identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
12018A00895DFF99649D071608A4FD80.text	12018A00895DFF99649D071608A4FD80.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Africaphes Jałoszyński 2025	<div><p>Africaphes gen. nov.</p><p>Type species: Africaphes quadripunctatus sp. nov. (here designated).</p><p>Diagnosis. Body (Figs 1–2) with distinct constriction between head and pronotum and between pronotum and elytra; thick bristles present on lateral genal regions and lateral regions of pronotum (Figs 8–9); head (Figs 3, 6) with transverse frontovertexal region; eyes situated closer to mandibular bases and antennal fossae than to occipital constriction (Figs 3, 6); antennal fossae narrowly separated at middle by distance subequal to width of scape; antenna (Figs 1–2) with trimerous club and rimmed circular structures of unknown function on all flagellomeres (Fig. 5); frontoclypeal groove lacking; frontal region in both sexes with pair of pits behind supraantennal tubercles (Figs 3, 6); posterior margin of vertex (Figs 3, 6) not bulging posterodorsad, weakly impressed at middle; submental region (Figs 8–9) demarcated laterally by deep and posteriorly converging lateral submental sulci, area between sulci subtriangular; hypostomal sulci (Figs 8–9) extending posterad behind hypostomal regions to level of posterior tentorial pits, each dividing genal region into setose lateral (outer) part and asetose inner part; occipital constriction much narrower than width of vertexal region (Figs 3, 6); maxillary palpomere 3 (Fig. 17) not enlarged (i.e., not clearly broader than scape); labrum with two sublateral ventral rows of peg-like labral sensilla (Fig. 13); pronotum (Figs 3, 6) approximately round, broadest near middle, with two pairs of antebasal pits, lacking lateral and sublateral carinae; prosternum (Fig. 18) with precoxal region much shorter than procoxal rests, with narrow, weakly elevated prosternal carina in natural position hidden between procoxae; notosternal sutures (Fig. 18) complete; procoxal cavities (Fig. 18) closed by postcoxal prosternal lobe overlapped by (but not fused with) postcoxal mesal lobe of hypomeron; hypomeral ridges (Fig. 18) lacking but inner region of each hypomeron differs from outer setose region in lacking setae; mesoventral intercoxal process (Figs 14–15) carinate, weakly elevated, narrow and extending from middle of mesoventral procoxal rests to level of posterior margins of mesocoxae, posteriorly fused with metaventrite; posterior margins of mesocoxal rests (Fig. 14) with dense setae; metaventral intercoxal process (Figs 14–15) composed of pair of narrow pointed katepisternal spines narrowly separating metacoxae; each elytron (Figs 3, 6) with two deep asetose basal foveae; mesoscutellar shield exposed between elytral bases (Fig. 3); aedeagus (Figs 20–25) with symmetrical thin-walled median lobe and asymmetrical endophallus composed of elongate sclerites, with subapical ventral ostium and with free, slender parameres. Males with secondary sexual characters on vertexal region of head, with circular or elongate glandular pits and/or tubercles.</p><p>Description. Body form (Figs 1–2) elongate, moderately slender, strongly convex, with distinct constrictions between head and pronotum and between pronotum and elytra; in known species body length ~ 1 mm.</p><p>Head (Figs 3, 6, 8–9, 17) only slightly narrower than prothorax, distinctly transverse; compound eyes situated much closer to mandibular bases and antennal fossae than to occipital constriction, genal regions in dorsal view (= tempora) rounded; vertex not bulging posterodorsad, confluent with frons, its posterior margin demarcated from occiput and slightly impressed at middle. Occipital constriction (Fig. 17; occ) much narrower than vertexal region; ‘neck’ broadening posterad. Antennal fossae small and situated anterolaterally, narrowly separated at middle by median longitudinal frontal area subequal to width of scape. Frontoclypeal groove lacking; clypeus short, subtrapezoidal, with lateral margins converging towards labrum. Genal region (Fig. 8; gen) on each side of head divided by long ventral longitudinal or oblique extension of hypostomal sulcus (Fig. 8; hs) into asetose inner and setose outer area. Gular plate (Fig. 8; gp) subtrapezoidal, covered with transverse microsculpture, with indistinct gular sulci clearly discernible only in anteriormost region; posterior tentorial pits (Fig. 8; ptp) situated slightly in front of transverse impression demarcating ventrally ‘neck’ region, each elongate and narrow.</p><p>Submentum (Fig. 8; smn) subtriangular, laterally demarcated by deep lateral submental sulci (Fig. 8; lss) converging posterad; mentum (Figs 10–11; mn) subtrapezoidal, broadest near base; prelabium (Fig. 17; prl) short, with pair of small lateral premental sclerites (Fig. 12; prms), each bearing minute seta; region between and anterior to insertions of labial palps (ligula) broader than width of palpiger and anteriorly bearing pair of setae; labial palpomere 1 (Fig. 12; lp1) annulate, palpomere 2 (Fig. 12; lp2) broader than 1 and strongly elongate, palpomere 3 (Fig. 12; lp3) narrow, elongate and distinctly shorter than 2, tapering distad and straight. Maxilla with small transverse cardo (Fig. 11; cd), subtriangular basistipes (Fig. 11; bst), elongate mediostipes (Fig. 11; mst), elongate lacinia (Fig. 11; lac) and galea (Fig. 11; gal), each with thick and not very numerous setae along distomesal margin, palpifer (Fig. 11; ppf) elongate, maxillary palp with distinctly elongate and subcylindrical palpomere 1 (Fig. 11; mxp1), long, clavate and weakly curved palpomere 2 (Fig. 11; mxp2), elongate and relatively slender palpomere 3 (Fig. 11; mxp3), and minute subconical and conspicuously slender palpomere 4 (Fig. 11; mxp4). Mandibles (Fig. 11; md) symmetrical, each broadly subtriangular, with short and broad minute mesal teeth. Labrum strongly transverse, slightly emarginate anteriorly, and with sparse, symmetrically distributed dorsal setae; ventrally with two sublateral groups of four peg-like labral sensilla (Fig. 13).</p><p>Antenna (Figs 1–2, 5) short in relation to body length, slender, with scape and pedicel each elongate and slightly broader than proximal flagellomeres; antennal clubs trimerous and slender. Each flagellomere with several large circular rimmed impressions of unknown function (modified campaniform sensilla?).</p><p>Males with secondary sexual modifications on vertexal region (Figs 4, 7) in form of presumably glandular pits or elongate inflections and/or modified setae.</p><p>Prothorax in dorsal view (Figs 3, 6) approximately round, broadest near middle. Anterior and lateral margins confluent, without anterior angles; lateral margins rounded; posterior corners feebly marked, blunt; posterior margin weakly bisinuate or weakly convex posteriorly. Pronotal base with two pairs of distinct circular antebasal pits, lacking sublateral and lateral carinae. Lateral regions of pronotum with thick bristles. Hypomeral ridges lacking, but asetose inner region of hypomeron clearly differs from setose outer region (Figs 8–9). Prosternum (Fig. 18; pst) about as long as 1/3 length of prothorax, with precoxal region much shorter than coxal rests; notosternal sutures (Fig. 18; nss) complete. Prosternal process developed as weakly elevated narrow carina in intact specimens hidden between procoxae; posterior margin of furcasternum with shallow median emargination. Procoxal cavities (Fig. 18; pcc) broadly closed; postcoxal hypomeral lobe overlapping (but not fused to) postcoxal prosternal lobe.</p><p>Mesonotum with broadly subtriangular scutellar shield (Fig. 3; mss) visible between elytral bases in intact specimens.</p><p>Mesoventrite transverse, with short prepectus, transverse and setose anteromedian impressions functioning as procoxal rests (Figs 14–15, 19; pcr) separated at middle by anterior extension of carinate mesoventral intercoxal process (Figs 14–15, 19; msvp), its posterior end fused with metaventrite.</p><p>Metanotum not studied.</p><p>Anterior region of pterothorax with one pair of moderately deep ventral foveae (Fig. 19; vf).</p><p>Metaventrite (Figs 14–15, 19) with anterior metaventral process completely fused with mesoventral process; metaventral intermetacoxal process (Figs 14–15, 19; mtvp) composed of pair of long, slender and pointed katepisternal spines narrowly separating metacoxae. Posterior margins of mesocoxal rests with dense setae. Metanepisterna and metepimera fully demarcated and narrow, not exposed in ventral view. Metendosternite with slender and strongly diverging lateral furcal arms (Fig. 19; lfa) inserted on short subquadrate stem.</p><p>Legs (Figs 1–2) short, lacking peculiar features, all femora clavate, tarsi (Fig. 16) with massive claws.</p><p>Elytra (Figs 1–2, 3, 6) together oval, with elongate humeral calli, each elytron with two deep asetose basal foveae (Fig. 3; bef) well-discernible in intact specimens.</p><p>Hind wings functional.</p><p>Abdomen lacking peculiar characters.</p><p>Aedeagus (Figs 20–25) with symmetrical and thin-walled median lobe, ventral subapical ostium, weakly sclerotized and asymmetrical endophallic structures with tubular flagellum, and slender parameres with setose apices.</p><p>Etymology. Africaphes combines the continent name with - phes used in generic names of several Stenichnini. Gender masculine.</p><p>Composition and distribution. Africaphes includes two nominal species known to occur in Burundi and Rwanda; one specimen presumably representing one more species from Rwanda was also seen.</p><p>Remarks. Africaphes is externally most similar to the cosmopolitan Microscydmus, but clearly differs in deep lateral submental sulci, which Microscydmus lacks. Moreover, Microscydmus has partly obliterated notosternal sutures visible only near the procoxal cavity and at the anterior prosternal margin (notosternal sutures complete in Africaphes), and complete hypomeral ridges (lacking in Africaphes) (for characters of Microscydmus, see Jałoszyński (2014, 2015b, 2017) and Jałoszyński &amp; Newton (2017)). The only other continental Afrotropical genus with lateral submental sulci is Stenichnodes . Morphology of this genus was described and illustrated in Jałoszyński (2015c). Stenichnodes differs from Africaphes in numerous characters:eyes situated posteriorly, closer to occipital constriction than to mandibular bases (anteriorly, closer to mandibular bases in Africaphes), the occipital constriction about as wide as vertexal region (much narrower in Africaphes), the lack of ‘stenichnine bristles’ (present in Africaphes), complete hypomeral ridges (lacking in Africaphes), each elytron with one vestigial basal fovea (two deep foveae in Africaphes), and a peculiar structure of the aedeagus, which has a basal membranous area bearing a median lentiform sclerotized plate internally connected to a long apodeme aligned with the central longitudinal axis of the aedeagus, to which a muscle (membranous area retractor) is attached (in Africaphes membranous area is ventral and subapical, lentiform sclerotized plate and internal apodeme are lacking).</p><p>Africaphes can be distinguished from all world Stenichnini genera by a combination of synapomorphies: antennae with trimerous clubs; eyes situated anteriorly; occipital constriction much narrower than vertex; genal regions divided by posterior extensions of hypostomal sulci into mesal asetose and outer setose portions; lateral submental sulci present; pronotum with two pairs of antebasal pits, lacking lateral carinae; prosternum demarcated from hypomera by complete notosternal sutures, precoxal region of prosternum vestigial; hypomeral ridges lacking; mesoventral intercoxal process carinate and fused with metaventrite; metaventral intermetacoxal process composed of a pair of spines narrowly separating metacoxae; each elytron with two asetose basal foveae; and aedeagus with parameres. The only structure found in Africaphes and not known in any other Stenichnini is a unique set of large circular rimmed impressions on each flagellomere, found by SEM studies in both included species.</p><p>Africaphes can be identified using the following keys (to facilitate identifications a separate key to only Afrotropical continental genera is given after a worldwide key). Reasons why some genera are not included in the key, and some names are treated as possible synonyms pending clarification, were discussed in Jałoszyński (2024a).</p></div>	https://treatment.plazi.org/id/12018A00895DFF99649D071608A4FD80	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jałoszyński, Paweł	Jałoszyński, Paweł (2025): Africaphes gen. nov., a new genus of Stenichnini in Burundi and Rwanda (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 5666 (2): 262-278, DOI: 10.11646/zootaxa.5666.2.7, URL: https://doi.org/10.11646/zootaxa.5666.2.7
12018A008955FF9D649D0393091DF9AA.text	12018A008955FF9D649D0393091DF9AA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Napochomorphus Franz 1980	<div><p>Updated key to world genera of Stenichnini (excl. Napochomorphus, Neuraphomorphus, Psepharobius and Taphroscydmus)</p><p>1 Frons and vertex with elevated subtriangular median ‘platform’ covered with scale-like microsculpture and narrowing posteriorly...................................... Afroeudesis Franz [Brazil, Ivory Coast, Tanzania, China, Nepal]</p><p>- Frons and vertex lacking subtriangular ‘platform’........................................................... 2</p><p>2 Pronotum deeply and rapidly constricted between base and middle, in dorsal view hourglass-shaped........................................................................................ Scydmepitoxis Jałoszyński [Australia]</p><p>- Pronotum not hourglass-shaped.......................................................................... 3</p><p>3 Pronotum with deep median longitudinal groove extending from near base to middle of pronotal disc........................................................................ Aenigmaphes Jałoszyński [Papua New Guinea, Sulawesi]</p><p>- Pronotum lacking median longitudinal groove.............................................................. 4</p><p>4 Labial palps conspicuously elongate, each longer than mandible, palpomere 3 about ten times as long as broad at base.......................................................................... Bellendenker Jałoszyński [Australia]</p><p>- Labial palps shorter than mandibles, palpomere 3 at most 6 times as long as broad................................. 5</p><p>5 Pronotum at each side near posterior corner with one long erect penicillus composed of several setae.................................... Penicillidmus Jałoszyński [Australia, Papua New Guinea, Philippines, undescribed species in SE Asia]</p><p>- Pronotum lacking posterolateral penicilli.................................................................. 6</p><p>6 Each femoral clava dorsally with longitudinal groove bearing submedian oval glandular opening...................... 7</p><p>- Longitudinal groove and oval opening on each femur absent................................................... 8</p><p>7 Constriction between antennomeres 10 and 11 similar as that between all remaining flagellomeres; antennae conspicuously long and slender, nearly filiform............................ Loeblites Franz [China, Philippines, Thailand, Malaysia]</p><p>- Antennomere 11 sits tightly on apex of 10 without constriction; antennae not filiform................................... Syndicus Motschulsky [Australia, Bhutan, Cambodia, China, India, Indonesia, Japan, Laos, Malaysia, Myanmar, Nepal, Philippines, Singapore, Sri Lanka, Thailand, Vietnam]</p><p>8 Pair of basal elytral foveae connected anteriorly by inversely U-shaped groove extending onto elytral articulating lobe.......... Horaeomorphus L.W. Schaufuss [Himalayas, SE Asia, China, Japan, Taiwan, New Guinea, Australia, Fiji, Mauritius]</p><p>- Basal elytral pits, if present, not connected anteriorly by inversely U-shaped groove................................ 9</p><p>9 Occipital constriction conspicuously narrow, as narrow as &lt;1/5 of head width across eyes, in dorsal view anteriorly delimited by posterolateral angulate projections of vertex and posteriorly by anterolateral subtriangular projections of occiput, and mesoventral process reaching middle of mesocoxae, clearly separated from short and narrow anterior metaventral process............................................................ Neuraphanax Reitter [Colombia, Peru, Venezuela]</p><p>- Occipital constriction as wide as 1/4 of head width or wider, lacking anteriorly-directed subtriangular posterolateral projections, or, if small projections are present, then mesoventral process is posteriorly fused with metaventrite and separates mesocoxae on their entire length.................................................................................... 10</p><p>10 Mesoventral intermesocoxal process anteriorly with curved lateral arms forming massive trident-like structure................................................................................... Tridensius Jałoszyński [Venezuela]</p><p>- Mesoventral process, if present, lacking anterolateral arms................................................... 11</p><p>11 Adcoxal region of prothoracic hypomeron forming conspicuous longitudinal elevation with anterior and posterior regions developed as subtriangular projecting lobes, with procoxal cavity situated on mesal surface of elevation; metaventrite with longitudinal groove along each lateral margin demarcating broad lateral area................................................................................................... Protoconnus Franz [Central and South America]</p><p>- Adcoxal regions of hypomera not strongly elevated, procoxal cavities situated on ventral surface of prosternum; metaventrite lacking lateral longitudinal grooves..................................................................... 12</p><p>12 At least antennomeres 10 and 11 tightly assembled, lacking constriction between them; head slightly to strongly transverse and flattened......................................................... Plaumanniola Costa Lima [South America]</p><p>- Antennomeres 10 and 11 with distinct constriction between them; head as long as broad or elongate, not flattened....... 13</p><p>13 Head in lateral view subtriangular, ventrally flattened, dorsally strongly convex and angulate, with the highest site above antennal fossae, strongly declining anterad and less strongly posterad.......................................... 14</p><p>- Head in lateral view rounded or subrectangular, not subtriangular.............................................. 17</p><p>14 Each lateral pronotal carina mesally accompanied by broad longitudinal impression with sharply marked mesal margin........................................................................... Siamitoides Jałoszyński [Indonesia]</p><p>- Pronotum lacking sublateral longitudinal impressions....................................................... 15</p><p>15 Vertex, prosternum, mesoventrite, metaventrite behind mesocoxae and abdominal sternites all with small tufts and/or rosettes of bristles; lateral submental sulci present.................... Siamites Franz [Indonesia, Malaysia, Thailand, Vietnam]</p><p>- Body lacking tufts and rosettes of bristles; lateral submental sulci lacking....................................... 16</p><p>16 Maxillary palpomere 4 conspicuously slender, rod-like; mesoventral process posteriorly fused with metaventrite behind mesocoxal rests............................... Elacatophora L.W. Schaufuss [India, Indonesia, Malaysia, Thailand]</p><p>- Maxillary palpomere 4 subconical; mesoventral process posteriorly reaching about half of mesocoxal rests, not fused with metaventrite............................................................... Lophioderus Casey [N America]</p><p>17 Head in ventral view with inversely U-shaped, sharply marked and usually glossy impression on submental area............................................................ Himaloconnus Franz [India, Japan, Nepal, Taiwan, Thailand]</p><p>- Head in ventral view lacking U-shaped impression......................................................... 18</p><p>18 Anterior margin of prosternum only half as wide as posterior margin........ Sibylloconnus Jałoszyński [Dominican Rep.]</p><p>- Anterior margin of prosternum about as wide as posterior margin.............................................. 19</p><p>19 Outer region of each metacoxa hidden under posterolateral metaventral lobe..................................... 20</p><p>- Outer regions of metacoxae exposed..................................................................... 24</p><p>20 Metafemur in both sexes with ventral subtriangular tooth; metacoxae narrowly separated.................................................................................................. Homoconnus Sharp [C &amp; S America]</p><p>- Metafemur in both sexes lacking ventral subtriangular tooth; metacoxae contiguous............................... 21</p><p>21 Mesocoxae separated by carinate mesoventral process....................................................... 22</p><p>- Mesocoxae not separated, mesoventral process lacking...................................................... 23</p><p>22 Pronotal base with two pairs of small pits, inner pair connected by transverse groove; lateral pronotal margin with conspicuously dense thick bristles; notosternal sutures complete; each elytron with one basal fovea; endophallus strongly asymmetrical....................................................................................... Delius Casey [USA]</p><p>- Pronotal base lacking paired pits and transverse groove; lateral pronotal margin only with thin setae; notosternal sutures largely obliterated, marked only as short notches near anterior prothoracic margin; each elytron with two minute basal foveae; endophallus lightly sclerotized and symmetrical....................................................................................................... Leptocharis Reitter [Algeria, Tunisia, Turkey, Corsica, Sardinia, Spain]</p><p>23 Pronotum clearly broadest in front of middle; aedeagus with darkly sclerotized, complex and strongly asymmetrical endophallus....................................................................... Neladius Casey [USA]</p><p>- Pronotum barrel-shaped, broadest near middle; endophallus lightly sclerotized and symmetrical............................................................................................... Turkocharis Jałoszyński [Turkey]</p><p>24 Mesocoxae not separated by mesoventral process (low vestigial carina can be hidden between mesocoxae, but does not separate them)............................................................................................. 25</p><p>- Mesoventral process present and posteriorly reaching at least middle of mesocoxae, clearly separating at least their anterior halves............................................................................................. 31</p><p>25 Metacoxae clearly separated and occipital constriction in dorsal view nearly as narrow as half of vertex.................................................................................. Contreuconnus Jałoszyński [Madagascar]</p><p>- Metacoxae contiguous or subcontiguous, in the latter case occipital constriction barely marked, about as wide as vertex.. 26</p><p>26 Width of occipital constriction in dorsal view subequal to width of head between eyes; tempora at most as long as eyes; hypomeral ridges absent; sides of pronotum lacking thick bristles.............................................. 27</p><p>- Occipital constriction distinctly narrower than frons between eyes; tempora longer than eyes; hypomeral ridges present; sides of pronotum with thick bristles......................................................................... 29</p><p>27 Tempora absent, eyes adjacent to occipital constriction; aedeagus in ventral view with symmetrical endophallus............................................................................ Austrostenichnus Franz [New Caledonia]</p><p>- Tempora short but distinct, eyes not adjacent to occipital constriction; endophallus strongly asymmetrical.............. 28</p><p>28 Pronotum with antebasal transverse groove but lacking pits; basal elytral foveae absent; procoxal cavities narrowly open posteriorly; pronotum widest near anterior third and distinctly narrowing towards base; mandible with mesal preapical tooth...................................................................... Palaeoscydmaenus Franz [Australia]</p><p>- Pronotum with antebasal transverse groove connecting outer antebasal pits; two shallow basal elytral foveae present; procoxal cavities closed posteriorly by overlapping postcoxal lobes of furcasternum and hypomeron; pronotum subequal in width in front of middle and at base, with shallow narrowing in posterior third; mandible lacking mesal preapical tooth................................................................................. Stenichnaphes Franz [New Zealand]</p><p>29 Procoxal cavities posteriorly broadly open, posterolateral region of prosternum not forming process directed towards hypomeron.................................................................................................... Magellanoconnus Franz partim [RSA, Zimbabwe, New Zealand, Argentina, Bolivia, Chile, Colombia, Ecuador]</p><p>- Procoxal cavities posteriorly closed by overlapping postcoxal processes of prosternum and hypomeron................ 30</p><p>30 Pronotum with distinct sublateral carinae in posterior 1/5–1/4, base with one pair of small inner pits in middle region separated by narrow longitudinal wrinkle or carina, and with variously distinct lateral impressions adjacent to sublateral carinae; precoxal region of prosternum subequal in length to procoxal rests; minute but well developed mesoventral process and anterior metaventral process hidden between mesocoxae; metafemur in males with long ventral process........................................................................................... Spinosciacharis Jałoszyński [Australia]</p><p>- Pronotum without sublateral carinae, base with transverse groove (complete or disrupted in middle), in some species also with two pairs of pits; precoxal region of prosternum as long as only half length of procoxal rests; between mesocoxae no traces of mesoventral process and anterior metaventral process; metafemora in males unmodified....................................................... Heterotetramelus Franz [Australia, Tasmania, New Caledonia; possibly also New Zealand]</p><p>31 Mesoventral process between mesocoxae distinctly broadened and broadening with flat smooth asetose surface with sharp lateral margins, posteriorly touching conspicuously broad subtriangular anterior metaventral process with truncate apex; mesocoxal rests conspicuously deep and filled with dense setae............................................... 32</p><p>- Mesoventral process nearly parallel-sided or gradually broadening posterad, lacking flat asetose region between mesocoxae and posteriorly fused with metaventrite, if not fused, then anterior metaventral process minute, subquadrate, and mesocoxal rests not filled with dense setae............................................................................. 35</p><p>32 Pronotum lacking thick bristles on lateral margins; pronotal base with two broad submedian longitudinal carinulae separated by shallow median impression and with two pairs of shallow lateral pits............. Bicarinulodes Jałoszyński [Venezuela]</p><p>- Pronotum with dense thick bristles on lateral margins; pronotal base with transverse groove and small paired pits, lacking longitudinal carinulae................................................................................ 33</p><p>33 Frons and vertex lacking median longitudinal impression; pronotal base lacking pair of outer pits outside transverse groove....................................................................... Meridaphes Jałoszyński [Venezuela]</p><p>- Frons and vertex with median longitudinal impression; pronotal base with pair of outer pits outside transverse groove.... 34</p><p>34 Posterior margin of vertex distinctly angulate and convex anteriorly; pronotum broadest slightly behind middle, lacking sublateral carinae............................................... Stenichnoconnus Franz [Suriname, Venezuela]</p><p>- Posterior margin of vertex weakly arcuate; pronotum broadest in front of middle, with distinct sublateral carinae....................................................................................... Pseudoraphes Franz [Peru]</p><p>35 Mesoventral process posteriorly reaches about middle of mesocoxae, its tip not fused with metaventrite............... 36</p><p>- Mesoventral process either reaching (or nearly reaching) posterior margin of mesocoxae or together with anterior metaventral process forming continuous carina separating mesocoxae on entire length, at most with transverse groove marking fusion site .................................................................................................. 47</p><p>36 Metacoxae broadly separated; pronotum with median pair of short and nearly parallel longitudinal grooves and with sublateral carinae reaching middle of pronotum and conspicuously widely separated from lateral pronotal margins......................................................................................... Stenichnoteras Scott [Seychelles]</p><p>- Metacoxae contiguous or subcontiguous; pronotum lacking median pair of longitudinal grooves, sublateral carinae absent.. .................................................................................................. 37</p><p>37 Procoxal cavities posteriorly broadly open, posterolateral region of prosternum not forming process directed towards hypomeron.................................................................................................... Magellanoconnus Franz partim [RSA, Zimbabwe, New Zealand, Argentina, Bolivia, Chile, Colombia, Ecuador]</p><p>- Procoxal cavities posteriorly closed by overlapping postcoxal processes of prosternum and hypomeron or very narrowly open, with narrow gap between these processes................................................................. 38</p><p>38 Pronotum lacking antebasal pits, grooves and impressions................................................... 39</p><p>- Pronotum with antebasal sculpture: pits, and/or transverse antebasal groove, and/or median longitudinal antebasal carina... .................................................................................................. 40</p><p>39 Occipital constriction only slightly narrower than head between eyes; pronotum with sinuate lateral margins, convex anteriorly and slightly concave posteriorly; elytral bases together about as wide as head and as pronotal base; each elytron with one basal fovea........................................................................... Catalinus Casey [USA]</p><p>- Occipital constriction much narrower than head between eyes; pronotum with lateral margins convex on entire length; elytral bases together much narrower than head and much narrower than pronotal base; basal elytral foveae lacking............................................................................... Clepsydraphes Jałoszyński [Madagascar]</p><p>40 Occipital constriction deep, much narrower than head between eyes; tempora in dorsal view subequal to or longer than eyes.................................................................. Neuraphoconnus Franz [New Caledonia]</p><p>- Occipital constriction shallow, indistinctly narrower than head between eyes (or eyes absent); tempora in dorsal view much shorter than eyes (or eyes absent)....................................................................... 41</p><p>41 Each elytron with one clearly visible densely setose basal fovea..................................................... Neuraphes Thomson [all Europe, N Africa, China, India, Japan, Kazakhstan, Nagorno-Karabakh, Nepal, Asian Russia, Asian Turkey]</p><p>- Basal elytral foveae, if present, not filled with dense setae.................................................... 42</p><p>42 Hypomeral ridge present.............................................................................. 43</p><p>- Hypomeral ridge absent............................................................................... 44</p><p>43 Lateral margins of pronotum with thick bristles; lateral pronotal edges present at least in posterior third; basal elytral fovea distinct.............. Scydmoraphes Reitter [all Europe, N Africa, Japan, Pakistan, Asian Russia, Asian Turkey, Canada]</p><p>- Lateral margins of pronotum with inconspicuous thin setae; lateral pronotal edges absent; basal elytral fovea vestigial, barely discernible.......................................................... Madagaphes Jałoszyński [Madagascar]</p><p>44 Pronotum with paired antebasal pits or transverse band of many punctiform pits but without transverse antebasal groove; mandibles conspicuously narrow and slender, sickle-shaped, lacking preapical teeth, but often microserrate.................. Stenichnus Thomson [all Europe, N Africa, China, India, Israel, Japan, Kazakhstan, Mongolia, Nepal, Pakistan, Asian Russia, Taiwan, Asian Turkey, Uzbekistan, Canada, USA]</p><p>- Pronotum with transverse antebasal groove and one pair of outer antebasal pits or impressions; mandibles subtriangular and each with one preapical tooth.......................................................................... 45</p><p>45 Frons in both sexes with conspicuous broad impression; aedeagus with strongly asymmetrical endophallus and lacking parameres............................................................. Brachycepsis Brendel [Canada, USA]</p><p>- Frons in both sexes convex or flattened, lacking conspicuous impression; aedeagus with symmetrical endophallus and with parameres.......................................................................................... 46</p><p>46 Lateral submental sulci posteriorly broadly separated, delimiting trapezoidal submental area................................................................................ Scydmaenilla King [Australia, Tasmania, New Zealand]</p><p>- Lateral submental sulci posteriorly approximate, delimiting triangular submental area.................................................................................................. Zeanichnus Jałoszyński [New Zealand]</p><p>47 Lateral submental sulci present......................................................................... 48</p><p>- Lateral submental sulci absent.......................................................................... 57</p><p>48 Outer antebasal pronotal pit connected with deep longitudinal lateral groove running along lateral pronotal margin and nearly reaching anterior pronotal margin............................................. Sinonichnus Jałoszyński [China]</p><p>- Pronotal disc lacking lateral longitudinal grooves.......................................................... 49</p><p>49 Antennae with sharply delimited dimerous clubs............................. Nogunius Jałoszyński [Japan, Taiwan]</p><p>- Antennae gradually broadened or with trimerous clubs...................................................... 50</p><p>50 Eyes anterior, situated much closer to mandibular bases than to occipital constriction.............................. 51</p><p>- Eyes posterior, situated closer to occipital constriction than to mandibular bases, or occupying entire area between antennal fossa and occiput.................................................................................... 53</p><p>51 Maxillary palpomere 3 at most as broad as scape; pronotal base with two pairs of pits; hypomeral ridges lacking; each elytron with two distinct basal foveae........................................ Africaphes Jałoszyński [Burundi, Rwanda]</p><p>- Maxillary palpomere 3 conspicuously broadened, much broader than scape; pronotal base with transverse groove connecting one pair of pits or lacking pits and groove; hypomeral ridges present; basal elytral foveae vestigial, barely discernible even in transparent slides.................................................................................... 52</p><p>52 Lateral submental sulci strongly converging posterad; hypostomal ridges reaching posteriorly middle between mentum and posterior tentorial pits; pronotal base with transverse groove connecting one pair of pits; proximal protarsomeres in males conspicuously broadened and with dense cushion of setae on tarsomere 1.. Mexiconnus Jałoszyński [Costa Rica, Mexico]</p><p>- Lateral submental sulci subparallel; hypostomal ridges reaching posterior tentorial pits; pronotal base lacking groove and pits; protarsi in males unmodified................................................. Palposcydmus Jałoszyński [India]</p><p>53 Eyes conspicuously large, occupying entire area between antennal fossa and occiput; body strikingly stout, almost suboval.......................................... Obesoconnus Jałoszyński [Thailand, Costa Rica, French Guiana, Mexico]</p><p>- Eyes small, not occupying entire length of head between antennal fossae and occiput; body slender................... 54</p><p>54 Lateral submental sulci posteriorly broadly separated, delimiting trapezoidal submental area............................... Alloraphes Franz [Caribbeans, Central &amp; South America] &amp; Stenichnodes Franz [Chad, Kenya, India, Indonesia, Sri Lanka] {possible synonyms}</p><p>- Lateral submental sulci posteriorly approximate, delimiting triangular submental area............................. 55</p><p>55 Lateral margins of pronotum with conspicuously dense thick erect bristles; hypomeral ridges complete; aedeagus with parameres........................................ Leptoderoides Croissandeau [Israel, Lebanon, ‘Syria’, Turkey]</p><p>- Lateral margins of pronotum only with thin setae; hypomeral ridges absent; aedeagus without parameres.............. 56</p><p>56 Pronotum with three pairs of antebasal pits................................... Parascydmus Casey [Canada, USA]</p><p>- Pronotum with two pairs of antebasal pits, inner pair separated by small median tubercle or wrinkle....................................................................................... Rutaraphes Jałoszyński [Japan, Taiwan]</p><p>57 Metaventral intermetacoxal process composed of a pair of elongate (at least twice as long as broad) pointed spines...... 58</p><p>- Metaventral intermetacoxal process lacking pair of strongly elongate pointed spines adjacent mesally, at most with short median notch separating triangular lateral lobes not longer than wide................................................. 66</p><p>58 Pronotum elongate subtrapezoidal, broadest at base and strongly narrowing anterad......................................................... Napoconnus Franz [equatorial &amp; subequatorial Africa, SE Asia, Nepal, Mauritius, S America]</p><p>- Pronotum rounded, broadest near middle or in front of middle, narrowing anterad and posterad...................... 59</p><p>59 Notosternal sutures complete.......................................................................... 60</p><p>- Notosternal sutures largely obliterated................................................................... 61</p><p>60 Pronotum broadest clearly in front of middle; aedeagus with symmetrical median lobe................................................................................... Oreoeudesis Franz [Cameroon, Kenya, Rwanda, Tanzania]</p><p>- Pronotum broadest near middle; aedeagus with strongly asymmetrical median lobe....... Schuelkelia Jałoszyński [China]</p><p>61 Anterior prosternal margin strongly concave; frons in males modified, with projections and/or impressions................................................................................ Protandroconnus Franz [C &amp; S America]</p><p>- Anterior prosternal margin straight; frons in males unmodified................................................ 62</p><p>62 Postcoxal prosternal process fused with postcoxal hypomeral process without trace of fusion site.................... 63</p><p>- Postcoxal prosternal process clearly demarcated from postcoxal hypomeral process............................... 64</p><p>63 Tempora shorter than eyes; pronotal base with antebasal transverse groove and one pair of longitudinally elongate lateral impressions close to lateral margins and not connected with groove, without sublateral carinae; each elytron with one distinct basal fovea................................................................. Perumicrus Jałoszyński [Peru] - Tempora shorter than eyes; pronotal base without sublateral carinae, with transverse groove, without pits; each elytron with two vestigial basal foveae..................................................... Amimoscydmus Jałoszyński [Brazil]</p><p>64 Inner regions of hypomera extremely narrow, barely discernible, so that procoxal cavities are nearly adjacent to hypomeral ridges; vestiges of notosternal sutures discernible only at anterior prosternal margin........ Mimoscydmus Franz [Brazil]</p><p>- Inner regions of hypomera clearly separating procoxal cavities from hypomeral ridges; notosternal sutures obliterated in their middle regions, so that parts of sutures are discernible both at anterior prosternal margin and at procoxal cavity......... 65</p><p>65 Tempora n dorsal view much longer than eyes; inner region of hypomeron at most as wide as procoxal cavity and in postcoxal area divided by longitudinal carina........................................ Heteroscydmus Franz [Bolivia, Brazil]</p><p>- Tempora n dorsal view at most as long as eyes (in most cases shorter); inner region of hypomeron much wider than procoxal cavity and not divided by longitudinal carina................... Microscydmus Saulcy &amp; Croissandeau [cosmopolitan]</p><p>66 Notosternal sutures entirely absent; anterior prosternal margin conspicuously deeply concave; in males frons with elevated median circular glandular ‘platform’ bearing median tuft of setae............. Trichocircus Jałoszyński [C &amp; S America]</p><p>- Notosternal sutures complete or obliterated at middle length, and then their anterior and posterior portions discernible; anterior prosternal margin indistinctly concave or straight; in males frons lacking median circular glandular ‘platform’.......... 67</p><p>67 Precoxal region of prosternum clearly more than half as long as procoxal rests and at most 3 times as wide as long....... 68</p><p>- Precoxal region of prosternum at most half as long as procoxal rests, in many cases much shorter or even vestigial, 4 and more times as wide as long................................................................................. 72</p><p>68 Metacoxae widely or narrowly separated by distinct subtrapezoidal or subtriangular metaventral process.............. 69</p><p>- Metacoxae contiguous or almost contiguous.............................................................. 71</p><p>69 Pronotum with long median longitudinal carina; lateral pronotal edges present in posterior half.......................................................................................... Kangarooconnus Jałoszyński [Australia]</p><p>- Pronotum lacking median longitudinal carina and lacking lateral edges......................................... 70</p><p>70 Only short rod-like apical regions of parameres discernible; antennae gradually thickened; males with small setal penicillus inserted on cuticular papilla on apex of each elytron......................... Euconnomorphus Franz [Bolivia, Peru]</p><p>- Rod-like parameres entirely separated from median lobe and discernible on entire length; antennae with sharply delimited trimerous or tetramerous clubs; elytral apices in males lacking papillae with penicilli........................................ Euconnus Thomson partim [mainly subgen. Borneoconnus Franz from Sabah, and a group of Sri Lankan species]</p><p>71 Body length over 2.5 mm; pronotum with paired antebasal pits, lacking transverse groove; posterior tentorial pits strongly elongate, slot-like.................................... Sciacharis Broun [Australia, New Zealand, South America]</p><p>- Body length below 1 mm; pronotum with antebasal transverse groove, lacking paired pits; posterior tentorial pits punctiform............................................................................. Microraphes Franz [Brazil]</p><p>72 Procoxal cavities posteriorly narrowly open (postcoxal processes of prosternum and hypomeron not overlapping); each elytron with distinct humeral denticle projecting anteriorly............................. Leascydmus Jałoszyński [Australia]</p><p>- Procoxal cavities posteriorly closed (postcoxal processes of prosternum and hypomeron overlapping); elytra lacking humeral denticles........................................................................................... 73</p><p>73 Aedeagus lacking parameres............................................. Parapseudoconnus Franz [S America]</p><p>- Aedeagus with parameres............................................................................. 74</p><p>74 Metacoxae clearly touching at middle.................................................................... 75</p><p>- Metacoxae at least slightly separated.................................................................... 76</p><p>75 Precoxal region of prosternum vestigial, barely discernible; parameres so broad that almost enclosing median lobe, in lateral view their basal halves as wide as median lobe.......................... Venezolanoconnus Franz [C &amp; S America]</p><p>- Precoxal region of prosternum as long as half-length of procoxal rests, clearly visible in front of procoxae; parameres slender, rod-like.............................................................. Archiconnus Franz [C &amp; S America]</p><p>76 Mesoventral process not developed in front of mesocoxae; entire hypomera densely setose up to procoxal cavities; hypomeral ridges entirely lacking.................................................. Scydmaenozila Jałoszyński [Australia]</p><p>- Mesoventral process extending anteriorly far in front of mesocoxae, where it reaches procoxal rests on anterior region of mesoventrite; each hypomeron divided into setose outer region and asetose inner (adcoxal) region, in most cases demarcated by hypomeral ridge; or entire hypomera asetose................................................................. Euconnus Thomson partim [worldwide], Anhoraeomorphus Franz [Madagascar], Noctophus Casey [USA] {possible synonyms}</p></div>	https://treatment.plazi.org/id/12018A008955FF9D649D0393091DF9AA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jałoszyński, Paweł	Jałoszyński, Paweł (2025): Africaphes gen. nov., a new genus of Stenichnini in Burundi and Rwanda (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 5666 (2): 262-278, DOI: 10.11646/zootaxa.5666.2.7, URL: https://doi.org/10.11646/zootaxa.5666.2.7
12018A008951FF9E649D07BD0DCEFE79.text	12018A008951FF9E649D07BD0DCEFE79.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neuraphomorphus Reitter 1882	<div><p>Key to genera of Stenichnini of continental Africa south of Sahara (excl. Neuraphomorphus)</p><p>1 Frons and vertex with elevated subtriangular median ‘platform’ covered with scale-like microsculpture and narrowing posteriorly........................................................ Afroeudesis Franz [Ivory Coast, Tanzania]</p><p>- Frons and vertex lacking subtriangular ‘platform’........................................................... 2</p><p>2 Procoxal cavities posteriorly broadly open, posterolateral region of prosternum not forming process directed towards hypomeron....................................................... Magellanoconnus Franz [RSA, Zimbabwe]</p><p>- Procoxal cavities posteriorly closed by overlapping postcoxal processes of prosternum and hypomeron................. 3</p><p>3 Eyes posterior, situated closer to occipital constriction than to mandibular bases....... Stenichnodes Franz [Chad, Kenya]</p><p>- Eyes anterior, situated much closer to mandibular bases than to occipital constriction, or lacking...................... 4</p><p>4 Lateral submental sulci present....................................... Africaphes Jałoszyński [Burundi, Rwanda]</p><p>- Lateral submental sulci lacking.......................................................................... 5</p><p>5 Metaventral intermetacoxal process lacking pair of strongly elongate pointed spines adjacent mesally........................................................................................... Euconnus Thomson [worldwide]</p><p>- Metaventral intermetacoxal process composed of a pair of elongate (at least twice as long as broad) pointed spines....... 6</p><p>6 Pronotum subtrapezoidal, broadest at base and strongly narrowing anterad............................................................................................... Napoconnus Franz [equatorial &amp; subequatorial Africa]</p><p>- Pronotum rounded, broadest near middle or in front of middle, narrowing anterad and posterad....................... 7</p><p>7 Notosternal sutures complete............................ Oreoeudesis Franz [Cameroon, Kenya, Rwanda, Tanzania]</p><p>- Notosternal sutures largely obliterated........................ Microscydmus Saulcy &amp; Croissandeau [cosmopolitan]</p></div>	https://treatment.plazi.org/id/12018A008951FF9E649D07BD0DCEFE79	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jałoszyński, Paweł	Jałoszyński, Paweł (2025): Africaphes gen. nov., a new genus of Stenichnini in Burundi and Rwanda (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 5666 (2): 262-278, DOI: 10.11646/zootaxa.5666.2.7, URL: https://doi.org/10.11646/zootaxa.5666.2.7
12018A008952FF9E649D00CA0C7CF80C.text	12018A008952FF9E649D00CA0C7CF80C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Africaphes quadripunctatus Jaloszynski 2025	<div><p>Africaphes quadripunctatus Jałoszyński, sp. nov.</p><p>(Figs 1, 3–5, 8, 10, 14, 16, 20–21)</p><p>Type material studied. Holotype (Burundi): ♂, two labels: “ Afrika / Burundi / Muramvya / 10 km NE Bugarama / 27.2.1985 / leg. Brachat /Mühle” [white, printed], „ AFRICAPHES \ quadripunctatus m. \ P. Jałoszyński, 2025 \ HOLOTYPUS“ [red, printed] (MNHW).</p><p>Diagnosis. Male: posterior vertexal region with two distinct pits separated by small tubercle and with adjacent long and thick modified setae; submental region elongate and strongly narrowing posterad; antennomere 8 strongly transverse.</p><p>Description. Body of male (Fig. 1) moderately dark brown with slightly darker head and slightly lighter appendages, setae yellowish brown; BL 1.15 mm.</p><p>Head (Figs 3–4, 8) broadest across eyes, HL 0.20 mm, HW 0.26 mm; temple in dorsal view (Fig. 3) slightly longer than eye; frontovertexal region strongly transverse; each eye large but weakly convex, coarsely faceted. Punctures on dorsum inconspicuous; setae short, sparse and suberect; outer genal regions with lateral and ventral bristles. Secondary sexual characters: posterior vertexal region with pair of circular pits separated by distance about twice as wide as their diameters, median area just anterior to pits with small elongate tubercles bearing two long and flat modified setae, two conspicuously long, thick, flattened and strongly curved setae inserted on median area just posterior to pits (Fig. 4). Ventrally, head with strongly elongate submental region (Fig. 8; smn) strongly narrowing posterad. Antenna (Figs 1, 5) slender but short in relation to body length, AnL 0.48 mm, antennomeres 1 and 2 each strongly elongate, 2.5 times as long as broad, 3 slightly transverse, with sharply marked proximal marginal ‘collar’, 4–7 each weakly elongate and gradually increasing in width and length, 8 as wide as 7 and much shorter, strongly transverse, 9 and 10 each about as long as broad and much wider than 8, 11 much shorter than 9 and 10 combined, as wide as 10 and about 1.5 times as long as broad.</p><p>Pronotum (Fig. 3) rounded, broadest near middle; PL 0.28 mm, PW 0.30 mm. Anterior and lateral margins strongly rounded; posterior corners obtuse-angled, posterior margin weakly bisinuate. Pronotal disc with two pairs of round and deep antebasal pits and with impression laterad each outer pit, on dorsolateral surface. Disc with fine, inconspicuous punctures; setae about as long, sparse and suberect as those on head dorsum, dorsally admixed with sparse bristles, which become denser on lateral pronotal regions and on outer hypomera.</p><p>Elytra (Figs 1, 3) together oval, broadest slightly in front of middle; EL 0.68 mm, EW 0.50 mm, EI 1.35; humeral calli distinct, elongate and mesally demarcated by shallow impressions, basal foveae deep and widely separated. Punctures and setae as those on pronotal disc.</p><p>Hind wings functional.</p><p>Legs short, lacking peculiar modifications.</p><p>Aedeagus (Figs 20–21) weakly elongate, AeL 0.13 mm; in ventral view with elongate subtriangular apex; endophallus with several curved elongate sclerites; parameres moderately slender, each with 2–3 apical setae.</p><p>Female. Unknown.</p><p>Distribution. SW Burundi (Muramvya Province).</p><p>Etymology. The adjective quadripunctatus refers to the head of male which bears two anterior pits and two presumably glandular posterior pits.</p><p>Remarks. Externally, males of A. quadripunctatus and A. bipunctatus sp. nov. clearly differ in the head structures (posterior glandular structures in the former are more elaborate and include two pits, a median tubercle and modified setae, whereas the latter species has only a simple median elongate ‘slot’), and in the shape of antennomere 8, which is strongly transverse in A. quadripunctatus and as long as wide in A. bipunctatus .</p></div>	https://treatment.plazi.org/id/12018A008952FF9E649D00CA0C7CF80C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jałoszyński, Paweł	Jałoszyński, Paweł (2025): Africaphes gen. nov., a new genus of Stenichnini in Burundi and Rwanda (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 5666 (2): 262-278, DOI: 10.11646/zootaxa.5666.2.7, URL: https://doi.org/10.11646/zootaxa.5666.2.7
12018A008953FF9F649D01B708A2F9CC.text	12018A008953FF9F649D01B708A2F9CC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Africaphes bipunctatus Jaloszynski 2025	<div><p>Africaphes bipunctatus Jałoszyński, sp. nov.</p><p>(Figs 2, 6–7, 9, 11–13, 15, 17–19, 22–25)</p><p>Type material studied. Holotype (Rwanda): ♂, two labels: “ Afrika / Rwanda / Bez. Cyangugu / Urw. v. Nyakabuye / 1700–1900m /14.– 27.2.1985 / leg. V.u. C.Brachat ” [white, printed], „ AFRICAPHES \ bipunctatus m. \ P. Jałoszyński, 2025 \ HOLOTYPUS“ [red, printed] (MNHW) . Paratypes (3 exx.): 1 ♂, same data as for holotype; 1 ♀, same data except for 22.4.1985, leg. Mühle; 1 ♀, same data except for 9.1.1986, leg. Mühle (cPJ) .</p><p>Diagnosis. Male: posterior vertexal region with median longitudinal ‘slot’; submental region subtriangular with all sides subequal in length; antennomere 8 as long as broad.</p><p>Description. Body of male (Fig. 2) moderately dark brown with slightly lighter appendages, setae yellowish brown; BL 0.95–1.00 mm.</p><p>Head (Figs 6–7) broadest across eyes, HL 0.20 mm, HW 0.23 mm; temple in dorsal view (Fig. 6) distinctly longer than eye; frontovertexal region strongly transverse; each eye moderately large and weakly convex, coarsely faceted. Punctures on dorsum inconspicuous; setae short, sparse and suberect; outer genal regions with lateral and ventral bristles. Secondary sexual characters: posterior vertexal region with single median longitudinal ‘slot’ (Fig. 7). Ventrally, head with submental region (Fig. 9) subtriangular with all its sides subequal in length. Antenna (Fig. 2) slender but short in relation to body length, AnL 0.45–0.48 mm, antennomeres 1 and 2 each strongly elongate, slightly more than twice as long as broad, 3 slightly transverse, with sharply marked proximal marginal ‘collar’, 4–6 each weakly elongate and gradually increasing in width and length, 7 weakly elongate, clearly wider than 6, 8 as wide as 7 and slightly shorter, about as long as broad, 9 and 10 each about as long as broad and much wider than 8, 11 much shorter than 9 and 10 combined, as wide as 10 and about 1.5 times as long as broad.</p><p>Pronotum (Fig. 6) rounded, broadest near middle; PL 0.25 mm, PW 0.25 mm. Anterior and lateral margins strongly rounded; posterior corners obtuse-angled, posterior margin indistinctly rounded. Pronotal disc with two pairs of round and deep antebasal pits and with barely discernible impression laterad outer pit, on dorsolateral surface. Disc with fine, inconspicuous punctures; setae about as long, sparse and suberect as those on head dorsum, dorsally admixed with sparse bristles, which become denser on lateral pronotal regions and on outer hypomera.</p><p>Elytra (Figs 2, 6) together oval, broadest near middle; EL 0.50–0.55 mm, EW 0.40 mm, EI 1.25–1.38; humeral calli distinct, elongate and mesally demarcated by shallow impressions, basal foveae deep and widely separated. Punctures and setae as those on pronotal disc.</p><p>Hind wings functional.</p><p>Legs short, lacking peculiar modifications.</p><p>Aedeagus (Figs 22–25) stout, AeL 0.11 mm; in ventral view with short subtriangular apex; endophallus with several curved elongate sclerites; parameres moderately slender, each with 2–3 apical setae.</p><p>Female. Externally similar to male, differs in lacking posterior vertexal modification and in slightly shorter antennae in relation to body length. BL 1.00 mm; HL 0.20 mm, HW 0.23–0.24 mm, AnL 0.43 mm; PL 0.25 mm, PW 0.25–0.28 mm; EL 0.55 mm, EW 0.40 mm, EI 1.38.</p><p>Distribution. SW Rwanda (formerly Cyangugu Province, currently part of Western Province).</p><p>Etymology. The adjective bipunctatus refers to the head of male which bears only two anterior pits.</p><p>Remarks. For differences between A. quadripunctatus and A. bipunctatus see Remarks for the former species.</p></div>	https://treatment.plazi.org/id/12018A008953FF9F649D01B708A2F9CC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jałoszyński, Paweł	Jałoszyński, Paweł (2025): Africaphes gen. nov., a new genus of Stenichnini in Burundi and Rwanda (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 5666 (2): 262-278, DOI: 10.11646/zootaxa.5666.2.7, URL: https://doi.org/10.11646/zootaxa.5666.2.7
