identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
0C3587E856568106FF3CF8DEFA9FFE97.text	0C3587E856568106FF3CF8DEFA9FFE97.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cisaris Townes 1969	<div><p>Cisaris Townes, 1970</p><p>Type species: Cisaris tenuipes Townes, 1970</p><p>Diagnosis: Head and mesosoma heavily punctate. Head comparatively large. Eye with sparse, moderately long hairs. Clypeus wide, in profile weakly convex with the margin reflexed, median section often slightly produced. Mandible with upper tooth much longer than lower tooth. Mesoscutum weakly convex, posterior edge without transverse groove. Notaulus not reaching to center of mesoscutum. Fore wing without an areolet, vein 2m-cu subvertical, with one bulla. Tergites polished. First tergite slender, spiracle far behind middle. Ovipositor compressed, its tip long and gradually tapered. Dorsal valve with a weak nodus. Ventral valve without ridges.</p></div>	https://treatment.plazi.org/id/0C3587E856568106FF3CF8DEFA9FFE97	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Pham, Nhi Thi;Cao, Nga Thi Quynh	Pham, Nhi Thi, Cao, Nga Thi Quynh (2025): First record of the genus Cisaris Townes (Ichneumonidae: Phygadeuontinae) from Vietnam, with descriptions of four new species. Zootaxa 5683 (4): 573-583, DOI: 10.11646/zootaxa.5683.4.7, URL: https://doi.org/10.11646/zootaxa.5683.4.7
0C3587E856578106FF3CFEF1FB95F87C.text	0C3587E856578106FF3CFEF1FB95F87C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cisaris carinatus Pham & Cao 2025	<div><p>Cisaris carinatus sp. nov.</p><p>(Figures 1, 2)</p><p>Material examined. Holotype: ♀ (IB), Son La Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.59145&amp;materialsCitation.latitude=21.328085" title="Search Plazi for locations around (long 103.59145/lat 21.328085)">Copia Nature Reserve</a>, 21°19’41.1”N 103°35’29.22”E, 1,519 m a.s.l., 27.iv–2.v.2016, Malaise trap, Hoang V. T. &amp; Nguyen V.B leg . Paratypes: 2♀ (IB), similar data as holotype, except 21°19’02.2”N 103°35’14.1”E, 1,591 m a.s.l.</p><p>Diagnosis. Antenna without white flagellomeres; scutellum smooth, with sparse punctures, lateral carina long and weak; propodeum with dense, large punctures, area superomedia fused with area basalis; ovipositor sheath 1.2–1.3× as long as hind tibia.</p><p>Description of holotype. Face 0.5× as high as wide, densely coarsely punctate (Fig. 1B). Clypeal suture distinct. Clypeus 2.5× as wide as high, basally sub-polished, with large punctures, subapical margin impressed (Fig. 1B). Mandible long, upper tooth sharp, 2.0× as long as lower tooth. Malar space matt, as wide as basal width of mandible (Fig. 1B). Gena wider than compound eye in lateral view, with dense punctures (Fig. 1D). Frons and vertex rugose punctate. Postero-ocellar distance 0.9× ocular-ocellar distance, 1.4× median ocellus diameter (Fig. 1C). Antenna with 17 flagellomeres, thickened toward apex, F1 2.0× as long as wide, 1.0× as long as F2. Occipital carina complete, ventrally curved before meeting hypostomal carina above base of mandible.</p><p>Mesosoma. Pronotum rugose punctate. Epomia present (Fig. 2A). Mesoscutum 0.8× as long as its maximum width, rugose punctate. Notaulus distinct anteriorly (Fig. 2B). Scutellum moderately convex, sub-polished, with sparse punctures, lateral carina weak and extending nearly to apex (Fig. 2C). Mesopleuron extremely rough, with dense, irregular wrinkles and coarse punctures; epicnemial carina with upper end reaching to subalar prominence. Sternaulus strong anteriorly, posteriorly weaker, reaching hind margin of mesopleuron, distinctly dorsad of lower posterior corner of mesopleuron. Metapleuron reticulate rugose (Fig. 2A). Propodeum with dense, large punctures, carinae distinct. Area superomedia fused with area basalis. Apico-median portion from posterior margin of area superomedial to apex of posterior corner of area petiolaris strongly concave longitudinally (Fig. 2D). Propodeal apophysis strong and compressed. Propodeal spiracle small, round.</p><p>Wings. Fore wing length ca. 2.8 mm, vein 1cu-a postfurcal to M&amp;RS, inclivous, postero-distal corner of subbasal cell ca. 60°, vein 3rs-m absent, vein 2rs-m shorter than length of M between 2rs-m and 2m-cu, length of CU between 1m-cu&amp;M and 2cu-a 2.4× length of 2cu-a. Hind wing with distal abscissa of CU present, length of CU between M and cu-a 4.0× length of cu-a.</p><p>Legs. Hind coxa largely punctate, except concave area dorsally smooth. Hind femur densely puncto-striate, 3.7× as long as its maximum width, 0.9× as long as hind tibia length. Hind tibia coarsely sculptured, 1.1× as long as hind tarsus. Hind basitarsus 0.4× as long as tarsus.</p><p>Metasoma.Tergites smooth and shining, with scattered, small punctures. T1 2.6× as long as apical width, median dorsal carina weak, present medially, dorsolateral carina complete, ventrolateral carina present, weak anteriorly; spiracle small, round. T2 1.1× as long as basal width, 0.6× as long as apical width (Fig. 2E). Laterotergite of T2 1.7× as long as apical width. Ovipositor straight. Ovipositor sheath 1.2× as long as hind tibia.</p><p>Colour. Black with reddish brown. Head black, scape, pedicel, first eight flagellomeres and mandible reddish, apical flagellomeres dark brown, palpi yellow. Metasomal tergites from base to apex reddish brown to reddish. Ovipositor reddish. Ovipositor sheath brown. Legs reddish, except hind femur reddish brown. Wings hyaline, wing veins dark brown.</p><p>Variation. Paratypes have fore wing length 2.8–3.0 mm, T1 2.6–2.7× as long as apical width, T2 0.6–0.7× apical width, ovipositor sheath 1.2–1.3× hind tibia length .</p><p>Male. Unknown.</p><p>Comparative notes. This species differs from other Cisaris species by having lateral carina of scutellum weak but extending nearly to apex.</p><p>Etymology. The specific name refers to the long lateral carina of scutellum.</p><p>Distribution. Currently known only from Son La Province, northwestern Vietnam.</p></div>	https://treatment.plazi.org/id/0C3587E856578106FF3CFEF1FB95F87C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Pham, Nhi Thi;Cao, Nga Thi Quynh	Pham, Nhi Thi, Cao, Nga Thi Quynh (2025): First record of the genus Cisaris Townes (Ichneumonidae: Phygadeuontinae) from Vietnam, with descriptions of four new species. Zootaxa 5683 (4): 573-583, DOI: 10.11646/zootaxa.5683.4.7, URL: https://doi.org/10.11646/zootaxa.5683.4.7
0C3587E856558102FF3CFF02FE20FEEE.text	0C3587E856558102FF3CFF02FE20FEEE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cisaris robustus Pham & Cao 2025	<div><p>Cisaris robustus Pham &amp; Cao, sp. nov.</p><p>(Figures 3, 4)</p><p>Material examined. Holotype: ♀ (IB), Lam Dong Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.69608&amp;materialsCitation.latitude=12.17032" title="Search Plazi for locations around (long 108.69608/lat 12.17032)">Bidoup-Nui Ba National Park</a>, 12°10’13.15”N 108°41’45.9”E, 1,477 m a.s.l., viii–x.2017, Malaise trap, Pham T.N. leg .; Paratypes: 1♀ (IB), similar data as holotype; 1♀ (IB), Son La Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.58725&amp;materialsCitation.latitude=21.317278" title="Search Plazi for locations around (long 103.58725/lat 21.317278)">Copia Nature Reserve</a>, 21°19’02.2”N 103°35’14.1”E, 1,591 m a.s.l., Malaise trap, Hoang V. T. &amp; Nguyen V.B leg .; 1♀ (IB), Lang Son Province, Trang Dinh district, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.413055&amp;materialsCitation.latitude=22.359446" title="Search Plazi for locations around (long 106.413055/lat 22.359446)">Chi Minh</a> commune, 22°21’34”N 106°24’47”E, 350 m a.s.l., 29.iv–5.v.2025, Malaise trap, Pham T.N., Cao T.Q.N., Cao, T.K. T., Dang T.H and Nguyen D.H leg.</p><p>Diagnosis. Antenna without white flagellomeres; scutellum sub-polished, with dense punctures, lateral carina present basally; propodeum with dense, large punctures, area superomedia fused with area basalis; ovipositor sheath 1.0–1.2× as long as hind tibia.</p><p>Description of holotype. Face 0.5× as high as wide, densely coarsely punctate (Fig. 3B). Clypeal suture distinct. Clypeus 2.7× as wide as high, with dense, large punctures, subbasal part weakly raised, subapical margin impressed (Fig. 3B). Mandible long, upper tooth sharp, 3.0× as long as lower tooth. Malar space matt, as wide as basal width of mandible (Fig. 3B). Gena wider than compound eye in lateral view, matt, with dense, coarse punctures (Fig. 3D). Frons and vertex rugose punctate. Postero-ocellar distance equal to ocular-ocellar distance, 1.3× median ocellus diameter (Fig. 3C). Antenna with 17 flagellomeres, thickened toward apex, F1 2.0× as long as wide, 1.0× as long as F2. Occipital carina complete, ventrally curved before meeting hypostomal carina above base of mandible.</p><p>Mesosoma. Pronotum rugose punctate. Epomia present (Fig. 4A). Mesoscutum 0.7× as long as its maximum width, rugose punctate. Notaulus distinct anteriorly (Fig. 4B). Scutellum moderately convex, sub-polished, with dense punctures, lateral carina present basally (Fig. 4C). Mesopleuron extremely rough, with dense, irregular wrinkles and coarse punctures; epicnemial carina with upper end reaching to subalar prominence. Sternaulus strong anteriorly, posteriorly weaker, reaching hind margin of mesopleuron, distinctly dorsad of lower posterior corner of mesopleuron. Metapleuron rugose punctate (Fig. 4A). Propodeum with dense, large punctures, carinae strong. Area superomedia fused with area basalis. Apico-median portion from posterior margin of area superomedial to apex of posterior corner of area petiolaris strongly concave longitudinally (Fig. 4D). Propodeal apophysis strong and compressed. Propodeal spiracle small, round.</p><p>Wings. Fore wing length ca. 3.8 mm, vein 1cu-a postfurcal to M&amp;RS, inclivous, postero-distal corner of subbasal cell ca. 70°, vein 3rs-m absent, vein 2rs-m shorter than length of M between 2rs-m and 2m-cu, length of CU between 1m-cu&amp;M and 2cu-a 2.2× length of 2cu-a. Hind wing with distal abscissa of CU present, length of CU between M and cu-a 3.0× length of cu-a.</p><p>Legs. Hind coxa densely punctate, except concave area dorsally smooth. Hind femur densely puncto-striate, 3.9× as long as its maximum width, 0.85× as long as hind tibia length. Hind tibia coarsely sculptured, 1.1× as long as hind tarsus. Hind basitarsus 0.4× as long as tarsus.</p><p>Metasoma.Tergites smooth and shining, with scattered, small punctures. T1 2.3× as long as apical width, median dorsal carina weak, present medially, dorsolateral carina complete, ventrolateral carina present, weak anteriorly; spiracle small, round. T2 0.9× as long as basal width, 0.6× as long as apical width. Laterotergite of T2 1.6× as long as apical width (Fig. 4E). Ovipositor straight. Ovipositor sheath 1.1× as long as hind tibia.</p><p>Colour. Black with reddish brown. Head black, scape, pedicel, first eight flagellomeres and mandible reddish, apical flagellomeres reddish brown, palpi yellow. Pronotum with narrow reddish anteriorly. Metasomal tergites from base to apex reddish brown to reddish. Ovipositor reddish. Ovipositor sheath reddish brown. Fore and mid legs reddish, hind leg reddish brown. Wings hyaline, wing veins black.</p><p>Variation. Paratypes have fore wing length 3.3–3.6 mm, malar space 0.7–0.8× as long as basal mandible width, T1 2.2–2.7× as long as apical width, laterotergite of T2 1.4–1.7×, ovipositor sheath 1.0–1.2× hind tibia length.</p><p>Male. Unknown.</p><p>Comparative notes. This species differs from Cisaris carinatus sp. nov. by its scutellum densely punctate with lateral carina present basally. In addition, this new species is larger (fore wing length 3.3–3.8 mm vs. 2.8–3.0 mm). Both new species differ from C. tenuipes by their area superomedia fused with area basalis.</p><p>Etymology. The specific name refers to its robust body.</p><p>Distribution. Currently known from Lam Dong Province, the Central Highlands, Son La and Lang Son provinces, northern Vietnam.</p></div>	https://treatment.plazi.org/id/0C3587E856558102FF3CFF02FE20FEEE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Pham, Nhi Thi;Cao, Nga Thi Quynh	Pham, Nhi Thi, Cao, Nga Thi Quynh (2025): First record of the genus Cisaris Townes (Ichneumonidae: Phygadeuontinae) from Vietnam, with descriptions of four new species. Zootaxa 5683 (4): 573-583, DOI: 10.11646/zootaxa.5683.4.7, URL: https://doi.org/10.11646/zootaxa.5683.4.7
0C3587E856538100FF3CFE23FBE6FF5F.text	0C3587E856538100FF3CFE23FBE6FF5F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cisaris rugosus Pham & Cao 2025	<div><p>Cisaris rugosus Pham &amp; Cao, sp. nov.</p><p>(Figures 5, 6)</p><p>Material examined. Holotype: ♀ (IEBR), Lao Cai Province, Hoang Lien National Park, 27.ix.2013, hand net, Pham T.N. leg.</p><p>Diagnosis. Antenna without white flagellomeres; scutellum sub-polished, with moderately dense punctures, lateral carina present basally; propodeum punctate to rugose, area superomedia separated from area basalis, area dentipara densely wrinkled; metasomal tergites 2 and 3 enlarged, almost fused together dorsally, following tergites mostly retracted under tergite 3; ovipositor sheath 1.3× as long as hind tibia.</p><p>Description of holotype. Face 0.5× as high as wide, rugose punctate (Fig. 5B). Clypeal suture distinct. Clypeus 2.9× as wide as high, with large punctures, subapical margin impressed (Fig. 5B). Mandible long, upper tooth sharp, 2.0× as long as lower tooth. Malar space matt, densely punctate, as wide as basal width of mandible (Fig. 5B). Gena wider than compound eye in lateral view, sub-polished, with moderately dense punctures (Fig. 5D). Frons and vertex rugose punctate. Postero-ocellar distance 0.7× as long as ocular-ocellar distance, 1.7× median ocellus diameter (Fig. 5C). Antenna with 17 flagellomeres, thickened toward apex, F1 2.0× as long as wide, 1.0× as long as F2. Occipital carina complete, ventrally curved before meeting hypostomal carina above base of mandible.</p><p>Mesosoma. Pronotum rugose punctate, except narrow impunctate area sub-dorsally. Epomia present (Fig. 6A). Mesoscutum 0.85× as long as its maximum width, sub-polished, with dense punctures laterally and rugose punctate medio-posteriorly. Notaulus distinct anteriorly. Scutellum moderately convex, sub-polished, with sparse punctures, lateral carina present basally (Fig. 6B). Mesopleuron extremely rough, with dense, irregular wrinkles and rugose punctures; epicnemial carina with upper end reaching to subalar prominence. Sternaulus strong anteriorly, posteriorly weaker, reaching hind margin of mesopleuron, distinctly dorsad of lower posterior corner of mesopleuron. Metapleuron reticulate rugose (Fig. 6A). Propodeum with carinae strong. Area superomedia sub-polished with irregular wrinkles laterally. Area basalis with moderately dense punctures. Area dentipara densely wrinkled. Apico-median portion from posterior margin of area superomedial to apex of posterior corner of area petiolaris strongly concave longitudinally (Fig. 6D). Propodeal apophysis strong and compressed. Propodeal spiracle small, round.</p><p>Wings. Fore wing length ca. 4.7 mm, vein 1cu-a postfurcal to M&amp;RS, inclivous, ventrally curved, vein 3rs-m absent, vein 2rs-m shorter than length of M between 2rs-m and 2m-cu, length of CU between 1m-cu&amp;M and 2cu-a 3.0× length of 2cu-a. Hind wing with distal abscissa of CU present, length of CU between M and cu-a 1.7× length of cu-a.</p><p>Legs. Hind coxa densely coarsely punctate, except concave area dorsally smooth. Hind femur sub-polished with dense punctures, 4.0× as long as its maximum width, 0.8× as long as hind tibia length. Hind tibia coarsely sculptured, slightly longer than hind tarsus. Hind basitarsus 0.45× as long as tarsus.</p><p>Metasoma.Tergites smooth and shining, with scattered, small punctures. T1 3.0× as long as apical width, median dorsal carina weak, present medially, dorsolateral carina complete, ventrolateral carina present, weak anteriorly; spiracle small, round. T2 and T3 enlarged, almost fused together dorsally (Figs 6C, E). Laterotergite of T2 2.3× as long as apical width (Fig. 6C). T4 onwards mostly retracted under tergite 3 (Fig. 6C). Ovipositor straight. Ovipositor sheath 1.3× as long as hind tibia.</p><p>Colour. Black with reddish brown. Head black, antenna reddish to reddish brown, mandible reddish, palpi yellow. Metasomal tergite 1 black, tergites 2 and 3 reddish brown. Ovipositor reddish. Ovipositor sheath reddish brown. Legs reddish brown. Wings hyaline, wing veins blackish brown.</p><p>Male. Unknown.</p><p>Comparative notes. This species differs from its congeners by having its T2 and T3 enlarged, almost fused dorsally, T4 onwards mostly retracted under tergite 3 and densely wrinkled area dentipara.</p><p>Etymology. The specific name refers to its densely wrinkled area dentipara.</p><p>Distribution. Currently known from Lao Cai Province, northwest Vietnam.</p></div>	https://treatment.plazi.org/id/0C3587E856538100FF3CFE23FBE6FF5F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Pham, Nhi Thi;Cao, Nga Thi Quynh	Pham, Nhi Thi, Cao, Nga Thi Quynh (2025): First record of the genus Cisaris Townes (Ichneumonidae: Phygadeuontinae) from Vietnam, with descriptions of four new species. Zootaxa 5683 (4): 573-583, DOI: 10.11646/zootaxa.5683.4.7, URL: https://doi.org/10.11646/zootaxa.5683.4.7
0C3587E85651810EFF3CFEB2FDF8FD0F.text	0C3587E85651810EFF3CFEB2FDF8FD0F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cisaris tergalis Pham & Cao 2025	<div><p>Cisaris tergalis Pham &amp; Cao, sp. nov.</p><p>(Figures 7, 8)</p><p>Material examined. Holotype: ♀ (IEBR), Son La Province, Son La, Quyet Tam, 5–15.v.2014, Malaise trap, Long D.K. coll.; Paratypes: 6♀ (IB), Lam Dong Province, Bidoup-Nui Ba National Park, 12°1047.34”N 108°41’45.9”E, 1,477 m a.s.l., viii–x.2017, Malaise trap, Pham T.N. leg .; 1♀ (IB), Lang Son Province, Trang Dinh district, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.413055&amp;materialsCitation.latitude=22.359446" title="Search Plazi for locations around (long 106.413055/lat 22.359446)">Chi Minh</a> commune, 22°21’34”N 106°24’47”E, 350 m a.s.l., 29.iv–5.v.2025, Malaise trap, Pham T.N., Cao T.Q.N., Cao, T.K. T., Dang T.H and Nguyen D.H leg.</p><p>Diagnosis. Antenna without white flagellomeres; scutellum sub-polished, with sparse punctures, lateral carina present basally; propodeum with moderately dense punctures, area superomedia separated from area basalis; metasomal tergites 2 and 3 almost fused together dorsally; ovipositor sheath 1.0–1.2× as long as hind tibia.</p><p>Description of holotype. Face 0.5× as high as wide, rugose punctate (Fig. 7B). Clypeal suture distinct. Clypeus 2.8× as wide as high, sub-polished with large punctures, subapical margin impressed (Fig. 7B). Mandible long, upper tooth sharp, 3.0× as long as lower tooth. Malar space matt, densely punctate, as wide as basal width of mandible (Fig. 7B). Gena wider than compound eye in lateral view, sub-polished, with moderately dense punctures (7D). Frons and vertex rugose punctate. Postero-ocellar distance 0.9× as long as ocular-ocellar distance, 1.3× median ocellus diameter (Fig. 7C). Antenna with 17 flagellomeres, slightly thickened toward apex, F1 2.2× as long as wide, 1.0× as long as F2. Occipital carina complete, ventrally curved before meeting hypostomal carina above base of mandible.</p><p>Mesosoma.Pronotum rugose punctate,except narrow impunctate dorsally.Epomia present(Fig.8A).Mesoscutum 0.7× as long as its maximum width, sub-polished, with large punctures laterally and rugose punctures medio-posteriorly. Notaulus distinct anteriorly. Scutellum moderately convex, sub-polished, with sparse punctures, lateral carina present basally (Fig. 8B). Mesopleuron extremely rough, with dense, irregular wrinkles and coarse punctures; epicnemial carina with upper end reaching to subalar prominence. Sternaulus strong anteriorly, posteriorly weaker, reaching hind margin of mesopleuron, distinctly dorsad of lower posterior corner of mesopleuron. Metapleuron with reticular wrinkles (Fig. 8A). Propodeum with moderately dense punctures, carinae strong. Area superomedia sub-polished with sparse punctures. Area basalis smooth, without punctures except punctate along surrounding carinae. Area dentipara densely punctate. Apico-median portion from posterior margin of area superomedial to apex of posterior corner of area petiolaris strongly concave longitudinally (Fig. 8D). Propodeal apophysis strong and compressed. Propodeal spiracle small, round.</p><p>Wings. Fore wing length ca. 2.9 mm, vein 1cu-a slightly postfurcal to M&amp;RS, inclivous, ventrally curved, vein 3rs-m absent, vein 2rs-m shorter than length of M between 2rs-m and 2m-cu, length of CU between 1m-cu&amp;M and 2cu-a 2.5× length of 2cu-a (Fig. 8E). Hind wing with distal abscissa of CU present, length of CU between M and cu-a 2.5× length of cu-a.</p><p>Legs. Hind coxa densely coarsely punctate, except concave area dorsally smooth. Hind femur sub-polished with moderately dense punctures, 3.9× as long as its maximum width, 0.8× as long as hind tibia length. Hind tibia coarsely sculptured, 1.1× as long as hind tarsus. Hind basitarsus 0.4× as long as tarsus.</p><p>Metasoma.Tergites smooth and shining, with scattered, small punctures. T1 2.5× as long as apical width, median dorsal carina weak, present medially, dorsolateral carina complete, ventrolateral carina present, weak anteriorly; spiracle small, round. T2 and T3 almost fused together dorsally (Fig. 8F). Laterotergite of T2 2.1× as long as apical width (Fig. 8C). Ovipositor straight (Fig. 7A). Ovipositor sheath 1.1× as long as hind tibia.</p><p>Colour. Black with reddish brown. Head black, antenna reddish, dorsal side of apical flagellomeres reddish brown, palpi yellow. Metasomal tergites from base to apex reddish brown to reddish. Ovipositor reddish. Ovipositor sheath reddish brown. Fore and mid legs reddish. Hind leg with coxa to femur reddish brown, tibia to tarsus reddish. Wings hyaline, wing veins blackish brown.</p><p>Variation. Paratypes have fore wing length 2.5–2.9 mm, malar space 1.0–1.2× as long as basal mandible width, T1 2.4–2.6× as long as apical width, laterotergite of T2 1.9–2.4×, ovipositor sheath 1.0–1.2× hind tibia length. The paratype from Lang Son Province has its F1 1.2× as long F2.</p><p>Male. Unknown.</p><p>Comparative notes. This new species resembles C. rugosus sp. nov. by having its T2 and T3 almost fused together dorsally but differs from the latter by having T4 onwards mostly exposed (vs. retracted under tergite 3) and punctate area dentipara (vs. densely wrinkled).</p><p>Etymology. The specific name refers to its characteristic of metasomal tergites 2–3.</p><p>Distribution. Currently known from Son La and Lang Son provinces, northern Vietnam and Lam Dong Province, the Central Highlands.</p></div>	https://treatment.plazi.org/id/0C3587E85651810EFF3CFEB2FDF8FD0F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Pham, Nhi Thi;Cao, Nga Thi Quynh	Pham, Nhi Thi, Cao, Nga Thi Quynh (2025): First record of the genus Cisaris Townes (Ichneumonidae: Phygadeuontinae) from Vietnam, with descriptions of four new species. Zootaxa 5683 (4): 573-583, DOI: 10.11646/zootaxa.5683.4.7, URL: https://doi.org/10.11646/zootaxa.5683.4.7
0C3587E8565F810EFF3CFD42FA63F8AE.text	0C3587E8565F810EFF3CFD42FA63F8AE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cisaris Townes 1969	<div><p>Key to the world species of Cisaris</p><p>(modified from Sun &amp; Sheng (2011) to insert four new species of Vietnam, males of the new species from Vietnam are unknown)</p><p>1. Female............................................................................................. 2</p><p>-. Male.............................................................................................. 10</p><p>2. T2 and T3 almost fused dorsally......................................................................... 3</p><p>-. T2 and T3 well separated from each other.................................................................. 4</p><p>3. T4 onwards fully exposed (Fig. 8C); area dentipara punctate (Fig. 8D)............................. C. tergalis sp. nov.</p><p>-. T4 onwards mostly retracted under tergite 3 (Fig. 6C); area dentipara densely wrinkled (Fig. 6D)....... C. rugosus sp. nov.</p><p>4. Antenna without white ring............................................................................. 5</p><p>-. Antenna with white ring.................................................................................7</p><p>5. Area superomedia well separated from area basalis............................................ C. tenuipes Townes</p><p>-. Area superomedia fused with area basalis (Figs 2D, 4D)...................................................... 6</p><p>6. Scutellum sub-polished, with sparse punctures, lateral carina weak, extending nearly to apex (Fig. 2C); smaller body size, fore wing length 2.8–3.0 mm................................................................ C. carinatus sp. nov.</p><p>-. Scutellum densely punctate, lateral carina present basally (Fig. 4C); larger body size, fore wing length 3.3–3.8 mm ............................................................................................. C. robustus sp. nov.</p><p>7. Terga red or reddish brown. Area superomedia and area petiolaris separated by a strong carina........................ 8</p><p>-. Terga black. If black with apical terga blackish brown (see Fig. 1 in Sun &amp; Sheng 2011), area superomedia and area petiolaris combined (see Fig. 1 in Sun &amp; Sheng 2011)................................................................ 9</p><p>8. Face 1.8 to 1.9× as wide as long at level of upper margin. Malar space approximately 0.6× as long as basal width of mandible. Costula originating from anterior corner of area petiolaris.................................... C. takagii Kusigemati</p><p>-. Face 2.2× as wide as long at level of upper margin. Malar space approximately 0.92× as long as basal width of mandible. Area superomedia with costula slightly behind its middle…........................................ C. mitis Pei &amp; Sheng</p><p>9. Area superomedia trapezoidal, anterior and posterior sides (carinae) very weak or almost absent, with costula at its posterior corner, transverse (see Fig. 5 in Sun &amp; Sheng 2011). Ocular-ocellar line at least 2.0× as long as largest diameter of ocellus (see Fig. 3 in Sun &amp; Sheng 2011)..................................................... C. canaliculatus Sun &amp; Sheng</p><p>-. Area superomedia hexagonal, with complete and strong carinae, costula originating from its posterior corner, leaning slightly forward laterally. Ocular-ocellar line 1.6× as long as largest diameter of ocellus..................... C. niger Kusigemati</p><p>10. Terga and hind leg entirely black (see Fig. 2 in Sun &amp; Sheng 2011). Malar space 0.42 to 0.47× as long as basal width of mandible. Area superomedia and area petiolaris completely combined.................... C. canaliculatus Sun &amp; Sheng</p><p>-. Terga brown or darkish brown. Hind leg reddish brown or darkish brown. Malar space at most 0.3× as long as basal width of mandible. Area superomedia separated from area petiolaris by strong carina...................................... 11</p><p>11. Area superomedia approximately 1.5× as wide as long, costula originating from its middle........... C. mitis Pei &amp; Sheng</p><p>-. Area superomedia approximately 2.7× as wide as long, costula originating from anterior corner of area petiolaris............................................................................................ C. takagii Kusigemati</p></div>	https://treatment.plazi.org/id/0C3587E8565F810EFF3CFD42FA63F8AE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Pham, Nhi Thi;Cao, Nga Thi Quynh	Pham, Nhi Thi, Cao, Nga Thi Quynh (2025): First record of the genus Cisaris Townes (Ichneumonidae: Phygadeuontinae) from Vietnam, with descriptions of four new species. Zootaxa 5683 (4): 573-583, DOI: 10.11646/zootaxa.5683.4.7, URL: https://doi.org/10.11646/zootaxa.5683.4.7
