inner foot wall at right posterior quarter of coelom. Spermatophore (Fig. 69 H). Long cylindrical, body long; test thin, short tapering section merging head with filamentous flagellum; head bluntly rounded, central whitE, WAvy core, longer thicker than translucent flagellum (head length = 7.09 mm, head width = 160 μm n = 1; flagellum incomplete); 2 tightly coiled SPMs in bursa of one specimen (AM C. 585040).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3382B3FCCAFF02FE42F9B6.taxon	diagnosis	Comparative remarks. In our mitochondrial phylogeny (Figs 1, 2), S. pravitas sp. nov. (atra group, unit 51) is the sister species of S. scabra (unit 50). Both species differ by COI distances of ≥ 8.2 %. From any other species S. pravitas differs by COI distances of ≥ 23.5 % (Table S 4). Siphonaria scabra differs by having a higher shell with less raised ribs and less developed edge scalloping, a larger AO, longer ED, smaller BC, a longer, narrower BD with distal loop, a longer SPM. We found S. pravitas sp. nov. in sympatry with eight congeners throughout its range. Six congeners are sympatric in Sydney Harbour, NSW: For comparisons with S. stowae, S. denticulata, S. diemenensis, S. funiculata, S. scabra, and S. zelandica refer to comparative remarks under these species. Two congeners are sympatric in LHI: Refer to comparative remarks under S. lentula and S. exulum.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3382B3FCCAFF02FE42F9B6.taxon	distribution	Distribution and habitat. Recorded as endemic to SE Australia (Fig. 73). In this study, found on sheltered rocky shores, at upper to mid littoral levels (Fig. 71 M).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3382B3FCCAFF02FE42F9B6.taxon	etymology	Etymology. From ‘ pravitas’ (Latin = ‘ crookedness, inequality, irregularity or deformity’) referring to the crooked waviness of the radial ribs on the shell of this species; noun in apposition.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3282B3FF68F902FB23F816.taxon	description	(Figs 71 C, 72 A)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3282B3FF68F902FB23F816.taxon	materials_examined	Material examined. Type material. Holotype, from NW side of Big Nusa Is., 02 ° 34.1 ’ S, 150 ° 46.7 ’ E, New Ireland, PNG; coll. KAVIENG 2014 expedition, KM 21, 28 Jun. 2014 (MNHN IM- 2013 - 55336 [M 534], Fig. 71 C).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3282B3FF68F902FB23F816.taxon	description	External morphology (preserved). Animal exterior evenly cream, foot sole darker, paler to foot edge; irregular black pigmentation on foot wall. Shell (Fig. 71 C; Table S 9). Small sized (max sl = 15.3 mm, n = 1), circular ovate, height tall; apex noticeably offset to posterior and left, hooked to posterior, apical sides convex, posterior straightening, protoconch direction homostrophic (n = 1, MNHN IM- 2013 - 55336 holotype), shell whorl dextral, shell thin; growth striae prominent and slightly raised; multiple radial colour bands, protoconch area dark brown, central band pale, shell fringe dark brown; rib count (35, n = 1), primary ribs far more prominent than secondary ribs; ~ 17 pale brown to off white primary ribs, ridges raised and rounded, broaden to shell edge; paired primary ribs form siphonal ridge, most primary ribs protrude strongly beyond shell lip with ends slightly raised to strongly scallop and corrugate shell edge; 0 – 3 finer secondary ribs between primary ribs, rib interstices darker. Interior shell margin and spatula dark chocolate brown; off white to cream rays on shell margin align under ribs, siphonal groove distinct, same colour as spatula, paler than margin; ADM scar distinct, CMS convex; no thickening of shell lip noted. Reproductive system (Fig. 72 A; n = 1). Positioned within right side of coelom, against foot wall on foot muscle, under the respiratory cavity; epiphallic parts lie over back of BM; GA medium, singular GP through foot wall; AO bulbous wide, tip blunt; ED short, wide, bent, joins to side of GA; single flagellum (F 1) short, twisted, as long as ED, appears as an extension of ED; GA, AO, ED all white muscular fibrous tissue; EG small, soft whitish tissue, slightly folded; BD and CD long, narrow, jointly connect into upper end of GA; BD with distal loop and MA, longer, centrally narrower than CD but wider at GA and MG junctions; both ducts smooth featureless, pass closely together through RAM (BD over CD) into folded, soft white tissues of large MG / AG complex; BC large, bulbous, thin translucent test, embedded in MG folds close to embedded SV; HD small, lobed, coiled, links AG to small yellowish granulated HG; AG larger than HG, sides match curvature of inner foot wall.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3282B3FF68F902FB23F816.taxon	diagnosis	Comparative remarks. In our mitochondrial tree (Figs 1, 2), S. recurva sp. nov. (atra group, unit 32) is the sister species of S. incerta (unit 72). Both species differ by COI distances of ≥ 16.7 %. From other species S. recurva differs by distances of ≥ 23.3 % (Table S 3). We found S. recurva sp. nov. in sympatry with two congeners in New Ireland, PNG. For comparisons with S. nusalikensis sp. nov. and S. normalis refer to comparative remarks under these species. The specimen figured as ‘ atra group, unit 32 ’ in Dayrat et al. (2014: fig. 5 I) is conspecific.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3282B3FF68F902FB23F816.taxon	distribution	Distribution and habitat. Recorded from Big Nusa Island, Kavieng, New Ireland, PNG and Tutuila, American Samoa, Western Pacfic Ocean (Fig. 73). Found on intertidal platforms and limestone cliffs.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF3282B3FF68F902FB23F816.taxon	etymology	Etymology. Derived from Latin adjective “ recurvus ” meaning ‘ curving back, crooked’ referring to the shape of the shell’s prominent protoconch.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2D82AEFF68FF02FC00FC36.taxon	description	(Figs 71 D – H, O – P, 72 B – D)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2D82AEFF68FF02FC00FC36.taxon	materials_examined	Material examined. Type material. Holotype, from Chinamans Rock, 27 ° 42.776 ’ S, 114 ° 09.361 ’ E, Kalbarri, WA; coll. B. W. Jenkins, WA 52 - 1, 20 Nov 2017 (WAM S 74049 [M 400], Fig. 71 D). Paratypes same as holotype (WAM S 74079 6 p, S 74048 p [M 100], Fig. 71 E, AM C. 584731 11 p, C. 585200 p [M 313], Fig. 71 F, C. 585919 p [SK 150], Fig. 71 G). Other, non-type material. Australia, WA: Chinamans Rock, Kalbarri, 21 ° 54.739 ’ S, 113 ° 58.706 ’ E; coll. B. W. Jenkins, WA 52 - 1, 20 Nov 2017 (WAM S 74047 p [M 099], Fig. 54 D, S 4078 p [SK 050]); Thompson Bay, Rottnest Is, RI 012 (AM C. 585201 p [SK 381], Fig. 54 E, C. 585202 d [SK 382], C. 595909 d [R 2004]); Tantabiddi, 21 ° 54.739 ’ S, 113 ° 58.706 ’ E, WA 42 - 1 (AM C. 585656 5 p, WAM S 74045 p [SK 036]); Cape Latouche Treville nr Gourdon Bay, 18 ° 27.101 ’ S, 121 ° 48.911 ’ E, WA 27 - 2 (AM C. 584687 5 p, WAM S 74135 5 p); 18 ° 27.457 ’ S, 121 ° 48.725 ’ E, WA 27 - 1 (AM C. 584711 5 p, WAM S 74136 2 p); Cape Keraudren, 19 ° 57.393 ’ S, 119 ° 46.358 ’ E, WA 29 - 2 (AM C. 584704 5 p, WAM S 74138 2 p); Withnell Bay Dampier, 20 ° 35.106 ’ S, 116 ° 47.252 ’ E, WA 33 - 1 (AM C. 584752 5 p, WAM S 74141 5 p); Point Samson, 20 ° 36.655 ’ S, 117 ° 11.472 ’ E, WA 32 - 2 (AM C. 584781 8 p, WAM S 74140 8 p); 20 ° 36.684 ’ S, 117 ° 11.303 ’ E, WA 32 - 1 (AM C. 584751 5 p, WAM S 74139 5 p); Dampier Hbr, 20 ° 39.920 ’ S, 116 ° 42.134 ’ E, WA 33 - 3 (AM C. 585575 3 p); Gnoorea Point, 1 20 ° 50.560 ’ S, 116 ° 21.804 ’ E, WA 36 - 1 (AM C. 584689 3 p, WAM S 74142 2 p); Beardon Pt Onslow, 21 ° 37.860 ’ S, 115 ° 06.573 ’ E, WA 37 - 1 (AM C. 584722 5 p, WAM S 74143 4 p); NW Cape Exmouth, 21 ° 48.360 ’ S, 114 ° 07.665 ’ E, WA 41 - 1 (AM C. 584698 6 p, WAM S 74144 3 p); Fly Island, Great Sandy Islands, 21 ° 48 ’ 19.548 ’ ‘ S, 114 ° 33 ’ 9.36 ’ ‘ E (WAM S 97276 p); Tantabiddi, 21 ° 54.739 ’ S, 113 ° 58.706 ’ E, WA 42 - 1 (AM C. 584690 5 p, d; WAM S 74145 2 p); Pt S of Bruboodjoo Pt Bateman Bay, 23 ° 02.991 ’ S, 113 ° 49.371 ’ E, WA 43 - 1 (AM C. 584732 12 p, WAM S 74146 5 p); Point Maud, 23 ° 08.322 ’ S, 113 ° 46.294 ’ E, WA 44 - 1 (AM C. 584753 5 p, C. 585919 p [SK 150], WAM S 74147 5 p); Coral Bay, S of Exmouth Gulf, 23 ° 08.322 ’ S, 113 ° 46.294 ’ E (AM C. 595919 p [SK 004], C. 959956 p [SK 001]); N of Point Quobba, 24 ° 26.288 ’ S, 113 ° 24.204 ’ E, WA 45 - 2 (AM C. 584727 7 p, WAM S 74149 5 p); Point Quobba, 24 ° 29.124 ’ S, 113 ° 24.501 ’ E, WA 45 - 1 (AM C. 584754 5 p, WAM S 74148 5 p); Bottle Bay Cape Peron, 25 ° 32.566 ’ S, 113 ° 29.467 ’ E, WA 49 - 1 (AM C. 584755 5 p, WAM S 74150 5 p); Islet off Eagle Bluff, Shark Bay, Peron Peninsula, 26 ° 05.5 ’ S, 113 ° 34.4 ’ E (WAM S 72339 6 p); Whalebone, 26 ° 07.835 ’ S, 113 ° 38.391 ’ E, WA 47 - 2 (AM C. 585657 5 p); Kells Rock Shark Bay, 26 ° 10.473 ’ S, 113 ° 12.415 ’ E, WA 50 - 3 (AM C. 584734 12 p, WAM S 74151 6 p); Pepper Point (Zuytdorp), 26 ° 23.826 ’ S, 113 ° 18.268 ’ E, WA 51 - 1 (AM C. 584725 6 p, d, WAM S 74152 6 p); Chinamans Rock, Kalbarri, 27 ° 42.776 ’ S, 114 ° 09.361 ’ E (AM C. 585489 p [M 053], C. 585490 p [M 398], C. 585491 p [M 399], C. 585920 p [SK 151]); Red Bluff, 27 ° 44.627 ’ S, 114 ° 08.576 ’ E, WA 52 - 2 (AM C. 595908 p [M 125], C. 585365 p [M 126]); Horrocks, 28 ° 21.469 ’ S, 114 ° 24.751 ’ E, WA 53 - 1 (AM C. 584680 2 p; WAM S 74153 2 p); Turtle Bay East Wallabi Is, 28 ° 25.804 ’ S, 113 ° 44.538 ’ E, WA 55 - 1 (AM C. 584758 5 p; WAM S 74155 5 p); Houtman Abrolhos Islands, 28 ° 25.804 ’ S, 113 ° 44.538 ’ E (AM C. 595971 2 p), S of Fish Point East Wallabi Is, WA 55 - 2 28 ° 25.816 ’ S, 113 ° 44.633 ’ E (AM C. 584720 7 p; WAM S 74156 2 p); Cape Burney Geraldton, 28 ° 52.084 ’ S, 114 ° 38.056 ’ E, WA 54 - 1 (AM C. 584749 5 p; WAM S 74154 5 p); Leander Point Port Denison, 29 ° 16.568 ’ S, 114 ° 54.858 ’ E, WA 57 - 1 (AM C. 584760 5 p, WAM S 74157 5 p); S end Leander Point Port Denison, 29 ° 16.725 ’ S, 114 ° 54.918 ’ E, WA 57 - 2 (AM C. 585517 d); Freshwater Point, 29 ° 36.256 ’ S, 114 ° 58.464 ’ E, WA 57 - 3 (AM C. 584693 3 p, C. 585009 p [SK 063], WAM S 74158 2 p); Illawong Bch rocks, 29 ° 42.198 ’ S, 114 ° 57.551 ’ E, WA 57 - 4 (AM C. 584683 2 p, WAM S 74159 2 p); Illawong, 29 ° 42.254 ’ S, 114 ° 57.542 ’ E, WA 57 - 5 (AM C. 584694 3 p; WAM S 74160 2 p); Jurien Bay, 30 ° 17.244 ’ S, 115 ° 02.482 ’ E, WA 58 - 1 (AM C. 584679 2 p; WAM S 74161 p); Grey, 30 ° 39.968 ’ S, 115 ° 08.072 ’ E, WA 58 - 2 (AM C. 585366 p); Cape Leschenault, 31 ° 17.508 ’ S, 115 ° 27.089 ’ E, WA 58 - 3 (AM C. 584712 5 p, WAM S 74162 5 p); Quinns Rock, 31 ° 39.822 ’ S, 115 ° 41.345 ’ E, WA 58 - 4 (AM C. 585368 p); Longreach Bay Point, Rottnest Is, 31 ° 59.333 ’ S, 115 ° 32.063 ’ E RI 01 (AM C. 585600 4 p, C. 585198 p [M 311], C. 585199 p [M 312], C. 585200 p [M 313], C. 584943 p [SK 154], Fig. 71 H); Point Brown Swan River, 32 ° 02.344 ’ S, 115 ° 45.471 ’ E, WA 59 - 5 (AM C. 585369 p [M 128]); Fremantle Hbr, breakwater, 32 ° 03.342 ’ S, 115 ° 43.987 ’ E, WA 58 - 5 (AM C. 585519 d); Yallingup, 33 ° 38.358 ’ S, 115 ° 01.481 ’ E, WA 60 - 9 (AM C. 584724 6 p, WAM S 74163 5 p); Cowaramup Point, 33 ° 51.934 ’ S, 114 ° 58.904 ’ E, WA 60 - 3 (AM C. 585370 p); Sarge Bay Cape Leeuwin, WA 60 - 4 (AM C. 585809 p); Yanchep, sheltered limestone cliffs, 34 ° 32.783 ’ S, 115 ° 37.962 ’ E (AM C. 595972 p).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2D82AEFF68FF02FC00FC36.taxon	description	External morphology (Fig. 71 O). Foot wall usually without irregular black blotches, may be weakly present in some individuals; edge, pneumostomal lobe and cephalic folds all evenly brownish yellow, without any black / darker pigmentation markings, foot sole maybe dark brown / grey, foot edge paler; mantle translucent to transparent, as wide as foot wall, weakly lobed, aligns with undulations of primary shell ribs; the mantle edge thickened with yellowish band, may show dark pigmentation markings aligned with rib interstices; pneumostome wide between right adductor muscles and within mantle; cephalic folds thickened. Shell (Figs 71 D – H, P; Table S 9). Medium sized (max sl mean = 17.5 mm SD = 2.7 mm, n = 19); height low to medium; ovate; apex central, apical sides weakly convex; protoconch direction weakly homostrophic (n = 1, Fig. 71 P), shell whorl dextral, protoconch area distinctly darker brown; growth lines distinct; rib count (mean = 42, SD = 8.7, n = 19), weakly raised pale white to cream, weakly extend beyond shell edge, rib interstices dark brown; 10 – 12 evenly spread primary ribs; siphonal ridge prominent raised rounded, formed by 3 primary ribs; shell lip uneven, scalloped. Interior shell lip cream with brown splashes aligning with rib interstices, paler than shell margin, ADM scar prominent; spatula cream sometimes bluish; shell lip often golden brown with dark brown markings flanking shallow golden brown siphonal groove; CMS weakly concave, similar but darker colouration to spatula and shell margin; thickening of inner shell lip and spatula occurs in larger specimens, whitening covers brown colouration of inner shell lip. Reproductive system (Figs 72 B, D; n = 2). Positioned within right side of coelom, against foot wall on foot muscle, under the respiratory cavity; epiphallic parts positioned between RAM and BM. GA relatively small, with singular GP through foot wall; AO medium sized, broad, bluntly pointed, central bend, joined to upper GA; ED elongated, broad, centrally twisted, joins to side of GA; GA, AO, ED all white muscular fibrous tissue; EG medium sized, soft whitish tissue, slightly folded, joins at junction of end of ED and extension of single broad long flagellum (F 1), end often looped; BD and CD connect closely together into GA between ED join and GP, both ducts long, slightly bent, smooth, narrow, whitish, featureless, pass closely together through outer side of RAM (BD over CD) into soft white folded tissues of MG; MG / AG complex large; CD connecting to ducts, BD without distal loop, often with loop or fold immediately in front of BC; BC embedded in MG folds, close to embedded SV; BC medium, thin whitish translucent test; HD distinct, short, coiled, links ducts in soft white folded tissues of AG to yellowish granulated HG; outer edge of MG lobbed; AG and HG of similar size, sides match curvature of inner foot wall. Spermatophore (Fig. 72 C). Thread-like, test thin, translucent (length = 6.38 ± 3.71, n = 3), flagella incomplete, head section cylindrical, tip bulbous bluntly rounded, containing a white gelatinous core, tapers into the filamentous transparent flagellum, both sections smooth, featureless, head much thicker than flagellum (length = 5.90 ± 3.02, n = 3, ~ 71 % of SPM length, head width = 119 ± 33 μm, n = 3), two SPM embedded in brown gelatinous mass in one bursa (AM C. 584943). Radula (Figs 83 I – L). Dentition formula 32: 1: 32 (n = 1, AM C. 320123); single central rachidian tooth short broad, flanked squarely by 32 half row laterals, 10 are inner, 6 mid and 16 outer laterals; number of transverse rows not counted; central tooth short wide with short unicuspid mesocone; inner laterals (without endo or ectocones) prominent, mesocones of inner and mid laterals bicuspidate (Figs 83 I – L), mid laterals with broad pointed ectocone; outer, laterals typically with a ‘ chisel’ shaped mesocone, often weakly bicuspidate, flanked by small, pointed single ecto and endocones, angle of separation of each cone from the mesocone varies and maybe inwardly curved (Fig. 83 K).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2D82AEFF68FF02FC00FC36.taxon	diagnosis	Comparative remarks. Siphonaria restis sp. nov. (atra group, unit 54) forms a well-differentiated lineage in clade G of the mitochondrial tree (Figs 1, 2). It differs from other species by COI distances of ≥ 21.8 % (Table S 4). Throughout its range, we found S. restis sp. nov. in sympatry with seven congeners. Two are species are sympatric in south-western WA: For comparisons with S. stowae and S. jeanae refer to comparative remarks under these species. Three are sympatric in northern WA: For comparisons with S. atra, S. viridis, and S. gemina sp. nov. refer to comparative remarks under these species One species is sympatric in western WA: Refer to comparative remarks under S. zelandica. The RS figure of ‘ S. luzonica’ in Hubendick (1955: 3, fig 3) from Rottnest Is, corresponds well with the RS of S. restis sp. nov. shown here (i. e., parallel junction of BD and CD into GA, relative size of AO and twist in BD close to BC, Fig. 72 B), and not that of S. sipho, the senior synonym of S. luzonica.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2D82AEFF68FF02FC00FC36.taxon	distribution	Distribution and habitat. Recorded as endemic to western and northern coasts of WA, Indian Ocean (Fig. 73). In this study, commonly found in sheltered positions on moderately exposed rocky shores, at upper and mid littoral levels.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2D82AEFF68FF02FC00FC36.taxon	etymology	Etymology. From the Latin ‘ restis’ meaning ‘ cord’ — referring to the cord-like primary ribs on the shell of this species.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2F82AFFCCAFB82FE70F876.taxon	description	(Figs 71 I, Q – R, 72 E – F)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2F82AFFCCAFB82FE70F876.taxon	materials_examined	Material examined. Type material. Holotype, from Rivière Banane 19 ° 11.25 ’ S, 63 ° 22.866 ’ E, N coast Rodrigues; coll. A. Meunier and O. Griffiths, RG 01 - 1, 20 Aug 2018 (AM C. 585197 [SK 330], Fig. 71 I). Paratype same data as holotype (AM C. 585196 p [M 427, SK 133, protoconch D 1], Fig. 71 Q). Other, non-type material. Rodrigues: Anse Quiter, SW coast, 19 ° 46.183 ’ S, 63 ° 22.866 ’ E, RG 02 - 1 (AM C. 585888 p).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2F82AFFCCAFB82FE70F876.taxon	description	External morphology. Foot wall, mantle, cephalic folds and pneumostome evenly cream, foot sole darker, paler to foot edge; mantle translucent, narrower than foot wall, edge weakly lobed, thickened with broad white edge band; uneven dark / black pigmentation along foot wall and mantle join, concentrated over centre of cephalic folds. Shell (Figs 71 I, Q; Table S 9). Small sized (max sl mean = 8.8 mm, SD = 1.3 mm, n = 2), ovate; height medium; centrally broad, thickness thin; exterior even, smooth; protoconch direction weakly homostrophic (n = 2, Fig. 71 Q), shell whorl dextral; apex offset to posterior and left of centre, apex offset growth reflected in apical ridge, apical banding fades from tan / brown at shell edge to pale protoconch; anterior and lateral apical sides convex, posterior weakly concave; ribs radiate from apex to shell lip, straight unraised, primary and secondary ribs indistinct, rib count (mean = 39.5, SD = 2.5, n = 2), primary ribs irregularly spaced, whitish, ridges narrow, dual ribs form siphonal ridge, secondary ribs brown / tan; shell edge un-scalloped, even; growth striae indistinct. Interior spatula glossy, white, inner margin to shell edge dark chocolate brown with narrow white rays aligning under primary ribs, extending from shell lip to inner margin; ADM scar indistinct; CMS straight; siphonal groove very weakly indented. Thickening or whitening of inner shell lip not observed. Reproductive system (Fig. 72 E; n = 4). Positioned within coelom under the respiratory cavity, epiphallic parts positioned between RAM and BM close to MG. ED joins at underside of small GA, AO joins underside of ED, short, blunt, narrower than ED, smaller than GA; ED thick, elongated, centrally twisted, broader at EG, narrower than AO; single broad curled stubby flagellum F 1, appears as extension of ED at connection with EG; AO, GA and ED all muscular white tissue; EG broad, relatedly large, soft white tissue; BD and CD connect in parallel to GA at opposite sides of GA, BD without distal loop or MA; BD longer and slightly thinner than CD, both ducts smooth and pass together between RAM and inner foot wall connecting into thick layered folds of MG (BD over CD); BC relatively large, spherical, embedded along with part of BD in AG / MG; SV embedded in AG under BC; HD short narrow, coiled, links large AG to a much smaller yellowish granulated HG, AG and MG folded, soft white tissue, with outer sides curved reflecting the close positioning to curvature of inner foot wall at right posterior quarter of coelom. Spermatophore (Fig. 72 F). Broad head with short flagellum (length = 1.62 ± 0.1 mm, n = 4); test thin, whitish, smooth, featureless; head section broad cylindrical, bulbous, centrally bent, rounded tip; short tapering section merges head to filamentous flagellum; head longer, wider than translucent flagellum (head length = 1.04 ± 0.02 mm, ~ 70 % of SPM length, head width = 99 ± 2 μm, flagellum width = 17 ± 0 μm, n = 4); 5 and 14 SPM tightly packed in BC (AM C. 585197, C. 585196).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2F82AFFCCAFB82FE70F876.taxon	diagnosis	Comparative remarks. In our mitochondrial phylogeny, S. rodriguensis sp. nov. (laciniosa group, unit 68) is the sister species of S. sipho (unit 24). Both species are closely related to S. viridis (Figs 1, 3). Siphonaria rodriguensis sp. nov. differs from S. sipho by COI distances of ≥ 14.9 % and from S. viridis by ≥ 13.9 % (Table S 6). We found S. rodriguensis sp. nov. in sympatry with S. fuliginata on Rodrigues, Indian Ocean. For comparative remarks see under S. fuliginata. The shell of S. basseinensis Melvill, 1893 described from Bombay (Fig. 14 M, not reviewed herein) is similar in having a smooth and even exterior, paired flush ribbing, but differs in having a prominent siphonal ridge.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2F82AFFCCAFB82FE70F876.taxon	distribution	Distribution and habitat. Recorded as endemic to Rodrigues Island, Indian Ocean (Fig. 73). In this study, found on exposed marine rocky shores, at upper and mid littoral levels (Fig. 71 R).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2F82AFFCCAFB82FE70F876.taxon	etymology	Etymology. Named after the type location of Rodrigues Island, Indian Ocean.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2E82A9FCCAFF02FDABFC56.taxon	description	(Figs 71 J – L, T, 72 G – H)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2E82A9FCCAFF02FDABFC56.taxon	materials_examined	Material examined. Type material. Holotype, from Reef in front Sud-Sud Hotel, Itampolo, 24 ° 41.431 ’ S, 43 ° 56.603 ’ E, Madagascar; coll. O. Griffiths, MA 03 - 1, July 2018 (AM C. 584952 [M 264], Fig. 71 J). Paratypes same data as holotype (AM C. 585376 10 + p, C. 584953 p [M 265 protoconch H 10], Fig. 71 K, C. 584954 p [M 266], Fig. 71 L, C. 584832 p [SK 402]). Other, non-type material. Madagascar: Ilot de Lokaro, 24 ° 56.5 ’ S, 47 ° 07.1 ’ E, TM 05 (MNHN IM- 2009 - 13792 p [M 579], IM- 2009 - 13769 p [M 581]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2E82A9FCCAFF02FDABFC56.taxon	description	External morphology. Foot wall, mantle, cephalic folds and pneumostome evenly cream, foot sole darker, paler to foot edge; mantle translucent, wide, thin, mantle edge weakly lobed with thickened whitish band; no dark / black pigmentation markings. Shell (Figs 71 J – L, T; Table S 9). Small sized (max sl mean = 15.6 mm, SD = 1.2 mm, n = 3), circular ovate; medium to tall, apex offset weakly to left and posterior of centre, apical sides strongly convex, posterior side straight to weakly convex; protoconch direction weakly homostrophic (n = 2, Figs 71 K), shell whorl dextral; growth lines distinct; ribs (count mean = 45, SD = 2.5, n = 3), raised, whitish, straight, rib ridges rounded, width strongly increases to shell lip; rib interstices narrow, dark brown; majority of ribs primary, extend from apex to slightly protrude beyond shell edge, secondary ribs fill between primary ribs, 3 primary ribs form siphonal ridge; shell lip uneven, scalloped, aligning with primary rib ends. Interior; white rays aligning under primary ribs extend from shell lip to inner shell margin to golden tan spatula, with dark chocolate brown infill between rays; muscle scar impression distinct, same as shell margin colouring, CMS straight; thickening or whitening of shell edge not observed. Reproductive system (Fig. 72 G; n = 1). Positioned withincoelomundertherespiratorycavity, hermaphroditic glands positioned to posterior against right foot wall and over foot sole, epiphallic parts embedded between RAM and BM close to MG, AG and BC. Join of AO and ED distinct together at top of GA, AO smaller than GA, elongated, blunt, centrally bent; ED thick, elongated, narrower than AO; AO, GA and ED all muscular white tissue; EG broad, soft white tissue, single looped narrow flagellum F 1 extends from ED; BD and CD connect side by side into GA, BD bulbous entry with short distal loop and MA in front of ED; CD enters inner side of GA; BD much longer with similar thickness to CD, both ducts smooth and pass together just inside outer side of RAM connecting into MG (BD over CD), BC white opaque test, relatively small, bulbous, embedded along with part of BD in folds of MG / AG; HD short narrow, coiled, links broad AG to an elongated wide yellowish granulated HG, AG and MG folded, soft white tissue, AG larger than HG, with outer sides curved reflecting the close positioning to curvature of inner foot wall at right posterior quarter of coelom; SV embedded on left side of AG close to BC. Spermatophore (Fig. 72 H). Body cylindrical, thread-like (length = 8.72 mm, n = 1, AL = 10.2 mm), test thin, translucent, smooth, featureless; head broad, containing a white core, laterally banded in front half, tip tapered bluntly rounded, tapers to a thin flagellum (head width = 121 μm, flagellum width = 17 μm, n = 1); single SPM tightly coiled in orange gelatinous mass in BC of the holotype.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2E82A9FCCAFF02FDABFC56.taxon	diagnosis	Comparative remarks. In our mitochondrial phylogeny (Figs 1, 2), S. striata sp. nov. (atra group, unit 74) forms a clade with S. alternata (unit 29) from the Caribbean and Gulf of Mexico and S. kurracheensis (unit 28) from Pakistan. It differs from S. kurracheensis by COI distances of ≥ 8.6 % and from S. alternata by ≥ 5.8 % (Table S 5). Notably, the genetically most similar species, S. alternata, is not known to occur in the Indian Ocean. Siphonaria kurracheensis differs in having a lower, paler shell with more prominent darker and broader interior rays extending to the spatula, a shorter F 1, shorter twisted ED, and BD without a bursal loop. We found S. striata sp. nov. in sympatry with two congeners in Madagascar: For comparisons with S. madagascariensis and S. itampoloensis sp. nov. refer to comparative remarks under these species. The epiphallic parts of RS are very similar to those in S. madagascariensis. However, the hermaphroditic parts differ markedly (i. e., arrangement of HG and AG, BC and BD smaller, SPM shorter in S. striata sp. nov.).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2E82A9FCCAFF02FDABFC56.taxon	distribution	Distribution and habitat. Recorded as endemic to Madagascar, Indian Ocean (Fig. 73). In this study, found on moderately exposed inner lagoon rocky reef, at upper and mid littoral levels.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2E82A9FCCAFF02FDABFC56.taxon	etymology	Etymology. Derived from Latin adjective “ striatus ” meaning ‘ striated / wrinkled / lined / grooved’ referring to the shell’s external striated sculpture.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2882AAFF68FA42FECFFCF6.taxon	description	(Figs 74 A – C, M, 75 A – B)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2882AAFF68FA42FECFFCF6.taxon	materials_examined	Material examined. Type material. Holotype, from nr Tagaqa 18 ° 11.802 ’ S, 177 ° 38.640 ’ E, S. coast of Viti Levu, Fiji; coll. B. W. Jenkins, FI 02 - 1, 22 Aug 2018 (AM C. 584852 [M 275, SK 182], Fig. 74 A). Paratypes: Same data as holotype (AM C. 586005 3 p, C. 584854 p [SK 287], Fig. 74 B, C. 584855 p [SK 297], Fig. 74 C). Other, non-type material. Fiji, Viti Levu: nr Tagaqa, SW coast, 18 ° 11.802 ’ S, 177 ° 38.640 ’ E, FI 02 - 1 (AM C. 584851 p [SK 298]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2882AAFF68FA42FECFFCF6.taxon	description	External morphology. Foot sole grey; foot wall, mantle, cephalic folds, pneumostome evenly cream, paler to foot edge; irregular small blotches of black pigmentation on foot wall and centre of cephalic lobes; mantle slightly narrower than foot wall, covers exposed inner shell lip, wider at anterior, mantle edge strongly lobed with bands of black pigmentation aligned with shell rib interstices; genital pore indistinct, located on foot wall to right anterior of right cephalic fold; small black epithelial eye spot centralised on each of centrally touching cephalic folds; pneumostomal lobe under mantle, unpigmented, between the right ADMs. Shell (Figs 74 A – C; Table S 9). Small to medium sized (max sl mean = 14.8 mm, SD = 1.7 mm, n = 4), ovate; height medium to tall; exterior very uneven, growth disjointed / discontinuous; apex offset central sightly posterior (usually eroded), apical sides convex, straight to concave at posterior; protoconch direction heterostrophic (n = 1), below apex, shell whorl dextral; growth striae prominent in bands, shell thickness thick; rib count (mean = 46, SD = 11, n = 4), marked size difference between primary and secondary ribs; 12 – 15 primary ribs white, prominent, fairly straight, increasingly raised rounded ridge, width broadens to and protrudes weakly beyond shell lip to unevenly scallop and corrugate the edge; paired slightly separated primary ribs form strongly raised siphonal ridge; secondary ribs fill spaces between primary ribs, fairly even in width, rib interstices narrow, darker; some irregular radial coloured banding, dark around apex, paler / cream with dark flecks between primary ribs in lower areas. Interior shell edge and margin white under ribs, brown rays / bands align under rib interstices extend from lip to ADM, siphonal groove distinct, whitish; spatula dark chocolate brown under apex, paler to ADM scar, CMS straight; thickening of shell lip apparent, infills and reduces lip scalloping. Reproductive system (Fig. 75 A; n = 1). Proportionally RS very large structure to animal size, positioned within coelom under the respiratory cavity, epiphallic parts positioned over back of BM, F 1 draped over left side of BM; very small GA, AO broad, bent, rounded, bluntly pointed, rests against MG, joins top of GA; ED long, narrow, slightly bent, joins side of GA; EG medium with folds, overlaps join of ED flagellum (F 1) join, F 1 long, narrow, appears as an extension of ED; AO, GA and ED all muscular white tissue; BD and CD (bulbous at join) connect closely together in opposing directions into GA close to ED and AO entries; BD with distal loop and MA, BD longer and slightly narrower than CD, both ducts smooth, narrow, both pass together through outer side of RAM (BD above CD), connecting into folds of MG; BC embedded in MG, size medium, bulbous test soft translucent, SV embedded within AG; HD lobed, links AG to small brownish granulated HG, MG and AG small folded soft white tissue, anterior edge of MG lobed, AG smaller than HG, sides match curvature of inner foot wall at right posterior quarter of coelom. Spermatophore (Fig. 75 B). Body cylindrical, elongate (length = 7.05 mm, n = 1); test thin, translucent; head bulbous, tip bluntly rounded, wide, evenly cylindrical, containing a core white gelatinous mass, tapers along the transparent flagellum to a thin tip; both sections smooth, featureless. Head section longer, much wider than flagellum (head length = 4.77 mm, n = 1, ~ 68 % of SPM length, head width = 1.33 mm, flagellum width = 13 μm). 2 SPM tightly coiled and embedded in brown gelatinous mass in BC of holotype.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2882AAFF68FA42FECFFCF6.taxon	diagnosis	Comparative remarks. Siphonaria tagaqaensis sp. nov. (laciniosa group, unit 66) represents a well-differentiated lineage in the phylogenetic tree (Figs 1, 3). It differs from other species by COI distances of ≥ 27 % (Table S 6). Siphonaria yagasaensis is anatomically similar concerning the shape of AO, ED, F 1 and SPM, but differs in having a lower, thicker, paler, whitish shell. We found S. tagaqaensis sp. nov. in sympatry with two congeners on Viti Levu: For comparison with S. namukaensis refer to comparative remarks under that species. Siphonaria vudaensis sp. nov. has a smaller, taller shell with stronger scalloped edge, flared siphonal ridge, darker interior, a narrower AO, and a larger BC, and BD without a bursal loop.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2882AAFF68FA42FECFFCF6.taxon	distribution	Distribution and habitat. Recorded as endemic to Viti Levu, Fiji (Fig. 74 M). In this study, found in sheltered positions on moderately exposed rocky shores at mid-littoral level.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2882AAFF68FA42FECFFCF6.taxon	etymology	Etymology. For the type locality of Tagaqa, S coast of Viti Levu, Fiji.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2B82ABFF68FCC2FEFEFC96.taxon	description	(Figs 74 D – F, O – P, 75 C – D)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2B82ABFF68FCC2FEFEFC96.taxon	materials_examined	Material examined. Type material. Holotype: from Okinawa, Tancha Bay, 26 ° 27.897 ’ N, 127 ° 49.131 ’ E; coll. B. W. Jenkins, JP 01 - 5, 20 March 2020 (AM C. 585615 [M 495, SK 314], Fig. 74 D). Paratypes: same data as holotype (AM C. 585949 15 p, C. 584912 p [SK 386], Fig. 74 E); Moon Bay (26 ° 26.653 ’ N, 127 ° 48.230 ’ E), Onna, Okinawa, Japan; coll. B. W. Jenkins, JP 01 - 4, 18 March 2020 (AM C. 584913 p [M 505, SK 326], Fig. 74 F). Other, non-type material. Japan, Okinawa: Cape Maeda, 26 ° 26.573 ’ N, 127 ° 46.113 ’ E, JP 01 - 2 (AM C. 585624 5 p); Moon Bay, Onna 26 ° 26.653 ’ N, 127 ° 48.230 ’ E, JP 01 - 4 (AM C. 595956 5 p); Sun Marina Beach, Onna, 26 ° 27.842 ’ N, 127 ° 48.755 ’ E, JP 01 - 1 (AM C. 585623 5 p, C. 595921 p [SK 537], C. 595924 p [SK 538], C. 595926 p [SK 536]); Tancha Bay 26 ° 27.897 ’ N, 127 ° 49.131 ’ E, JP 01 - 5 (AM C. 584853 p [SK 399]); Tancha Bay 2, rocky point 26 ° 27.941 ’ N, 127 ° 49.194 ’ E, JP 01 - 6 (AM C. 585628 4 p).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2B82ABFF68FCC2FEFEFC96.taxon	description	External morphology (Fig. 74 O). Foot sole dark grey; foot wall, mantle, cephalic folds, pneumostome lobe evenly grey, paler to foot edge; irregular small blotches of black pigmentation on foot wall and centre of cephalic lobes; mantle translucent, wider than foot wall, covers inner shell lip, wider at anterior, mantle edge strongly lobed with bands of black pigmentation aligned with shell rib interstices; genital pore indistinct, located on foot wall to right anterior of right cephalic fold; small black epithelial eye spot centralised on each of centrally touching cephalic fold; pneumostomal lobe under mantle. Shell (Figs 74 D – F; Table S 9): medium sized (max sl mean = 19.8 mm, SD = 2.3 mm, n = 4), circular ovate; tall; apex offset central and sightly left, apical sides convex; protoconch direction homostrophic (n = 2; holotype C. 585615 [M 495], paratype C. 584913 [M 505]), shell whorl dextral; shell exterior and edge uneven; growth striae prominent; pale brown radial banding with irregular darker flecks; shell thickness thick; rib count (mean = 43.3, SD = 0.8, n = 4), 10 – 12 fairly evenly spread primary ribs, pale white, weakly bent and uneven, strongly raised rounded ridges (some> 1 mm), widen to shell edge, protrude beyond shell lip to unevenly scallop between primary ribs and corrugate the edge, ends of primary ribs may be flared creating uneven roughness on rib ridges; between primary ribs brown flecks / bands with 0 – 6 finer whitish secondary ribs (commonly 2), rib interstices narrow darker; siphonal ridge formed by three adjacent ribs, no more raised than other primary ribs. Interior shell margin white to cream, with fine dark brown markings on shell edge under rib interstices, may extend over shell margin, white rays align on shell edge under primary / secondary ribs, ADM scar distinct, brown to white as spatula; cephalic muscle straight to weakly concave; thickening and vivid whitening of shell lip common (Fig. 74 F). Reproductivesystem (Fig. 75 C; n = 2). Proportionally RS very large structure to animal size, positioned within coelom under the respiratory cavity, epiphallic parts compacted, positioned over back of BM to side of RAM, F 1 draped over left side of BM; very small GA, AO broad, bent, rounded, bluntly pointed, rests against MG, joins top of GA; ED long, narrow, bent, joins side of GA; EG medium with folds, overlaps join of ED flagellum (F 1) join, F 1 long, narrow, appears as an extension of ED; AO, GA and ED all muscular white tissue; BD and CD (bulbous at join) connect closely together in opposing directions into GA ED and AO junction; BD with distal loop and MA, BD longer and slightly narrower than CD, both ducts smooth, narrow, pass together through outer side of RAM (BD above CD), connecting into folds of MG; BC embedded in MG, smallish, bulbous test soft translucent, SV embedded within AG; HD lobed, links AG to small brownish granulated HG, MG and AG small folded soft white tissue, anterior edge of MG lobed, sides match curvature of inner foot wall at right posterior quarter of coelom. Spermatophore (Fig. 75 D). Broad head with short flagellum (length = 1.67 mm, n = 1); head section cylindrical, bulbous, centrally bent, tip rounded; test thin, smooth, featureless, translucent encasing a white opaque thin central core; short looped tapering section merges head to filamentous flagellum, head longer, wider than flagellum (head length = 1.41 mm, flagellum length = 0.26 mm, n = 1, 85 % of SPM length, head width = 172 μm, flagellum width = 21 μm); 2 SPMs tightly coiled in brown gelatinous mass in BC of one specimen [SK 538].	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2B82ABFF68FCC2FEFEFC96.taxon	diagnosis	Comparative remarks. Siphonaria tanchaensis sp. nov. (laciniosa group, unit 20) represents a well-differentiated lineage in the mitochondrial tree (Figs 1, 3). It differs from its sister species S. poindimiensis sp. nov. (unit 19) by COI distances of ≥ 21.8 % (Table S 6). We found S. tanchaensis sp. nov. in sympatry with four congeners on Okinawa: For comparative remarks see under S. camura sp. nov., S. rucuana, S. subatra, and S. sipho, respectively. Generally, shell features of this species closely resemble those of other members of the laciniosa group (i. e., few prominent, weakly extending primary ribs, secondary ribs patterning and colouration, internal colouration, irregular weakly scalloped shell edge, thickened white shell margin lip). Figured specimens of ‘ S. laciniosa ’ in Kira (1962: pl. 69, fig. 11) from central Honshu and of ‘ laciniosa group, unit 20 ’ (Dayrat et al. 2014: fig. 4 B) from Okinawa corresponds well with typical features of S. tanchaensis sp. nov.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2B82ABFF68FCC2FEFEFC96.taxon	distribution	Distribution and habitat. Recorded as endemic to Okinawa, Japan (Fig. 73). In this study, found on exposed rocky shores at mid littoral level (Fig. 74 P).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2B82ABFF68FCC2FEFEFC96.taxon	etymology	Etymology. Named after the type locality of Tancha Bay, Okinawa, Japan.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2A82A5FF68FC22FF56FF36.taxon	description	(Figs 74 G – J, N, Q – R, 75 E – H)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2A82A5FF68FC22FF56FF36.taxon	materials_examined	Material examined. Type material. Holotype, from S end Tanguisson Beach, 13 ° 32.549 ’ N, 144 ° 48.443 ’ E, Guam; coll. B. W. Jenkins, GM 02 - 1, 27 Dec 2018 (AM C. 584877 [M 417, SK 102 (RS, SPM)], Fig. 74 G). Paratypes same data as holotype (AM C. 585897 7 p, C. 584878 p [M 438, SK 163], Fig. 74 I; C. 584875 p [M 342], Fig. 74 H; C. 584881 p [SK 253], Fig. 74 J). Other, non-type material. Guam: Tanguisson Beach, S end 13 ° 32.549 ’ N, 144 ° 48.443 ’ E GM 02 - 1 (AM C. 585995 10 p, C. 584876 d [M 349], C. 584879 p [M 341], C. 584880 p [M 350], C. 584882 p [SK 141 protoconch D 10]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2A82A5FF68FC22FF56FF36.taxon	description	External morphology (Fig. 74 R). Foot sole and foot wall evenly pale grey, paler at foot / wall edge; foot wall with patches of black pigmentation, mantle edge with vertical bands of black pigmentation aligned with shell rib interstices; mantle weakly lobed to unlobed, non-translucent, narrower than width of foot wall, covers exposed inner shell lip; genital pore noticeable, located on foot wall to right anterior of right cephalic fold; two small black epithelial eye spots centralised on two centrally touching black pigmented cephalic folds; pneumostomal lobe long, under the mantle, unpigmented, between the right ADMs. Shell (Figs 74 G – J; Table S 9): small sized (max sl mean = 12.9 mm, SD = 0.6 mm, n = 7), circular ovate; height low; apex offset central and to left; apical sides weakly convex, posteriorly weakly concave, protoconch direction homostrophic (n = 1, Fig. 74 Q), shell whorl dextral; growth striae not prominent; radial colour bands, protoconch area dark, paler central band, edge band dark; in some individuals central band is eroded creating concave sides (Fig. 74 H); shell thick, pale grey; rib count (mean = 36.3, SD = 3.9, n = 7), primary ribs white, fairly straight, flattened, increasingly broaden (up to double width) and often raised to shell edge, weakly protrude beyond uneven shell lip to corrugate the edge; 1 – 2 secondary ribs between primary ribs, rib interstices narrow, black to dark brown; paired primary ribs (often fused) form siphonal ridge, end raised at shell edge. Interior shell margin and shell lip off-white under primary / secondary ribs, dark chocolate brown rays align under rib interstices; siphonal groove distinct, white to golden brown; spatula and distinct ADM scar dark brown to mottled tan, CMS convex; thickening of shell lip not noted. Reproductive system (Figs 75 E, G; n = 3). Positioned within coelom under the respiratory cavity, hermaphroditic glands positioned to posterior against right foot wall and over foot sole, epiphallic parts positioned to anterior between BM and RAM. Join of AO and GA indistinct, AO elongated, bluntly pointed, centrally bent with no prominent MA, larger than GA, much thicker than ED; ED relatively long, coiled, twisted, thickened; EG white, folded, elongated; single flagellum F 1 on EG, very long, reasonably straight, unlooped, positioned over BM, thin; AO, GA and ED all muscular white tissue; BD and CD with opposing bulbous connections to GA between ED, AO and GP; BD wrinkled, not twisted, much longer and thinner than short CD with prominent looping on anterior side, both ducts smooth and pass together through RAM connecting into MG (BD above CD), BC translucent, white test, small sized and bulbous; HD short, coiled, links AG to a small elongated yellowish finely granulated HG; MG and AG relatively large, folded, soft white tissue; SV embedded on left side of AG, AG larger than HG, sides match curvature of inner foot wall at right posterior quarter of coelom. Spermatophore (Figs 75 F, H). Relatively long (length 12.7 ± 4.4 mm, AL = 9 mm, n = 2), test thin, translucent, over half-length comprises a translucent cylindrical head section (~ 64 % of SPM length), lip bluntly rounded, containing a white gelatinous thread-like core; tapers into a filamentous transparent flagellum; head section longer and much thicker than flagellum (head length = 7.9 ± 0.4 mm, head width = 78 ± 12 μm, flagellum width = 14 ± 4 μm, n = 2), both sections smooth, featureless; SPM tightly coiled, embedded in red-brown gelatinous mass [SK 102, SK 414].	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2A82A5FF68FC22FF56FF36.taxon	diagnosis	Comparative remarks. In our mitochondrial phylogeny (Figs 1, 3), S. tanguissonensis sp. nov. (plicata group, unit 59) is the sister species of S. lirata (unit 58) also from Guam. Both species differ by COI distances of ≥ 11.6 % (Table S 7). We found S. tanguissonensis sp. nov. in sympatry with three congeners on Guam: For comparative remarks see under S. guamensis, S. lirata, and S. normalis, respectively. The shell sculpture and ribbing in S. tanguissonensis sp. nov. is coarser and more raised than in the former species. The rayed colouration of the internal shell lip in S. tanguissonensis sp. nov. resembles that of S. normalis, S. guamensis (darker), and S. lirata (paler, finer).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2A82A5FF68FC22FF56FF36.taxon	distribution	Distribution and habitat. Recorded only from northern Guam (Fig. 73). In this study, found in sheltered positions on moderately exposed rocky shores, at upper littoral level (Fig. 74 N).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2A82A5FF68FC22FF56FF36.taxon	etymology	Etymology. For the type, of Tanguisson Beach, Guam.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2482A5FF68FE82FBBCFA56.taxon	description	(Figs 74 K – L, S – T, 75 I – J)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2482A5FF68FE82FBBCFA56.taxon	materials_examined	Material examined. Type material. Holotype, from Ethel Beach 10 ° 27.827 ’ S, 105 ° 42.497 ’ E, Christmas Is, Australia, Indian Ocean; coll. B. W. Jenkins, CI 02 - 1, 11 Sept 2018 (AM C. 584664, [M 305, SK 256], Figs 74 K, T). Paratype: E side Smith Point Flying Fish Cove 10 ° 25.749 ’ S, 105 ° 39.957 ’ E, Christmas Is, Indian Ocean; coll. B. W. Jenkins, CI 01 - 2, 9 Sept 2018 (WAM S 74050 [SK 120], Fig. 74 L). Other, non-type material. Christmas Is: E side Smith Point Flying Fish Cove, 10 ° 25.749 ’ S, 105 ° 39.957 ’ E CI 01 - 2 (AM C. 585979 p, C. 585980 p [SK 413]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2482A5FF68FE82FBBCFA56.taxon	description	Externalmorphology (Fig. 74 T). Footedge, footwall, mantle, cephalic folds and pneumostomal lobe all yellow in colour with indistinct darker pigmentation markings; foot sole darker yellow to grey; mantle translucent, wider than foot wall, wider over anterior, strongly lobed with a thickened edge, tips of the major and minor mantle lobes align under primary ribs, wide bands of dark grey pigmentation align under the dark brown colouration of rib interstices; mantle lobes large, mould / extend to cover area of shell lip and align with undulations of primary shell ribs; pneumostome narrow, positioned between right adductor muscles and within mantle. Shell (Figs 74 K, L; Table S 9). Small to medium sized (max sl mean = 17.6 mm, SD = 1.9 mm, n = 2), elongate ovate; height very low; apex offset to posterior and left, apical sides concave, protoconch direction undetermined, shell whorl dextral; growth striae indistinct, shell thin; rib count (mean = 28.5, SD = 1.5, n = 2), exterior evenly dark chocolate brown, pale white where rib ridges eroded, ribs straight, ridges rounded, larger broaden to and protrude beyond shell lip to strongly scallop and weakly corrugate the edge, some ends squared off; smaller secondary ribs fill gaps between primary ribs, rib interstices narrow; bent paired primary ribs on siphonal ridge. Interior evenly dark chocolate brown, some off-white rays under outer ends of primary ribs, siphonal groove shallow distinct; ADM scar distinct, CMS straight; thickening of shell lip not observed. Reproductive system (Fig. 75 I; n = 3). Positioned against inside of foot muscle and foot wall on the right side within coelom, under the respiratory cavity. GA, EG and ED positioned between BM and RAM. AO small, bluntly pointed, joins to larger GA with singular GP; ED relatively long, broad, straight, much larger than AO, joins to upper GA; GA, AO, ED all white muscular fibrous tissue; EG small soft whitish with single wide adjacent flagellum (F 1) appearing as an extension of ED and of similar length; BD and CD jointly but in opposing positions connect to GA between AO and GP; BD with a prominent distal loop and large MA to inner body wall is longer and slightly thinner than CD; both ducts smooth, relatively long and pass closely together through outer side of RAM (BD over CD); CD connects into MG; BC small, bulbous, with thin translucent test, embeds in lower folds of MG / AG close to SV; HD short, bulbous, orange coloured, coiled, links AG to a smaller, yellow / orange, granulated HD; MG and AG large, folded, soft white tissue; AL = 11.78 mm. Spermatophore (Fig. 75 J). Body cylindrical, elongate (length = 11.8 mm, AL = 13 mm, n = 1); test thin, translucent; head tip bluntly rounded, wide, evenly cylindrical, containing a core white gelatinous mass, tapers along the transparent flagellum to a thin tip; both sections smooth, featureless. Head section longer, much wider than flagellum (head length = 6.7 mm, n = 1; ~ 56 % of SPM length, head width = 103 μm, flagellum width = 17 μm); SPM tightly coiled, embedded in whitish gelatinous mass [SK 413].	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2482A5FF68FE82FBBCFA56.taxon	diagnosis	Comparative remarks. In our mitochondrial phylogeny (Figs 1, 2), S. tenebrae sp. nov. (atra group, unit 92) is the sister species of S. subatra (unit 38). Both species differ by COI distances of ≥ 15.1 % (Table S 3). Both species also differ in shell characteristics, RS structure, and external morphology. Siphonaria tenebrae has been found in sympatry with five congeners on CI: Siphonaria umbra sp. nov. has a less scalloped shell edge, more raised and prominent ribs, a paler interior with a darker spatula, and a larger AO and BC. For comparisons with S. alba, S. incerta, S. christmasensis sp. nov., and S. delicata sp. nov. refer to comparative remarks under these species.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2482A5FF68FE82FBBCFA56.taxon	distribution	Distribution and habitat. Known distribution endemic to CI, Australia, Indian Ocean (Fig. 78). In this study, found in sheltered positions on moderately exposed and exposed limestone rocky platforms, at mid littoral levels (Fig. 74 S).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2482A5FF68FE82FBBCFA56.taxon	etymology	Etymology. Derived from Latin noun ‘ tenebrae’ meaning ‘ dark’, referring to the evenly dark interior of the shell for this species.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2482A7FCCAFA62FA02FC96.taxon	description	(Figs 76 A – C, J – K, 77 A – B)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2482A7FCCAFA62FA02FC96.taxon	materials_examined	Material examined. Type material. Holotype, from Poindimie, 21 ° 55.901 ’ S, 165 ° 19.672 ’ E, NC; coll. B. W. Jenkins, NC 03 - 2, 23 Oct 2018 (AM C. 584999 [M 352, SK 128], Fig. 76 A). Paratypes, same data as holotype (AM C. 585837 p [SK 129], Fig. 76 B); Tiari, 21 ° 05.644 ’ S, 165 ° 26.646 ’ E, NC; coll. B. W. Jenkins, NC 04 - 3, 27 Oct 2018 (AM C. 585002 p [SK 261], Fig. 76 C). Other, non-type material. NC: Tiari, 21 ° 05.644 ’ S, 165 ° 26.646 ’ E, NC 04 - 3 (AM C. 585010 p [SK 262]); Poum 2, 20 ° 13.754 ’ S, 164 ° 01.699 ’ E, NC 05 - 3 (AM C. 585998 6 p). Taxonomic remarks. The record of S. laciniosa from NC (Hubendick 1946: 47) possibly refers to this species. External morphology. Foot sole grey; foot wall evenly dark cream, paler to foot edge; irregular black pigmented blotches on foot wall; cephalic folds thick narrow; black pigmentation darker over centre of cephalic folds, paler to foot; mantle wide thin with heavy broad lobes, wider extended at anterior, edge thickened with black pigmentation aligning with rib interstices; pneumostome small, under mantle. Shell (Figs 76 A – C, J; Table S 9). Medium to large sized (max sl mean = 12.9 mm, SD = 1.6 mm, n = 3), elongate ovate; medium to tall; apex strongly offset to left posterior, apex hooked, curved to posterior; apical sides strongly convex, weakly concave to straight on posterior side; shell growth often irregular; protoconch direction central to homostrophic (n = 2, Fig. 76 J) shell whorl dextral; growth striae prominent, shell thick, exterior uneven; rib count (mean = 45, SD = 6.2, n = 3), marked difference between primary and secondary ribs, 9 – 11 primary ribs solidly raised, rounded ridge, weakly crooked, pale white, strongly protrude beyond shell lip to unevenly scallop and corrugate the shell edge; paired ribs form siphonal ridge; areas of finer secondary ribs dark brown / black between primary ribs, often with narrow white flecks / bands. Interior shell lip and lower margin white, changing to dark brown in spatula; dark brown bands extending from shell edge to spatula under secondary ribs and rib interstices; siphonal groove distinct, same colour as shell edge; spatula evenly dark chocolate; ADM scar distinct, CMS straight; thickening of shell lip not apparent, the number of primary ribs may be reduced to 7 in one specimen (Fig. 76 C). Reproductive system (Fig. 77 A; n = 2). Positioned within coelom under the respiratory cavity, hermaphroditic complex (HG, AG and MG) to posterior against right foot wall and over foot sole, epiphallic parts positioned to anterior between RAM and over BM; AO very large, elongated, centrally bent, bluntly bulbous to pointed, merges to upper part of indistinct GA, singular very small GP; ED relatively long, broad, twisted, longer than AO, joins to lower side of GA; GA, AO, ED all white muscular fibrous tissue; EG soft whitish, slightly folded, smaller than AO; single long blunt twisted flagellum (F 1); BD and CD jointly but opposing connections to GA between AO and GP; BD long narrow with prominent distal loops (may loop behind ED) and MAs to inner body wall; CD broad short, wider than BD; both ducts smooth and pass closely together through outer side of RAM (BD over CD); CD connects into MG; BD connects to small bulbous BC with thin translucent test, embedded in folds of MG close to embedded SV; HD short, thick, white, coiled, links AG to smaller yellowish granulated HD; MG and AG folded, soft white tissue. Spermatophore (Fig. 77 B). Test thin, translucent (length 4.11 mm, n = 1), head bulbous, tip bluntly rounded, containing a white gelatinous mass; taper region into the filamentous transparent flagellum is extended; both sections smooth, featureless; head longer and much thicker than flagellum (head length = 3.29 ± 0.49 mm, ~ 72 % of SPM length, head width = 74 ± 8 μm, flagellum width = 1 μm, n = 2); SPM tightly coiled in a brown gelatinous mass. Comparative remarks. In our mitochondrial phylogeny (Figs 1, 3), S. poindimiensis sp. nov. (laciniosa group, unit 19) is the sister species of S. tanchaensis sp. nov. (unit 20). Both species differ by COI distances of ≥ 21.8 % (Table S 6). We found S. poindimiensis sp. nov. in sympatry with seven congeners in NC. For comparisons with S. atra, S. namukaensis sp. nov., S. normalis, S. bourailensis, S. hienghenensis sp. nov., S. caledonica sp. nov., and S. viridis refer to comparative remarks under these species. The external morphology of S. poindimiensis closely resembles that of S. vudaensis sp. nov. (unit 37) from Fiji. The specimen figured as ‘ laciniosa group, unit 19 ’ by Dayrat et al. (2014: fig. 4 A) exhibits morphological characters typical for S. poindimiensis sp. nov.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2482A7FCCAFA62FA02FC96.taxon	distribution	Distribution and habitat. Recorded as endemic to eastern coast of NC, Pacific Ocean (Fig. 78). In this study, found on moderately exposed rocky shores, at upper and mid littoral levels (Fig. 76 K). Etymology. For the type locality, Poindimie, NC.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2682A2FCCAFC22FD66FEB6.taxon	description	(Figs 76 D – E, L – M, 77 C – D)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2682A2FCCAFC22FD66FEB6.taxon	materials_examined	Material examined. Type material. Holotype, from Ethel Beach 10 ° 27.827 ’ S, 105 ° 42.497 ’ E Christmas Is, Indian Ocean; coll. B. W. Jenkins, CI 02 - 1, 11 Sept 2018 (AM C. 584672 [M 306], Fig. 76 D). Paratypes, same data as holotype, WAM S 74051 p [M 409, SK 084], Fig. 76 E). Other, non-type material. Australia. Christmas Is: CI 01 - 1 E side Smith Point Flying Fish Cove (AM C. 585320 p [SK 022], C. 585321 p [SK 070]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2682A2FCCAFC22FD66FEB6.taxon	description	External morphology. Foot wall, foot edge and cephalic folds evenly yellow, foot sole darker yellow to grey; thick narrow; cephalic folds thin. large; mantle translucent, thin, mantle edge thickened lobed, white edge band; pneumostome long, under mantle. Shell (Figs 76 D – E, L; Table S 9). Small to medium sized (max sl mean = 12.14 mm SD = 0.16 mm, n = 3); height low to medium; elongate ovate; apex offset weakly posterior and left, often eroded; apical sides weakly convex anterior, weakly concave posterior, straight to concave lateral; protoconch direction weakly homostrophic (n = 2, Fig. 76 L), shell whorl dextral; growth lines uneven wavy distorted; radial banding prominent; shell lip uneven, anterior slightly protruded; rib count (mean = 38, SD = 1.6, n = 3), primary and secondary ribs white, raised, extend slightly beyond corrugated shell lip, flare upwardly; often only one small or no secondary rib between primary ribs, rib interstices grey to brown; prominent primary rib gaps either side of and 3 – 4 times as wide as siphonal ridge, filled with 3 – 5 very small secondary ribs; siphonal ridge prominent raised rounded with 3 siphonal ribs; interior shell lip and margin dark brown with white rays of varying lengths aligning under primary ribs, extend to white to brown spatula; dark brown / black bands line either side of spatula, siphonal groove and ADM scar prominent; CMS straight to convex, similar but darker colouration to spatula and shell margin, shell margin noticeably wider at anterior; thickening of inner shell lip and spatula occurs in larger specimens, white layering coats and covers brown / black colouration of inner shell lip. Reproductive system (Fig. 77 C; n = 1). Relatively large to animal size; positioned against inside of foot muscle and foot wall on the right side within coelom, under the respiratory cavity. GA, EG and ED positioned between BM and RAM. AO large, elongated, bluntly bulbous, joins to small white muscular fibrous GA with singular GP; ED relatively short, straight, narrow, smaller than AO, joins to side of GA; GA, AO, ED all white muscular fibrous tissue; EG soft whitish, slightly folded, similar size to AO, with two flagellum, F 1 long (similar length to ED), bent, appears as an extension of ED, internally lays on top of the BM; F 2 short, straight, close to EG. BD and CD jointly connect to GA between ED, AO and GP; BD slightly longer and thinner than CD, with a prominent loop over and with MA to mid ED; both ducts smooth and pass closely together through RAM connecting into MG (BD over CD), BC large, spherical, thin translucent test, (2 SPM in one BC), embedded in lower folds of MG / AG close to SV; HD short, narrow, coiled, links smallish AG to a large yellowish granulated HD; MG and AG small, folded, soft white tissue; AG larger than HG, sides match curvature of inner foot wall [M 409]. Spermatophore (Fig. 77 D). Thread-like (length = 10.4 ± 2.1 mm, n = 6), test thin, translucent, comprises a translucent cylindrical body section containing a white gelatinous thread-like core, tapers rapidly into a filamentous transparent flagellum (head length = 10.3 ± 0.61 mm, ~ 78 % of SPM length, head width = 140 ± 30 μm, flagellum width = 20 ± 0 μm, n = 2), head section thicker than flagellum, head tip bluntly rounded, both sections smooth, featureless; 4 SPM coiled, embedded in red-brown gelatinous mass [M 409].	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2682A2FCCAFC22FD66FEB6.taxon	diagnosis	Comparative remarks. In our mitochondrial phylogeny (Figs 1, 2), S. umbra sp. nov. (atra group, unit 46) is the sister species of S. radians (unit 95). Both species are genetically well-differentiated differing by COI distances of ≥ 21.8 % (Table S 4). We found S. umbra sp. nov. in sympatry with five congeners on CI: For comparisons with S. alba, S. incerta, S. tenebrae sp. nov., S. christmasensis sp. nov., and S. delicata sp. nov. refer to comparative remarks under these species. Shell colour patterns resemble that of S. gemina sp. nov.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2682A2FCCAFC22FD66FEB6.taxon	distribution	Distribution and habitat. Recorded as endemic to CI, Australia, Indian Ocean (Fig. 78). In this study, found in sheltered positions on moderately exposed rocky shores, at mid to upper littoral levels (Fig. 76 M).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2682A2FCCAFC22FD66FEB6.taxon	etymology	Etymology. From ‘ umbra’ (Latin = shadow) for the two prominent axial dark areas both externally and internally at either side of the siphonal ridge and spatula groove in the shell of this species; noun in apposition.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2382A3FF68FBE2FF63FE56.taxon	description	(Figs 76 F – I, N – O, 77 E – F)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2382A3FF68FBE2FF63FE56.taxon	materials_examined	Material examined. Type material. Holotype, from Waikoloa Beach, 19 ° 55.456 ’ N, 155 ° 53.491 ’ W, Big Island, Hawaii; coll. B. W. Jenkins, HA 04 - 2, 25 June 2018 (AM C. 584901 [M 294, SK 213], Fig. 76 F). Paratypes same data as holotype (AM C. 585587 4 p, C. 584903 p [SK 257], Fig. 76 G, C. 584904 [SK 258], Fig. 76 H, C. 585913 p [SK 388], Fig. 76 I). Other, non-type material. Hawaii, Big Island: Waikoloa Beach, 19 ° 55.026 ’ N, 155 ° 53.282 ’ W, HA 04 - 1 (AM C. 585375 10 p, 19 ° 55.456 ’ N, 155 ° 53.491 ’ W, HA 04 - 2 (AM C. 584905 p [SK 205]).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2382A3FF68FBE2FF63FE56.taxon	description	External morphology (Fig. 76 P). Foot edge, foot wall, cephalic folds and pneumostomal lobe all evenly yellowish / green, foot sole cream; irregular black pigmentation markings around foot wall, concentrated over central cephalic lobes; mantle translucent narrow thickened edge strongly lobed / folded, mould / extend to cover area of shell lip and align with undulations of primary shell ribs; genital pore indistinct, located on foot wall to right anterior of right cephalic fold; small black epithelial eye spot centralised on each of centrally touching cephalic folds; pneumostome narrow, positioned between right adductor muscles and within mantle. Shell (Fig. 76 I – L, N; Table S 9). Small sized (max sl mean = 13.2 mm SD = 0.8 mm, n = 3); height medium; circular ovate, exterior uneven with discontinuities in growth; apex offset posterior and left; apex slightly hooked, curled to posterior; apical sides weakly convex, weakly concave at posterior; protoconch direction weakly homostrophic (n = 2, Fig. 76 N), shell whorl dextral; growth lines indistinct, rib growth uneven; radial banding prominent; shell lip uneven, anterior slightly protruded; primary and secondary ribs similar, white, width narrow, very crooked, slightly raised, ridges rounded, not extending greatly beyond scalloped / corrugated shell lip, rib count (mean = 45, SD = 3.3, n = 4); siphonal ridge not prominent, formed by dual primary ribs; often only one small or no secondary rib between primary ribs, rib interstices narrow, brown / black; interior shell lip and margin white under primary ribs, dark chocolate brown rays aligning under rib interstices extend from shell lip to evenly dark chocolate brown spatula; siphonal groove prominent, often with calcification; CMS straight, ADM scar distinct, paler than spatula and shell margin, golden colour; thickening of inner shell lip not apparent in larger specimens. Reproductive system (Fig. 77 E; n = 1). Positioned within right side of coelom, against foot wall on foot muscle, under the respiratory cavity; epiphallic parts (GA, EG and ED) positioned between BM and RAM, GA medium, with singular GP through foot wall; AO medium, bluntly rounded, joined at base to upper GA, rests against MG; ED long broad twisted centrally bent, joins to side of GA and base EG; GA, AO, ED all white muscular fibrous tissue; EG small, soft folded whitish tissue; single long narrow flagellum (F 1) centrally bent and looped, appears as an extension of much wider ED; BD and CD connect closely into GA between ED-join and GP, both ducts tightly aligned, broad whitish curved smooth featureless, pass closely together through outside of RAM (BD over CD) into soft white folded tissues of MG / AG complex; CD short and curves to connect with AG duct, BD with distal loop and flat MA to inner foot wall, embeds in folds of AG; BC medium bulbous thin whitish translucent test, 2 SPM in BC (n = 1); HD broad short coiled whitish, links ducts in soft white folded tissues of AG to granulated yellowish small HG; AG much larger than HG. Spermatophore (Fig. 77 F). Thread-like, test thin, translucent, comprises a translucent cylindrical body section containing a white gelatinous thread-like core, tapers rapidly into a filamentous transparent flagellum (head length = 17.33 mm, n = 1), flagellum incomplete, head section much thicker than flagellum (head width = 222 μm, flagellum width = 74.1 μm, n = 1); head tip narrow bluntly rounded; both sections smooth, featureless; 2 SPM tightly coiled, embedded in dark-brown gelatinous mass [M 294].	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2382A3FF68FBE2FF63FE56.taxon	diagnosis	Comparative remarks. Siphonaria undans sp. nov. (plicata group, unit 61) is a well differentiated lineage in the mitochondrial tree (Figs 1, 3). It differs from its sister species S. mauiensis sp. nov. (unit 60) by COI distances of ≥ 29.6 % (Table S 7). We found S. undans sp. nov. in sympatry with S. waikoloaensis on Big Island, Hawaii. Both species are similar, but S. waikoloaensis differs by having a less scalloped shell edge, darker shell lip with paler interior, a slightly larger BC, and shorter SPM.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2382A3FF68FBE2FF63FE56.taxon	distribution	Distribution and habitat. Recorded from Big Island, Hawaii only (Fig. 78). In this study, found in sheltered positions (mainly rock crevices) on moderately exposed fine-algal covered volcanic-rock shores, at upper littoral level.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2382A3FF68FBE2FF63FE56.taxon	etymology	Etymology. Derived from Latin adjective ‘ undans’ meaning ‘ wavy’, referring to the prominent waviness and unevenness of the primary ribs in the shell of this species.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2282A3FF68FE62FC22F8D6.taxon	description	(Figs 79 A – C, K – L, M, 80 A – B)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2282A3FF68FE62FC22F8D6.taxon	materials_examined	Material examined. Type material. Holotype, from Heal of foot, First Landing, near Vuda Point 17 ° 40.753 ’ S, 177 ° 23.006 ’ E, Viti Levu, Fiji; coll. B. W. Jenkins, FI 03 - 1, 25 Aug 2018 (AM C. 584785 [SK 119 (RS and SPM], Fig. 79 A). Paratypes, same data as holotype (AM C. 585523 21 p, C. 584859 p [M 288], C. 584860 p [M 289], Fig. 79 B, C. 584861 p [M 291]; C. 584862 p [M 292], Fig. 79 C). Other, non-type material. Fiji, Viti Vevu: Heal of foot First Landing, 17 ° 40.753 ’ S, 177 ° 23.006 ’ E FI 03 - 1 (AM C. 584863 p [SK 122], C. 584995 p [SK 121]); Vuda Point Marina seawall, 17 ° 40.878 ’ S, 177 ° 23.009 ’ E, FI 03 - 2 (AM C. 585324 p).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2282A3FF68FE62FC22F8D6.taxon	description	External morphology (Fig. 79 K). Foot sole and foot wall evenly cream, paler to foot edge; uneven black pigmented blotches on foot wall; cephalic folds thick narrow; black pigmentation darker over centre of cephalic folds, paler to foot; mantle wide thin with heavy broad lobes, edge thickened with black pigmentation aligning with rib interstices; pneumostome small, under mantle. Shell (Figs 79 A – C, J, L; Table S 9). Medium to large sized (max sl mean = 22.5 mm, SD = 3.3 mm, n = 5), ovate; low, profile very flat; apex offset slightly posterior and left, apical sides convex; shell whorl dextral, protoconch direction homostrophic (n = 1, Fig. 79 L); growth striae apparent, shell thin; rib count (mean = 44.6, SD = 10.4, n = 5), fairly even number of primary and secondary ribs, 14 – 15 primary ribs pale white, fairly straight often bent, slightly raised, protrude 1 – 2 mm beyond shell lip to unevenly scallop a flat edge; siphonal ridge formed by 3 close primary ribs shows greatest extension beyond lip; 1 – 2 secondary ribs between primary, rib interstices white at apex to dark brown black at shell edge. Interior shell margin dark brown with pearly lustre, fine white rays extend over shell margin to spatula aligning under primary / secondary ribs, siphonal groove distinct, slightly curved to lateral; spatula dark chocolate brown to mottled white; ADM scar distinct, darker than margin, same colour as spatula; CMS concave, paler than shell lip; thickening of shell lip occurs, translucent pearly thickening rather than over covering pigmentation or whitening; juvenile specimens thin often translucent; small specimens evenly dark chocolate brown. Reproductive system (Fig. 80 A; n = 1). Positioned within coelom under the respiratory cavity, hermaphroditic complex (HG, AG and MG) positioned to posterior against right foot wall and over foot sole, epiphallic parts positioned to anterior firmly between BM and RAM. AO very large, elongated, centrally bent, bluntly bulbous, joins to upper part of prominent GA, singular GP; ED relatively short, broad, shorter than AO, joins to side of GA; GA, AO, ED all white muscular fibrous tissue; EG soft whitish, slightly folded, smaller than AO; single short blunt twisted flagellum (F 1); BD and CD jointly but opposing connections to GA between ED, AO and GP; BD long narrow, with distinct folds prior to GA connection and prominent distal loop with MA to inner body wall; CD broad short, wider than BD; both ducts smooth and pass closely together through RAM (BD over CD); CD connects into MG; BD loops prior to connection to small bulbous BC with thin translucent test, embedded in folds of MG / AG close to embedded SV; HD short, thick, coiled, links AG to smaller yellowish granulated HD; MG and AG folded, soft white tissue. Spermatophore (Fig. 80 B). Thread-like, relatively long (length = 11.4 mm) and thin, translucent, over half-length comprises a translucent cylindrical body section (head length = 7.2 mm, ~ 63 % of SPM length) containing a white gelatinous thread-like core, tapers rapidly into a filamentous transparent flagellum; head section much thicker than flagellum (head width = 133 μm, flagellum width = 13.1 μm); head tip tapered bluntly rounded, both sections smooth, featureless; 2 SPM coiled, embedded in red-brown gelatinous mass [SK 119].	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2282A3FF68FE62FC22F8D6.taxon	diagnosis	Comparative remarks. In our mitochondrial phylogeny (Figs 1, 2), S. vudaensis sp. nov. (atra group, unit 37) is the sister species of a clade containing two species, S. tenebrae sp. nov. (unit 92) and S. subatra (unit 38). Siphonaria vudaensis differs from these two species by COI distances of ≥ 10.9 % (S. tenebrae) and ≥ 13.3 % (S. subatra) (Table S 3). We found this species in sympatry with three congeners on Viti Levu: For comparisons with S. namukaensis sp. nov., S. normalis, and S. tagaqaensis sp. nov. refer to comparative remarks under these species. Specimens from Tuituila, Samoa and Viti Levu figured as ‘ unit 37, atra group’ in Dayrat et al. (2014: figs 5 P, Q) belong to this species. The specimen figured as ‘ S. atra ’ in Cernohorsky (1972: 210, pl. 60, fig. 1) is a specimen of S. vudaensis sp. nov.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2282A3FF68FE62FC22F8D6.taxon	distribution	Distribution and habitat. Recorded from Viti Levu, Fiji and Tutuila, American Samoa (Fig. 78). In this study, found in sheltered positions on moderately exposed rocky boulder shores, at mid littoral level.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2282A3FF68FE62FC22F8D6.taxon	etymology	Etymology. For the type locality, Vuda Point, Viti Levu, Fiji.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2282DCFCCAF8E2FB86FC56.taxon	description	(Figs 79 D – F, N – O, 80 C – D)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2282DCFCCAF8E2FB86FC56.taxon	materials_examined	Material examined. Type material. Holotype, from Waikoloa Beach, 19 ° 55.456 ’ N, 155 ° 53.491 ’ W, Big Island, Hawaii; coll. B. W. Jenkins, HA 04 - 2, 25 June 2018 (AM C. 584907 [M 295, SK 214], Fig. 79 D). Paratypes: same data as holotype (AM C. 585327 10 + p; Waikoloa Beach 19 ° 55.026 ’ N, 155 ° 53.282 ’ W. HA 04 - 1, 25 June 2018 (AM C. 585327 p [SK 254], Fig. 79 E; C. 584673 p [SK 391], Fig. 79 F). Other, non-type material. Hawaii. Big Island: Waikoloa Beach, 19 ° 55.456 ’ N, 155 ° 53.491 ’ W, HA 04 - 2 (AM C. 584906 p [SK 206]); 19 ° 55.026 ’ N, 155 ° 53.282 ’ W, HA 04 - 1 (AM C. 595958 2 d).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2282DCFCCAF8E2FB86FC56.taxon	description	External morphology. Foot edge, foot wall, cephalic folds and pneumostomal lobe all evenly yellowish / green, foot sole cream; irregular small blotches of black pigmentation on foot wall and centre of cephalic lobes; mantle translucent narrow thickened edge strongly lobed with bands of black pigmentation aligned with shell rib interstices; genital pore indistinct, located on foot wall to right anterior of right cephalic fold; small black epithelial eye spot centralised on each of centrally touching cephalic folds; pneumostomal lobe under mantle, unpigmented, between the right ADMs. Shell (Figs 79 D – F, N; Table S 9). Small sized (max sl mean = 10.6 mm, SD = 1.4 mm, n = 3), ovate; height tall, shell thin; apex offset weakly posterior and left, apex often hooked / curled to posterior, apical sides strongly convex, protoconch direction homostrophic to central (n = 2, Fig. 79 O), shell whorl dextral; 3 radial colour bands, protoconch brownish, central pale brown and outer showing black interstices; rib count (mean = 45, SD = 7.8, n = 3), ~ 12 – 14 prominent primary ribs, white, crooked, raised, rounded ridges, width broads to shell lip, protrude beyond shell lip with a raised roll to unevenly scallop and corrugate the edge; interstices between primary ribs with 3 – 4 smaller secondary ribs; paired primary ribs form siphonal ridge, no more prominent than other primary ribs. Interior shell evenly dark chocolate brown, narrow irregular white rays extend from shell lip to margin aligning under primary / secondary ribs, siphonal groove distinct; ADM scar distinct, CMS straight; thickening of shell lip not observed. Reproductive system (Fig. 80 C; n = 1). Positioned within right side of coelom, against foot wall on foot muscle, under the respiratory cavity; RS proportionally large to animal size compared to other species; epiphallic parts (GA, EG and ED) positioned in between BM and RAM, GA medium, with singular GP through foot wall; AO medium, bluntly rounded, joined at base to upper GA, rests against MG; ED long broad twisted centrally bent, joins to side of GA and base EG; GA, AO, ED all white muscular fibrous tissue; EG large, soft folded whitish tissue; single long broad flagellum (F 1) centrally bent, appears as an extension of much wider ED; BD and CD connect closely in opposite directions into GA between ED join and GP, both ducts narrow whitish curved smooth featureless, pass closely together through outside of RAM (BD over CD) into soft white folded tissues of MG / AG complex; CD curves and broadens to connect with AG duct, BC long with distal loop and flat MA to inner foot wall, embeds in folds of AG; BC small bulbous thin whitish translucent test, 2 SPM in BC (n = 1); HD very small short coiled whitish, links ducts in soft white folded tissues of AG to granulated small HG; AG much larger than HG. Spermatophore (Fig. 80 D). Relatively short, test thin, translucent, comprises a translucent cylindrical body section containing a white gelatinous thread-like core, tapers into a filamentous transparent flagellum (head length = 8.111 mm n = 1, flagellum incomplete), head section much thicker than flagellum (head width = 185 μm, flagellum width = 74.1 μm n = 1), head tip bluntly rounded; both sections smooth, featureless; 2 SPM tightly coiled, embedded in dark-brown gelatinous mass in holotype.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2282DCFCCAF8E2FB86FC56.taxon	diagnosis	Comparative remarks. Siphonaria waikoloaensis sp. nov. (plicata group, unit 55) forms a well-differentiated lineage in the mitochondrial tree (Figs 1, 3). It differs from other species by COI distances of ≥ 28 % (Table S 7). We found S. waikoloaensis sp. nov. in sympatry with S. undans on Big Island, Hawaii. For a comparison see under this species.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2282DCFCCAF8E2FB86FC56.taxon	distribution	Distribution and habitat. Recorded from Big Island, Hawaii, USA, Pacific Ocean (Fig. 78). In this study, found in sheltered positions (mainly rock crevices) on moderately exposed fine-algal covered volcanic-rock shores, at upper littoral level (Fig. 79 N).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF2282DCFCCAF8E2FB86FC56.taxon	etymology	Etymology. For the type locality, Waikoloa Beach, Big Island, Hawaii.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF5D82DEFCCAFC62FC84FAB6.taxon	description	(Figs 79 H – J, 80 E – F)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF5D82DEFCCAFC62FC84FAB6.taxon	materials_examined	Material examined. Type material. Holotype, from Oneata Is, 18 ° 26.5166 ’ S, 178 ° 29.583 ’ E, Fiji. SWP 17 - 116 (AM C. 584827 [M 437, SK 226], Fig. 79 H). Paratypes, same data as holotype (AM C. 584710 5 p); Yagasa, small island to the SW, 18 ° 57.783 ’ S, 178 ° 29.333 ’ E, Tuvana-i-Ra Island, Fiji. SWP 17 - 103 (AM C. 584828 p [M 436], Fig. 79 J); Tuvana-i-Ra Island 21 ° 02.24 ’ S, 178 ° 45.01 ’ E, Fiji. SWP 17 - 77 (AM C. 608181 p [M 435], Fig. 79 I). Other, non-type material. Fiji: Tuvana-i-Ra Island, SWP 17 - 77, 21 ° 02.24 ’ S, 178 ° 45.01 ’ E (AM C. 585937 3 p); Yagasa, small island, SW of Tuvana-i-Ra Island, SWP 17 - 103, 18 ° 57.783 ’ S, 178 ° 29.333 ’ E (AM C. 585936 2 p).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF5D82DEFCCAFC62FC84FAB6.taxon	description	External morphology (preserved). Foot sole and foot edge without dark pigmentation; foot wall, cephalic folds and pneumostomal lobe evenly cream with irregular blotches of black pigmentation, concentrated over centre of cephalic lobes; mantle translucent, wider than foot wall, edge weakly lobed, without pigmentation. Shell (Fig. 79 H – J; Table S 9). Medium sized (max sl mean = 16.7 mm, SD = 4.6 mm, n = 3), elongate ovate; low to medium; apex offset central slightly posterior and left, mostly externally coated with algae similar to Lithothamnion; apical sides weakly concave, protoconch direction undetermined, shell whorl dextral, shell thick; rib count (mean = 33, SD = 4.9, n = 3), continuum of forms from primary to secondary ribs; primary ribs pale white, straight, slightly protrude beyond shell lip to unevenly scallop and corrugate the edge; 1 – 2 interspersed pale white finer secondary ribs, rib interstices narrow; paired primary ribs form siphonal ridge, no more prominent than other primary ribs. Interior shell lip / margin white of irregular widths and lengths, aligning under primary / secondary ribs; spatula dark chocolate brown to ADM, siphonal groove deep and distinct, paler than shell margin and spatula; ADM scar distinct, CMS straight, same colour as spatula; thickening of shell lip common, often in smaller specimens (Figs 79 H, I), infills and reduces lip scalloping, becomes pale brown to dark cream. Reproductive system (Fig. 80 E; n = 1). Positioned within coelom under the respiratory cavity, hermaphroditic complex (HG, AG and MG) to posterior against right foot wall and over foot sole, epiphallic parts between BM and RAM, F 1 lays over posterior of BM; AO large, elongated, centrally bent, blunt, merges to upper part of indistinct GA, singular GP; ED short, wide, twisted, short than AO, joins to lower side of GA and AO; GA, AO, ED all white muscular fibrous tissue; EG soft whitish, folded, smaller than AO; single short broad flagellum (F 1), possible F 2; BD and CD closely but in opposing directions connect to side of GA between AO and GP; both ducts smooth and pass through outside of RAM (BD over broader CD); BD long narrow with prominent distal loop and MA attached to inner body wall in front of BM, joins to BC with thin transparent test; CD shorter wider than BD; CD connects into MG; BC and CD embed in folds of AG and MG; HD broad, lobed, brown markings, under AG, links AG to much smaller yellowish granulated HG; MG and AG folded, soft white tissue. Spermatophore (Fig. 80 F). Thread-like (length = 3.04 mm, n = 1), translucent, test thin; head section bluntly rounded, evenly cylindrical, containing a white gelatinous core, tapers along the transparent flagellum to a thin tip; both sections smooth, featureless; head section longer wider than flagellum (head length = 2.48 mm, flagellum length = 0.55 mm, head width = 56 μm, flagellum width = 11 μm, n = 1); SPM tightly coiled in bursa, embedded in brown gelatinous mass.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF5D82DEFCCAFC62FC84FAB6.taxon	diagnosis	Comparative remarks. In our mitochondrial phylogeny (Figs 1, 3), S. yagasaensis sp. nov. (laciniosa group, unit 67) is the sister species of S. namukaensis sp. nov. (unit 22). Siphonaria yagasaensis sp. nov. differs from other species by COI distances of ≥ 13.9 % (Table S 6). For a comparison with S. namukaensis refer to comparative remarks under this species. Siphonaria yagasaensis sp. nov. exhibits a similar shell morphology with other species in the plicata group.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF5D82DEFCCAFC62FC84FAB6.taxon	distribution	Distribution and habitat. Recorded exclusively to Oneata, small island SW of Yagasa and Tuvana-i-Ra Islands, southern Fiji (Fig. 78). In this study, found on exposed and moderately exposed rocky shores, at upper littoral level.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF5D82DEFCCAFC62FC84FAB6.taxon	etymology	Etymology. For the type locality, Yagasa Island, Fiji.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF5F82DEFF68FA02FB42FA76.taxon	type_taxon	Type species Ancylus gussoni Costa, 1829, by monotypy.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF5F82DEFF68FA02FB42FA76.taxon	discussion	Taxonomic remarks. Our morphological and molecular studies confirm that Siphonaria and Williamia are closely related. Indeed, their sister group relationship has recently been confirmed by a phylogenomic study of the Panpulmonata (Krug et al. 2022). Morphologically, the patelliform shell of Williamia is distinct from that of Siphonaria. Williamia is distinguished by the combination of the following characteristics: Fragile, externally no ribs or pale axial bands, interior ADM complete circular. By contrast, Siphonaria exhibiting a much greater specific diversity, is quite variable in many shell characteristics, such as geometry, colour, position of apex and ribbing, variable thickness, strongly to unraised external ribs. Most significantly, Siphonaria differs from Williamia by having an internal, horseshoe-shaped ADM. Several Siphonaria exhibit shells that somewhat resemble that of Williamia (e. g., S. oblia, S. lateralis, S. thersites, S. compressa Allanson, 1958 [not examined]). However, these species clearly differ in anatomical characters. The external morphology is similar in both Siphonaria and Williamia. Both groups have a head anterior to posterior shell apex, dual cephalic lobes with central eye spots, mantle, right lateral position of siphon and rectum, pneumostomal lobe (although it is longer in Williamia), and similar sizes of ducts and spermatophore (e. g., S. obliquata, S. sipho to S. radiata.). The radula dentition differs between Siphonaria and Williamia. In Williamia, the central tooth is small and unicuspid, and demarcation of inner and outer lateral teeth is more prominent, inner laterals strongly bicuspid and outer laterals single plates. Hubendick (1946: 8, 18) recognized Williamia as a distinct genus within Siphonariidae distinguished from Siphonaria by a combination of anatomical and shell characters. However, not all differences are confirmed herein for all species. For example, the BC is in front of the adductor muscle [ADM] in Williamia but also in S. camura sp. nov. The shape of the adductor muscle impression in Siphonaria differs from Williamia having an unbroken circular scar, but this is also seen in S. radiata. The BD in Siphonaria runs through or outside this the RAM is correct for most species, but not for S. camura sp. nov. However, it never runs through the RAM in Williamia.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF5F82DAFCCAFA42FE3CFCB6.taxon	description	(Figs 81 A – H, J, 80 G)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF5F82DAFCCAFA42FE3CFCB6.taxon	materials_examined	Material examined. Type material. Lectotype of Tectura radiata Pease, 1860 from ‘ Sandwich Islands’ [Hawaii], Cuming Collection, date? (NHMUK 1962837; Fig. 81 A). One paralectotype, same data as lectotype (NHMUK 1962838). Holotype of Capulus nutatus Hedley, 1908 from Balmoral Beach [Sydney, NSW, Australia]; coll. C. T. Starkey, pre. 1907 (AM C. 029106; Fig. 81 B). Twelve paratypes, same data as holotype (AM C. 143080; Fig. 81 C largest paratype). Holotype of Roya kermadecensis Iredale, 1912 from Sunday Island, Kermadec Group (CM M. 5464; Fig. 81 D). Holotype of Williamia polynesica Rehder, 1980 from off Waikiki, Oahu, Hawaii (USNM 757897; Fig. 81 E). Other, non-type material. NZ: Kermadec Ids, Raoul (Sunday) Is 29 ° 15 ’ S, 177 ° 52 ’ W (AM C. 30230 8 d); 4 kms N of Cape Rodney, 36 ° 15 ’ S, 174 ° 49 ’ E (AM C. 110823 4 d); Off Poor Knights Islands 35 ° 28.3 ’ S, 174 ° 44.13 ’ E (NMNZ M. 100391 / 1 p [M 601, SK 555]). NC: Loyalty Is, Lifou Is (AM C. 25728 2 d; AM C. 335728 d). PNG: Ella Beach, Port Moresby, 9 ° 29 ’ S, 147 ° 9 ’ E (AM C. 335729). Australia: Coral Sea, Northeast Herald Cay, 16 ° 56 ’ S, 149 ° 11 ’ E (AM C. 335727). Elizabeth Reef, 29 ° 57.2 ’ S, 159 ° 1.18 ’ E (AM C. 336005 d), 29 ° 54.78 ’ S, 159 ° 2.78 ’ E (AM C. 336013 d), 29 ° 54.78 ’ S, 159 ° 2.78 ’ E (AM C. 336015 d); Middleton Reef, 29 ° 23.93 ’ S, 159 ° 6.7 ’ E (AM C. 461846 d), 29 ° 24.02 ’ S, 159 ° 6.2 ’ E (AM C. 461845 p), 29 ° 24 ’ S, 159 ° 5.16 ’ E (AM C. 461844 d), 29 ° 24.03 ’ S, 159 ° 4.37 ’ E (AM C. 461843 d). NI: 29 ° 2 ’ S, 167 ° 57 ’ E (AM C. 59405 p [SK 006]). Qld: Michaelmas Cay, 16 ° 36 ’ S, 145 ° 59 ’ E (AM C. 110822 2 d); SW side Euston Reef, 16 ° 40 ’ S, 146 ° 13 ’ E (AM C. 335730 9 d); Bunker Group, Lady Musgrave Is, 23 ° 54 ’ S, 152 ° 25 ’ E (AM C. 398297 d). Caloundra, 26 ° 49 ’ S, 153 ° 10 ’ E (AM C. 335994 d). LHI: 31 ° 46.55 ’ S, 159 ° 13.4 ’ E (AM C. 461858 d), 31 ° 32.5 ’ S, 159 ° 3.75 ’ E (AM C. 59697 d). NSW: Angourie Point, 29 ° 29 ’ S, 153 ° 22 ’ E (AM C. 112574 2 d); Clarence River (mouth), 29 ° 25.5 ’ S, 153 ° 21 ’ E (AM C. 110824 5 d); Woolgoolga, 30 ° 6.7 ’ S, 153 ° 12.3 ’ E (AM C. 110818 2 d); Point Halliday, 32 ° 4.5 ’ S, 152 ° 33 ’ E (AM C. 117308 2 d, AM C. 336021 4 d); North Fingal Bay, 32 ° 44.75 ’ S, 152 ° 10.5 ’ E (AM C. 110814 11 d, AM C. 117310 d); Port Stephens, 32 ° 42 ’ S, 152 ° 5 ’ E (AM C. 110812 5 d, C. 595937 p [SK 562]); Sydney, Balmoral Beach, Middle Harbour, 33 ° 49.7 ’ S, 151 ° 15.02 ’ E (AM C. 143080 12 d, AM C. 29106 d); Coogee Bay, 33 ° 55.5 ’ S, 151 ° 15.4 ’ E (AM C. 110813 2 d); Kurnell, Botany Bay, 33 ° 57 ’ S, 151 ° 12.37 ’ E (AM C. 107325 8 d, AM C. 110817 d, AM C. 117309 d, AM C. 112575 19 d); Bate Bay, Cronulla Beach, 34 ° 2.5 ’ S, 151 ° 10 ’ E (AM C. 110815 2 d). Shell Harbour, 34 ° 35 ’ S, 150 ° 52 ’ E (AM C. 398294 d); Gerringong, 34 ° 44.28 ’ S, 150 ° 50 ’ E (AM C. 110816 d); Ulladulla, 35 ° 21.5 ’ S, 150 ° 28.5 ’ E (AM C. 398293 d); Twofold Bay, 37 ° 5.11 ’ S, 149 ° 55.35 ’ E (AM C. 50019 d). Tas: Flinders Island, Fotheringate Beach, 40 ° 12.948 ’ S, 148 ° 2.106 ’ E (TMAG E 36516 d, TMAG E 36516 p). WA: Turtle Beach, W side of North West Cape, 21 ° 48 ’ S, 114 ° 10 ’ E (AM C. 110825 d); North West Cape, 21 ° 49 ’ S, 114 ° 11 ’ E (AM C. 110821 2 d); Point Quobba, N of Carnarvon, 24 ° 29 ’ S, 113 ° 25 ’ E (AM C. 398298 d); Off Chinaman’s Bluff, Kalbarri, 27 ° 42 ’ S, 114 ° 9 ’ E (AM C. 336007 d); South of Cowaramup, 33 ° 53 ’ S, 114 ° 59 ’ E (AM C. 110820 d).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF5F82DAFCCAFA42FE3CFCB6.taxon	discussion	Taxonomic remarks. The lectotype of Tectura radiata has been designated by Kay (1965). The name W. radiata Kuroda & Habe, 1961 (in Habe 1961) is a junior secondary homonym of W. radiata (Pease, 1861). Habe (1962: 96, pl. 44, fig. 14) introduced W. japonica as a replacement name (Art. 12.2 of the Code). Subsequently, Habe (1964: 144) treated Williamia japonica as a junior synonym of W. radiata, which was confirmed by Marshall (1981: 488). Marshall (1981) recognised two geographic subspecies; W. radiata radiata (Pease, 1861) and W. radiata nutata (Hedley, 1908), but suspected that both taxa would ‘ ultimately prove to be synonyms ’.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF5F82DAFCCAFA42FE3CFCB6.taxon	description	External morphology (preserved). Animal evenly dark yellowish, black pigmentation absent; foot wall and mantle narrow, mantle edge thickened unlobed, singular cephalic fold, wide, front wide, striated, 2 black eye spots separately located within cephalic fold (Fig. 80 G). Shell (Figs 81 A – H, J; Table S 9). Circular ovate, small sized (max sl mean = 6.7 mm, SD = 0.8 mm, n = 8), tall, shell thickness thin; exterior smooth, evenly yellowish brown, growth striae indistinct; apex offset weakly posterior and central, lateral and anterior apical sides strongly convex, posterior side concave, protoconch below shell apex, direction weakly heterostrophic (n = 1, Fig. 81 J), shell whorl dextral; ribs pale axial bands, fairly straight, increasingly broaden to shell lip, periostracum extends beyond uncorrugated shell edge. ribs unraised, rib count (mean = 24, SD = 1.9, n = 8), Interior paler than exterior. ADM circular and complete (Figs 81 D, F), thinner over siphonal opening and CMS. Reproductive system (Fig. 80 G; n = 1). Positioned within coelom under the respiratory cavity, epiphallic parts (AO, EG, ED, F 1) positioned over and behind BM, hermaphroditic complex (HG, AG and MG) to posterior against right foot wall under intestine and over foot sole; singular GP positioned immediately behind right side of cephalic fold; ED long wide, joins to lower side of small GA, AO indistinct; EG large, soft, folded, flagellum (F 1) indistinct; BD and CD jointly connect to side of GA, CD passes between foot wall and outside RAM, CD short wide flat, passes on inner side of RAM, both BD and CD connect into folds of AG and MG; BC medium spherical embedded under MG; HD long, broad, unfolded, links AG to smaller HG; MG and AG folded, soft white tissue.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF5F82DAFCCAFA42FE3CFCB6.taxon	diagnosis	Comparative remarks. Throughout the known range of W. radiata no sympatric congener has been recorded. The shells figured in Kay (1979: figs 157 K – L), Marshall (1981: fig. 2 B, D, F) correspond well with shells figured herein (Figs 81 B – H) as well as the lectotype figured in Kay (1965: pl. 11, figs 6, 7; Fig. 81 A herein). The circular and complete ADM scar (Fig. 81 F) corresponds well with the scar figured in Iredale (1915: pl. 9, fig. 10) as ‘ Roya kermadecensis ’ from Sunday Is, Kermadecs. The figured horseshoe shaped ADM scar of Williamia in Hubendick (1946: fig. 9) is incorrect. The RS of ‘ W. radiata’ described and figured in Ruthensteiner et al. (2007: figs 1 A – C) corresponds well with that figured herein (Fig. 80 G). The record of Roya sp. in Powell (1934: 155) likely refers to W. radiata.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF5F82DAFCCAFA42FE3CFCB6.taxon	distribution	Distribution and habitat. Recorded from Kermadec Islands, NZ, NC, PNG, Indian Ocean, E and SW Australia (Fig. 83) in subtidal depths.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF5B82DAFCCAFA02FB17F7F6.taxon	discussion	Taxonomic remarks. The original description and original figure as the only indication to the identity of S. alternicosta provide insufficient information to identify this species. No type specimen (s) are known to exist (Philippe Bouchet; pers. comm.). Menke (1844: 54) considered S. alternicosta (incorrectly attributed to Quoy & Gaimard) as an accepted species occurring on the W coast of Australia. The original figure resembles S. restis sp. nov. from WA somewhat, but this similarity is not considered taxonomically significant. We consider S. alternicosta as a nomen dubium.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF5B82DAFF68FC42FB3AFEB6.taxon	discussion	Taxonomic remarks. The species description was attributed by Reeve to Nuttall, but Reeve is the author of the name. Initially, the name was first mentioned by Jay (1839: 39) as a nomen nudum without a description (Art. 12.1 of the Code) for a specimen from the ‘ Sandwich Isles’ (Hawaii). The name has not been made available subsequently by Catlow & Reeve (1845: 100) or Jay (1850: 104) (no description, Art. 12.1). Carpenter (1864 a: 646) correctly recognised Reeve as the author of the name and stated that the species was from Hawaii. Similarly, White & Dayrat (2012: 61) indicated the distribution to be ‘ very likely Hawaii’. Suter’s (1913: 599) reference to ‘ Hutt. ’ as author of the name is likely in error for ‘ Nutt [all] ’. Suter (1909 b: 258, 1913: 599) reported S. amara from ‘ New Guinea’, which is considered incorrect and stated that it ‘ is also a near relation of S. cookiana’ (= S. propria Jenkins, 1983) from NZ, which is also not accepted herein. Hubendick’s (1946: 66 – 67) included S. amara in the list of nomina nuda due to uncertainty of type locality (California or Hawaii?), but it is not a nomen nudum. Examination of the holotype of S. amara (NHMUK 1981016 revealed some similarly with S. waikoloaensis sp. nov. from Hawaii. However, the external surface is more uneven, and the ribbing appears significantly broader, projecting at edge. The siphonal ridge is less prominent in S. amara.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF5B82DAFCCAFE02FCDDFAF6.taxon	discussion	Taxonomic remarks. The original description, original figures and type locality provide the only indication to the identity of S. coreensis. No types of S. coreensis could be located at the UKNHM (J. Ablett, pers. comm.). The identity of the species cannot be clarified based on the original description and figures alone and remains indeterminable. The identity of this species may be resolved in the future when suitable topographic material should become available. The original figures are not of S. sirius (Figs 36 E, F), which differs in shell morphology (i. e., multiple ribs on siphonal ridge) nor of S. japonica (Figs 7 A – E), which differs in shell geometry. Subsequently, Reeve (1856) listed S. coreensis as a synonym of S. atra. However, Reeve’s description and figure are based on a misidentification and herein attributed to S. sirius (Figs 36 E 0 F). Lischke (1871: 105) and Cernohorsky (1972: 210) followed Reeve’s treatment as a synonym of S. atra.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF5582D5FF68FF02FDDAFC56.taxon	description	(Fig. 84 A)	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF5582D5FF68FF02FDDAFC56.taxon	discussion	Taxonomic remarks. Utter confusion exists regarding the existence of a possible type specimen, the type locality, and the identity of S. laciniosa. Linnaeus (1758: 781) provided a brief description (‘ ribs raised unequal, the darker outwardly blunted’) and a type locality (‘ India’) referring to figures in pre-Linnean works, such as Rumphius (1705: 121, pl. 40, fig. C, 1711: 8, 1741: 121; pl. 40, fig. C) and D’Argenville (1757: 184, pl. 2, fig. O). The shells depicted in both works are of the ‘ atra group’. Linnaeus (1767: 1258) and Gmelin (1791: 3695) expanded the original description and stated that P. laciniosa ‘ lives in Indian seas’ (translation), re-citing the engravings of Rumphius and D’Argenville but adding references to Knorr (1789: 11, pl. 30, figs 2 – 4, 7 – 8), showing patellids but not siphonariids, and Martini & Chemnitz (1769 [in 1769 – 1829]: pl. 10, fig. 81), showing a modified copy of Rumphius’ (1705: 121) engravings. We conclude that Linneaus was uncertain of the identity of P. laciniosa and how it related to shells described by pre-Linnean authors as these represented different species. The alleged type specimen of Patella laciniosa Linneaus, 1758: 781 UUZM # 912 (specimen label without locality and provenance; Fig. 84 A) has been accessioned by the UUZM with the following information: “ Protologue: 1758: 781. Donation: Gustav IV Adolf. Preparation: dry. Swartz label: laciniosa ”. It is a shell of the ‘ laciniosa group’ and clearly a distinct from the species figured by Rumphius’ as ‘ two-eyed spotted species’, which is of the ‘ atra group’ (Fig. 6 N). Dillwyn (1817: 1021) and Hanley (1855: 417) opined that the name P. laciniosa was essentially based on the figure of Rumphius (1705 – 1741). This would mean that the UMMZ specimen cannot be a type. Hanley (1855) was uncertain of the identity of the shell figured in Rumphius (1741). According to Sherborn (1940), Benthem Jutting (1959) and Dance (1967), most of Rumphius’ specimens have been lost. We could not trace any information on a specimen of ‘ Patella tertia ’ or ‘ two-eyed spotted’ figured by Rumphius (1741). It is possible, perhaps even likely, that Rumphius’ depicted a shell from Indonesia. Dillwyn (1817: 1021) stated that the species ‘ inhabits the coasts of Amboyna’ [Ambon, Indonesia]. This might be the type locality of P. laciniosa if Linneaus’s statement on the origin of this species was a reference to Rumphius. Morrison (1972: 58) suggested that the ‘ Eye spots’ are likely due to erosion of the shell apex, a feature confirmed herein to be commonly observed in species of the ‘ atra group’. Hubendick’s (1946) was the first to attribute type status to the UUMZ specimen mentioned above. Hubendick (1946: 48) stated that ‘ the type of S. laciniosa is in the keeping of the Zool. Mus., Uppsala’, but confusingly identified it as a specimen of S. laciniosa forma exigua referring to a name introduced by Sowerby I, 1823. Subsequently, Dance (1967: 80) stated that no type specimen of P. laciniosa was found in the Linnaean collection and considered it to be lost. However, according to Odhner (1954: 26) and Wallin (2001: 89) there is a type of P. laciniosa in the UUZM catalogue of Linnaean type specimens. The identity of this alleged type of P. laciniosa cannot be ascertained. It is likely a siphonariid of the ‘ laciniosa group’, but its features are insufficient to identify this species. Moreover, the generic type locality ‘ India’ may refer to any place in the Tropical parts of the Indian Ocean, including the Indonesian archipelago and is too vague to allow to narrow the selection down to a single species. Morrison (1972) considered Hubendick’s claim of a type specimen as an invalid type designation stating that ‘ Linne didn’t indicate that he had ever seen a specimen up to 1767 ’. Alternatively, Morrison (1972: 58) designated the non-binomial ‘ the two-eyed spotted species’ figure in Rumphius (1705: 125, pl. 40, fig. C; Fig. 84 B) as ‘ the type of laciniosa ’. This type designation has subsequently been acceptedbyChristiaens (1980 a: 78). However, Hubendick’s and Morrison’s type concepts are irreconcilable with each other contributing to the confusion around this name. If we accept Hubendick’s claim that the UMMZ specimen is the type, then S. laciniosa is an unidentifiable species in the ‘ laciniosa group’ and distinct from the species depicted by Rumphius. However, if we accept Morrison’s view, then S. laciniosa is an unidentifiable species of the ‘ atra group’ possibly from Indonesia. We are of the view that the ambiguity surrounding this name cannot be resolved without a ruling of the commission and therefore we treat the name P. laciniosa herein as a nomen dubium. This name has been used throughout the body of taxonomic literature, but because of the ambiguity surrounding its identity, subsequent uses invariably were hypothetical and often contradictory. The earliest taxonomists effectively reiterated the name referring to the original description of Linnaeus (1758: 380) without contributing any clarification of the original concept (e. g., Schrӧter 1784: 403; Röding 1798: 5; Roissy 1805: 214; Dillwyn 1817: 1021; Lamarck 1819: 325; Sowerby I 1825: 29; Hanley 1855: 417). Pilsbry (1892 [in 1891 – 1892]: 160) listed P. laciniosa as ‘ unidentified Patellidae ... possibly = P. stellaeformis ’. Eventually, Hubendick (1946: 47) transferred the species to Siphonaria based on examination of the probable type. However, his delimitation of the species is speculative and not accepted herein. Specimens identified as ‘ S. laciniosa’ by Hubendick (1946: 47) are not from the type locality and many examples comprise mixed lots.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF5482D5FCCAFA42FBB8F836.taxon	discussion	Introduced by Jay (1839: 39) as a nomen nudum and attributed to Nuttall without description this is an unavailable name (Art. 12.1 of the Code; see also White & Dayrat, 2012: 62). It was listed as a synonym and therefore not made available subsequently by Catlow & Reeve (1845: 100), Jay (1850: 104), Reeve (1856: pl. 2, species 9), Paetel (1873: 117, 1875: 92, 1883: 178, 1889: 428), Pilsbry (1920 b: 379), Schrenck (1867: 306), Hutton (1880: 36), and Hubendick (1946: 47). Galindo (1977: 416) misspelled the name as ‘ crebicostata ’. Siphonaria crebidentata Smith in Galindo (1977: 416) is a separate emendation referring to this nomen nudum. Because S. crebricosta is not an available name, the lectotype designation by Baker (1964: 159) is also invalid (Art. 74.2 and 5 of the Code).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF5482D5FF68FC62FE2AF9B6.taxon	discussion	Taxonomic remarks. The original description and original figure of Michelin (1832) do not provide enough detail to resolve the identity of this species. No type specimen (s) are known to exist (Philippe Bouchet; pers. comm.). While the original figure in Michelin (1832: pl. 17) matches the features of the atra group, particularly that of S. atra and S. alba, this species remains indeterminable. Hubendick (1946: 69) incorrectly rejected S. sowerbyi as a nomen nudum (followed by White & Dayrat, 2012: 68). Morrison (1972: 56 – 58) treated S. sowerbyi Michelin, 1832, S. sowerbyi Guerin, 1832 and S. sowerbyi Michelin, 1868 as synonyms of S. laciniosa based on similarity in shell form and ‘ common reproductive development’. These synonymies are not accepted here. This nominal species remains a nomen dubium.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF5482D5FF68F902FA2BFDB6.taxon	discussion	Taxonomic remarks. The original description and original figure provide the only indication for the identity of S. stellata (Helbling, 1779). No type specimen (s) are known to exist. Martens (1868: 235) transferred Patella stellata Helbling, 1779 to Siphonaria, indicating that S. exigua Sowerby was the ‘ best’ name reversing the priority. This transfer to Siphonaria rendered S. stellata Blainville, 1827 as a junior secondary homonym (Art. 57.3; White & Dayrat, 2010: 68). Examination of Helbling’s figure (1779) reveals a shell that resembles specimens from Hawaii (e. g., S. waikoloaensis sp. nov., Figs 72 D – F; S. mauiensis sp. nov., Figs 56 G – I) or Mauritius (e. g., S. incerta; Figs 35 H – L) and is unlike the shell of S. exigua (Fig. 3 J). The identity of S. stellata is currently unknown and there is no prospect of clarifying its identity based on the original description alone. The use of the name S. stellata (Helbling) for material from the Bay of Bengal in Nagabhushanam & Krishnan (1993: 480) is therefore questionable. We consider this taxon as a nomen dubium.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF5482D5FCCAFD42FACDFA16.taxon	discussion	This species was described by Blainville (1827: 297) and attributed to ‘ de Roissy’ from an unknown origin. It is a secondary junior homonym of Patella stellata Helbling, 1779 and therefore invalid (Art. 57.3 of the Code). The name has subsequently been referred to by Hanley (1858 b: 153) and Paetel (1889: 429). Hubendick (1946: 48) and Morrison (1972: 56) treated it as a junior synonym. The listing of S. stellata in Adcock (1893: 11) as a synonym of S. luzonica (from SA, Australia; misidentified S. zelandica) is incorrect and a reversal of priority. Figures of ‘ S. stellata ’ RS in Hubendick (1945: figs 42, 53) are here attributed to S. javanica Figs 6 H, I. Shells of ‘ S. stellata’ figured in Hubendick (1946: 91, pl. 3, fig. 20 – 23) are of various accepted species, such as S. viridis (fig. 20 from Thursday Island, fig. 22 from WA) and S. javanica (fig. 21 from Mindanao, fig. 23 from Java). A shell figured in Satyamurti (1952: 252, pl. 34, fig. 2 a, b) resembles Helbling’s figure of P. stellata. Morrison (1972: 56 – 58) treated S. stellata Blainville 1827 along with 29 other nominal species as a synonym of S. laciniosa. The identity of S. stellata listed in Ravenish & Biju Kumar (2015: 38) is unknown (unfigured, no reference specimens).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF5782D6FF68F842FBF5FEF6.taxon	discussion	Legosiphon mulinus was introduced by Iredale (1940: 437) as a nomen nudum without description (Art. 13.1 of the Code). The features mentioned refer to Legosiphon. We have not found any subsequent usage of this name. It remains an unavailable name.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF5782D6FCCAFEE2FB0DFDF6.taxon	discussion	Mestosiphon parmelas was introduced by Iredale (1940: 437) as a nomen nudum without description (Art. 13.1 of the Code). It was not made available subsequently by Cernohorsky (1972: 210; synonym of S. atra), Morrison (1972: 57; no description) nor Short & Potter (1987: 122). It remains an unavailable name.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF5782D6FF68F942FDEBF816.taxon	discussion	Parellsiphon commixtus was introduced by Iredale (1940: 437 as a nomen nudum without description (Art. 13.1 of the Code). The name was not made available subsequently by Morrison (1972: 57; no description) nor White & Dayrat (2012: 61; not intentionally declared as new, Art. 86.2 of the Code). It remains an unavailable name.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF5782D6FF68F982FE67F916.taxon	discussion	Introduced by Davis (1904: 127) as a variety of S. alternata (Say, 1827), the name is an objective junior homonym of S. sipho intermedia Schrenck, 1867 and permanently invalid (Art. 57.2 of the Code).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF5782D6FF68FE02FE67FDB6.taxon	discussion	Described from Otago (Tavaï-Pounamou, Nouvelle-Zélande) [New Zealand] without figure by Hombron & Jacquinot (1841: 192), this name is a primary junior homonym of S. costata Sowerby, 1835 and permanently invalid (Art. 52.2 of the Code).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF5782D6FCCAFDE2FCECFCD6.taxon	discussion	Siphonaria crebridentata was introduced by Galindo (1977: 437) and attributed to Smith as a nomen nudum without description (Art. 13.1 of the Code). The name was not used again subsequently and remains an unavailable name.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF5782D6FF68FBC2FEC0FAF6.taxon	discussion	Introduced by Paetel (1873: 117) as a nomen nudum without description and incorrectly attributed to Crosse this is an unavailable name (Art. 12.1 of the Code). It has not been made available subsequently by Paetel (1883: 178, 1889: 428).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF5782D6FF68FCC2FD32FB96.taxon	discussion	Introduced by Paetel (1873: 117) as a nomen nudum without description and attributed to Dunker this is an unavailable name (Art. 12.1 of the Code). Not made available subsequently by Paetel (1883: 178, 1889: 428). It remains an unavailable name. Hubendick (1946: 68) suggested it may be a misspelling of S. laciniosa.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF5782D6FF68FAE2FDD6F9D6.taxon	discussion	Introduced by Paetel (1889: 429) as a nomen nudum without description and incorrectly attributed to ‘ Quoy’ (= Quoy and Gaimard, 1833), this is likely an incorrect subsequent spelling of S. punctata. However, Paetel (1889: 429) separately listed S. punctata.	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF5782D6FF68FD22FE4DFC96.taxon	discussion	Introduced by H. Adams & A. Adams (1855 [in 1853 – 1858]: 271) as a replacement name for S. radiata Sowerby I, 1835, this name is permanently invalid as it is an objective junior homonym of S. sowerbyi Michelin, 1832 (Art. 52.2 of the Code).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
0D49832FFF5782D6FF68FF02FEF6FE56.taxon	discussion	Introduced by Jay (1839: 39) as a nomen nudum and attributed to Petit without description and unfigured this is an unavailable name (Art. 12.1 of the Code). It was not made available subsequently by Catlow & Reeve (1845: 100), Paetel (1889: 429), Hubendick (1946: 69) nor White & Dayrat (2012: 62).	en	Jenkins, Bruce, Köhler, Frank (2024): Hidden in plain sight: Systematic review of Indo-West Pacific Siphonariidae uncovers extensive cryptic diversity based on comparative morphology and mitochondrial phylogenetics (Mollusca, Gastropoda). Megataxa 13 (1): 1-217, DOI: 10.11646/megataxa.13.1.1
