identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
5433833C8EC5564D9B935F38877B5AB0.text	5433833C8EC5564D9B935F38877B5AB0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tropodiaptomus kampucheaensis Koompoot & Sanoamuang 2025	<div><p>Tropodiaptomus kampucheaensis sp. nov.</p><p>Figs 10, 11, 12, 13, 14, 15, 16</p><p>Tropodiaptomus sp.: Chaicharoen and Sanoamuang (2022): 1, 6, 7, 9, 11, 12, 13.</p><p>Material examined.</p><p>Holotype: Cambodia • one ♂ (adult), Cambodia, Kampong Cham Province, Tbong Khmum District, 12°32'34"N, 105°36'94"E; 16 June 2006; water temperature 28.9 ° C, pH 7.7, and conductivity 108.6 µS cm - 1. W. Mahasrap leg.; a rice field (Fig. 1); accession number: THNHM -lv-19360; dissected, mounted on one slide in glycerol, covered with a coverslip, and sealed with nail polish . Allotype: Cambodia • one ♀ (adult); location, date and collectors as for holotype; accession number: THNHM -lv-19361, completely dissected, mounted on one slide in glycerol, covered with a coverslip, and sealed with nail polish . Paratypes: Cambodia • three ♂ (adult) and three ♀ (adult); date and collectors as for holotype; accession number: THNHM -lv-19362, undissected and preserved in 70 % ethanol .</p><p>Additional occurrence locality.</p><p>A temporary roadside canal, Kandal province, Cambodia (11°38'72"N, 104°20'19"E); 16 June 2006; water temperature 29.2 ° C, pH 8.2, and conductivity 94.1 µS cm - 1.</p><p>Description of adult male.</p><p>Total body length, measured from anterior margin of rostrum to posterior margin of caudal rami, 0.94–0.95 mm (mean 0.947 mm, n = 3) (Figs 10 A, 12 A). Body smaller and slender than in female. Prosome ~ 2.5 × as long as urosome (Fig. 10 A, 12 A). Pdg 4 separated dorso-laterally from Pdg 5. Lateral wings of Pdg 5 small, symmetrical, with one tiny spine at distal corner and one inner sensillum-like spine (Figs 10 C, 12 B).</p><p>Urosome (Figs 10 A – C, 12 A – C) with five somites, unornamented. Genital somite dilated postero-laterally on both sides (Fig. 12 C), shorter than wide. Urosomite 2 ~ as long as wide, urosomites 3 and 4 shorter than wide. Urosomites 2 and 3 without hairs on ventral side (Figs 10 B, 12 C). Urosomite 4 with expanded right dorso-posterior margin. Anal somite asymmetrical, left side slightly longer than right side. Caudal rami symmetrical, each ramus ~ 1.5 × as long as wide, inner margins hairy (Fig. 12 C). Ventral surfaces of both caudal rami without any prominent structures. Each ramus armed with five plumose setae and one base dorsal seta.</p><p>Antennules: asymmetrical, long, reaching to posterior end of genital somite. Left antennule (Fig. 13 A): 25 - segmented. Armature formulae as in Table 4. Right antennule geniculated (Figs 10 D, 12 E – G), consisting of 22 segments, strongly dilated between segment XIII and segment XVIII. Segment XIII with largest strong spinous process, one seta, and one aesthetasc. Antepenultimate segment (segment XX) longer than next segment. Spinous process on antepenultimate segment straight with an outwardly curved tip, reaching 2 / 3 of next segment (Figs 10 D, 12 G). Armature formulae as in Table 5.</p><p>Antenna (Fig. 13 B) biramous. Coxa and basis with one and two bare setae on distal corner, respectively. Enp two-segmented; Enp - 1 with two inner median setae and small spinules on distal outer margin; Enp - 2 bilobed, bearing eight setae on inner lobe and six setae on outer lobe. Exp seven-segmented, longer than Enp. Exp 1–6 with setal formulae 1, 3, 1, 1, 1, and 1, respectively. Exp 7 with one inner seta and three apical setae.</p><p>Mandible (Fig. 13 C): coxal gnathobase with cutting edge bearing eight well-chitinized teeth and one setulose seta. Basis with four inner setae; one located proximally and three distally. Enp two-segmented; Enp - 1 with four setae on inner distal corner; Enp - 2 with eight setae apically. Exp four-segmented; Exp - 1–3 each with one seta on inner margin; Exp - 4 with three setae apically; all setae bare.</p><p>Maxillule (Fig. 13 D): precoxal arthrite with nine strong setae laterally, four slender submarginal setae, and one anterior seta. Coxal epipodite with nine setae; coxal endite with four setae. Basal endites each with four setae proximally and four setae distally; basal exite with one outer seta. Enp with eight setae distally. Exp with six setae.</p><p>Maxilla (Fig. 13 E): praecoxa fused to coxa. Proximal and distal endites on praecoxa each with three setae apically. Two coxal endites each with three setae apically. Allobasis with three setae apically. Enp two-segmented, each with three setae.</p><p>Maxilliped (Fig. 13 F) praecoxa and coxa fused, three medial lobes on syncoxa: setal formulae 2, 3, 3, respectively; subdistal inner margin produced into a spherical lobe with a patch of tiny spinules. Basis with three setae at distal inner margin, with a row of tiny spinules proximately. Enp six-segmented, with 2, 3, 2, 2, 1 + 1, and 4 setae, respectively.</p><p>Swimming legs (P 1 – P 4) (Fig. 14 A – D) biramous, with three-segmented rami, except for two-segmented Enp on P 1. Exp longer than Enp. Each coxa on P 1 – P 4 with one pinnate seta at innermost distal corner. P 4 coxa with an extraordinarily long inner seta (Fig. 14 D). P 1 and P 2 bases without setae. P 3 and P 4 bases with a bare outer seta. P 2 Enp - 2 with a Schmeil’s organ on posterior surface (Fig. 14 B). Armature formulae of P 1 – P 4 as in Table 6.</p><p>P 5 (Figs 11 A – F, 12 D) highly asymmetrical. Intercoxal sclerite reduced, inner distal margin not produced. Coxae fused to intercoxal sclerite. Right P 5 (Figs 11 A – C, F, 12 D): right coxa larger than left one, with a triangular spine on outer subdistal corner of posterior surface. Basis (Figs 11 A, B, F, 12 D) rectangular, ~ 1.5 × as long as wide, ornamented with a thin, longitudinal hyaline membrane located at mid-length of inner margin (Figs 11 B, 12 D), and a short seta at subdistal outer corner. Exp - 1 (Figs 11 A – C, 12 D) shorter than wide, produce into acute process on outer distal corner. Exp - 2 (Figs 11 A, C, 12 D) elongated, somewhat rectangular, ~ 1.9 × as long as wide, with a thin crescent-shaped lamella near base of principal lateral spine (Figs 11 C, 12 D). Principal lateral spine robust, slightly curved, acutely pointed, ~ 0.9 × as long as Exp - 2 segment, located sub-distally, close to end claw, with spinules on distal outer and inner margins. End claw long, slender, and sickle-shaped, with a pointed tip and ornamented with spinules on inner margin; ~ 1.6 × as long as Exp - 2. Enp one-segmented, conical, reaching of Exp - 1, and bearing a row of apical spinules.</p><p>Left P 5 (Figs 11 A, B, D, E, 12 D) slightly bent inwards, reaching to 1 / 4 of right Exp - 2. Coxa as long as wide, with one triangular spine inserted on posterior lobe, at mid-distal outer margin. Basis (Figs 11 B, 12 D) rectangular, elongated, ~ 1.6 × as long as wide, with one short seta near outer corner distally; inner margin straight, without hyaline lamella. Exp (Fig. 11 D, E) flattened, elongated, ~ 1.0 × as long as basis; inner margin one-lobed, with uniform serration; anterior surface ornamented with two hairy pads in center, proximal one located under Enp; apex of Exp with usual combination of digitiform appendix and spinulate seta. Enp (Fig. 11 E) one-segmented, slightly cylindrical, extending to mid-length of Exp; apex rounded with a row of subapical spinules.</p><p>Description of adult female.</p><p>Total body length, measured from anterior margin of rostrum to posterior margin of caudal rami, 1.53–1.55 mm (mean 1.54 mm, n = 3). Prosome: urosome ratio ~ 2.3: 1. Prosome similar to that of male. Rostrum (Fig. 15 B) symmetrical, with moderate strong and acutely pointed, paired filaments. Pdg 4 and Pdg 5 completely fused. Last pedigerous somite (Figs 15 D, F, 16 B, C) with nearly symmetrical postero-lateral wings, each wing armed with one postero-lateral spine.</p><p>Urosome (Figs 15 D, F, 16 A – D) two-segmented. Genital-double somite (Figs 15 A, D, F, 16 B – D) asymmetrical, ~ twice as long as wide, longer than anal somite and caudal rami combined; left proximal margin slightly dilated, with ~ 2 / 3 of right proximal margin expanded; distal part narrower than proximal part. Genital area on ventral surface with opercular pad protecting gonopores, and with rectangular and semicircular expansion (Fig. 16 C). Anal somite ~ 1.2 × as long as caudal rami. Caudal rami (Figs 15 F, 16 B) symmetrical, ~ 2.1 × as long as wide, parallel, with hairy outer and inner margins, each ramus with six setae; dorsal setae jointed, longer than principal setae.</p><p>Left antennule, antenna, mandible, maxillule, maxilla, maxilliped, and P 1 – P 4 similar to those of the male.</p><p>P 5 (Figs 15 E, G, 16 E) symmetrical. Coxa roughly spherical, with a stout, triangular spine anteriolaterally. Basis (Fig. 16 E) with one short, smooth sensory seta on distolateral margin (Fig. 16 E). Exp (Figs 15 E, G, 16 E) three-segmented; Exp - 1 cylindrical, ~ 2 × as long as wide, with almost straight inner and outer margins. Exp - 2 sub-triangular, with a row of strong spinules along both margins; Exp - 3 fused into a small prominence on Exp - 2, armed with one short spine and one long seta apically. Enp (Fig. 16 E) one-segmented, ~ 0.8 × as long as Exp - 1, armed with two strong, unequal spiniform setae; outer seta longest, with a row of tiny spinules apically.</p><p>Distribution and ecology.</p><p>Currently, T. kampucheaensis sp. nov. has been found only in two temporary-water habitats: a temporary pond in Kampong Cham Province and a shallow canal in Kandal Province, Cambodia. Representatives of the new species were found in two sites from the 147 sampled locations in five provinces (Kampong Chnang, Kampong Cham, Kendal, Prey Veng, and Takeo) in Cambodia. This species is rare and currently endemic to Cambodia. The newly discovered species was observed alongside two diaptomid species, namely Allodiaptomus raoi Kiefer, 1936, and Mongolodiaptomus mekongensis Sanoamuang &amp; Watiroyram, 2018 .</p><p>Etymology.</p><p>The specific name kampucheaensis refers to the name Kampuchea, which is the native name for Cambodia in the Khmer language, where the type locality is located. The suffix - ensis originates from Latin and indicates the specific origin.</p></div>	https://treatment.plazi.org/id/5433833C8EC5564D9B935F38877B5AB0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Koompoot, Kamonwan;Sanoamuang, Laorsri	Koompoot, Kamonwan, Sanoamuang, Laorsri (2025): Two new species of Tropodiaptomus Kiefer, 1932 (Copepoda, Calanoida, Diaptomidae) from temporary waters in Thailand and Cambodia with a key to Southeast Asian species. ZooKeys 1249: 339-369, DOI: 10.3897/zookeys.1249.157214
34FD9972891B582C86F3C68F4834969E.text	34FD9972891B582C86F3C68F4834969E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tropodiaptomus lannaensis Koompoot & Sanoamuang 2025	<div><p>Tropodiaptomus lannaensis sp. nov.</p><p>Figs 2, 3, 4, 5, 6, 7, 8, 9</p><p>Tropodiaptomus sp.: Sanoamuang and Dabseepai 2021: 5, 8, 16, 18, 23.</p><p>Material examined.</p><p>Holotype: Thailand • one ♂ (adult), northern Thailand, Mae Hong Son Province, Mae La Noi District, 18°33'00.28"N, 97°91'01.42"E; 8 December 2012; water temperature 23.1 ° C, pH 8.75, conductivity 386 µS cm - 1, and dissolved oxygen 8.84 mg L - 1. P. Dabseepai and K. Koompoot leg.; a temporary pond with aquatic plants near the road No. 108 (Fig. 1); accession number: THNHM -lv-18787; dissected, mounted on one slide in glycerol, covered with a coverslip, and sealed with nail polish . Paratypes: Thailand • three ♂ (adult) and two ♀ (adult); date and collectors as for holotype; accession number: THNHM -lv-18789, undissected and preserved in 70 % ethanol .</p><p>Description of adult male.</p><p>Total body length, measured from anterior margin of rostrum to posterior margin of caudal rami, 1.16–1.21 mm (mean = 1.15 mm, n = 3) (Fig. 2 B). Body smaller and slender than in female. Prosome ~ 2.2 × as long as urosome (Figs 2 B, 3 A). Rostrum (Fig. 2 A) well-developed, with two spiniform processes. Pdg 4 and Pdg 5 separated by distinct septum dorso-laterally. Lateral wings of Pdg 5 small and slightly asymmetrical; right wing larger than left one; both wings with one tiny spine at distal corner and one inner sensillum-like spine (Fig. 3 B).</p><p>Urosome (Figs 2 C, 3 A – C) with five somites, unornamented. Genital somite slightly dilated postero-laterally on right side, shorter than wide. Urosomite 2 longer than wide, urosomite 3 approximately as long as wide, urosomite 4 with expanded right dorso-posterior corner (Fig. 3 A). Urosomites 2 and 3 without ventral hairs (Figs 2 C, 3 C). Urosomites 3 and 4, anal somite, and caudal rami bent to right side. Anal somite asymmetrical, shorter than preceding urosomites. Caudal rami symmetrical, each ramus ~ 2.3 × as long as wide, inner margin hairy (Fig. 3 C). Ventral surface of right caudal ramus without any prominence structures. Each ramus armed with six setae, subequal in length and size, plumose: dorsal seta bare and thinner than others.</p><p>Antennules asymmetrical, extending to approximately posterior end of genital somite. Left antennule (Fig. 4 A): 25 - segmented. Armature formulae as in Table 1. Right antennule geniculated (Fig. 3 D – F), consisting of 22 segments, strongly dilated between segment XIII and segment XVIII (Fig. 3 E). Segment XIII with largest strong spinous process. Antepenultimate segment (segment XX) longer than next segment. Spinous process on antepenultimate segment straight and slightly bent at distal end, reaching 2 / 3 of next segment (Fig. 3 F). Armature formulae as in Table 2.</p><p>Antenna (Fig. 4 B) biramous. Coxa and basis with one and two bare setae on inner distal corner, respectively. Enp two-segmented; Enp - 1 with two setae along inner margin and spinules on distal part of outer margin; Enp - 2 with nine setae along inner margin, seven setae apically; all setae bare. Exp seven-segmented: Exp - 1–6 with 1, 3, 1, 1, 1, 1, setae along inner margin; Exp - 7 with one seta on inner margin and three setae apically; all setae bare.</p><p>Mandible (Fig. 4 C): ~ 6 cuspidate teeth dorsally and one seta on coxal gnathobase dorsally. Basis with two serrate setae and two bare setae along inner margin. Enp - 1 with four setae on inner distal corner. Enp - 2 with eight setae apically. Exp four-segmented. Exp - 1–3 each with one seta on inner margin; Exp - 4 with three setae apically; all setae bare.</p><p>Maxillule (Fig. 4 D): praecoxal arthrite with nine strong setae laterally and four slender submarginal setae. Coxal endite with four setae; coxal epipodite with nine setae. Basal endites fused to segment bearing them: proximal and distal endites, each with four setae apically; basal exite with one short seta. Enp - 1 and Enp - 2 each with four setae apically, proximal segment fused to basis. Exp unsegmented with six setae apically and hairy inner margin.</p><p>Maxilla (Fig. 4 E): praecoxa fused to coxa. Proximal and distal endites on praecoxa, each with three setae apically. Two coxal endites, each with three setae apically. Allobasis with four setae apically. Enp two-segmented, each with three setae.</p><p>Maxilliped (Fig. 4 F): four medial lobes on syncoxa: setal formulae 1, 2, 3, 3, respectively; subdistal inner margin produced into a spherical lobe with a patch of tiny spinules. Basis with three setae along distal inner margin, with a row of tiny spinules proximately. Enp six-segmented, with 2, 3, 2, 2, 2, and 4 setae, respectively.</p><p>Swimming legs (P 1 – P 4) (Fig. 5 A – D) biramous, with three-segmented rami, except for two-segmented Enp on P 1. Exp longer than Enp. Each coxa on P 1 – P 4 with one pinnate seta at innermost distal corner. P 1 – P 3 bases without setae. P 4 basis with one outer seta (Fig. 5 D). P 2 Enp - 2 with Schmeil’s organ on posterior surface of (Fig. 5 B). Armature formulae of P 1 – P 4 as in Table 3.</p><p>P 5 (Figs 2 D – H, 6 A, B): intercoxal sclerite fused to coxa, produced into a triangular lobe distally (Fig. 6 A). Coxae asymmetrical, right part more prominent, each bears a stout spine on posterior surface; right spine larger than left spine. Right P 5 (Figs 2 D, 6 A, B), basis rectangular, ~ 1.5 × as long as wide; ornamented with a small semicircular knob at proximal inner margin and a larger, triangular hyaline knob occurring close to sub-proximal inner margin (Figs 2 G, 6 A); subdistal outer margin with a thin seta. Exp - 1 shorter than wide, with a hyaline lobe on inner margin; outer distal corner with an acute tip (Fig. 6 A, B). Exp - 2 somewhat cylindrical, ~ 1.5 × as long as wide, carrying accessory slender spine located at proximal 1 / 3 of outer margin and ~ 3 / 4 as long as outer distal spine (Figs 2 D, 6 A, B); outer distal spine moderately strong, slightly curved and acutely pointed, ornamented with spinules along inner margin, ~ 0.8 length of Exp - 2, and located at anterior 2 / 3 of segment (Fig. 6 A, B). Distal accessory element minute, situated close to insertion of end claw (Figs 2 D, 6 A). Enp one-segmented, conical shaped, reaching ~ 1 / 3 of Exp - 2, bearing a row of apical spinules.</p><p>Left P 5 (Figs 2 D – F, H, 6 A, B) slightly bent inwards, reaching to mid-length of right Exp - 2. Coxa with relatively short spine inserted on ~ mid-length posterior lobe at outer margin. Basis rectangular, elongated, ~ 1.8 × as long as wide, bearing a small smooth seta at outer distal margin (Figs 2 D, 6 A); inner margin almost straight, without hyaline outgrowths. Exp somewhat semicircular-shaped, flattened and ~ 1.6 × as long as wide; outer margin curved, inner margin two-lobed, both lobes with large denticles, small denticles near distal end only (Fig. 2 E, H); apex of Exp with typical digitiform appendix and spinulate seta, while anterior surface displays two elongate, hairy pads (Fig. 2 E, H). Enp one-segmented and dilated proximally, extending to mid-length of Exp; apex rounded with a row of subapical spinules (Fig. 6 A, B).</p><p>Description of adult female.</p><p>Total body length, measured from anterior margin of rostrum to posterior margin of caudal rami, 1.27–1.41 mm (mean 1.35 mm, n = 3) (Figs 7 A, 8 A). Prosome: urosome ratio ~ 2.1: 1. Rostrum (Fig. 7 B) symmetrical, with moderately long point and ornamented with a pair of sensilla. Pdg 4 and Pdg 5 completely fused. Last pedigerous somite with asymmetrical posterolateral wings (Figs 7 D, 8 A, B); left wing larger and longer than right wing. Each wing with one hyaline spine (Figs 7 D, 8 B, D).</p><p>Urosome (Figs 7 D, E, 8 A – D) two-segmented. Genital double-somite (Figs 7 D, 8 A, B) asymmetrical, longer than urosomite 2, anal somite, and caudal rami combined (Fig. 8 A, B); right proximal region slightly expanded, right distal corner produced into a small lobe; left margin slightly curved, with a small spine at proximal 1 / 3 of somite (Fig. 8 B). A pair of gonopores and copulatory pores located centrally at ~ 1 / 3 length of genital double-somite (Figs 7 E, 8 C). Urosomite 2 symmetrical, shorter than wide. Anal somite symmetrical, as long as length of caudal rami (Fig. 8 B). Caudal rami parallel, symmetrical; each ramus ~ 1.5 × as long as wide and with hairy outer and inner margins (Fig. 8 A, B). All principal caudal setae slightly dilated anteriorly.</p><p>Left antennule, antenna, mouthparts, and P 1 – P 4 similar to those of the male. Antennules symmetrical.</p><p>P 5 (Figs 7 F, G, 9 A, B) asymmetrical. Intercoxal sclerite forming a narrow, elongate triangle. Coxa massive, with broadly strong process situated anterolaterally on distal outer margin; coxal spine on right side larger and longer than left side (Figs 7 F, 9 B). Basis with a bare, minute sensory seta on outer margin (Figs 7 F, 9 B). Exp (Figs 7 F, G, 9 A, B) three-segmented. Exp - 1 cylindrical, ~ 1.9 × as long as wide, convex outer margin and almost straight inner margin; Exp - 2 sub-triangular, with a row of strong spinules along both margins; Exp - 3 fused into small prominence on Exp - 2, armed with one short spine and one long seta apically. Enp one-segmented (Figs 7 F, 9 A, B), ~ 0.7 × as long as Exp - 1, armed with two strong, unequal spiniform setae; outer seta longest, with a row of apical spinules.</p><p>Distribution and ecology.</p><p>Tropodiaptomus lannaensis sp. nov. has so far been found exclusively in its type locality, a temporary pond in Mae Hong Son Province, northern Thailand. At the time of the new species’ collection, no other calanoid copepods were present at the same site. Representatives of the new species were recorded solely once among more than 5,000 sampled locations in Thailand. This species is rare and currently endemic to Thailand.</p><p>Etymology.</p><p>The specific epithet lannaensis is derived from Lanna, the name of the area in present-day northern Thailand from the 13 th to 18 th centuries in which the type locality is located. The name is an adjective in the nominative singular, feminine gender. The suffix - ensis originates from Latin and indicates the specific origin.</p></div>	https://treatment.plazi.org/id/34FD9972891B582C86F3C68F4834969E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Koompoot, Kamonwan;Sanoamuang, Laorsri	Koompoot, Kamonwan, Sanoamuang, Laorsri (2025): Two new species of Tropodiaptomus Kiefer, 1932 (Copepoda, Calanoida, Diaptomidae) from temporary waters in Thailand and Cambodia with a key to Southeast Asian species. ZooKeys 1249: 339-369, DOI: 10.3897/zookeys.1249.157214
